Lipovskaya gully (Donskaya Luka): Olenekian, Russian Federation

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Lypbalkodus gladius
Novikov 2010
Coelacanthimorpha - Coelacanthiformes - Latimeriidae
? Holophagus sp. Egerton 1861
Novikov 2010
Dipnoi - Ceratodontidae
Ceratodus jechartiensis Minikh 1977
Novikov 2010
? Microceratodus sp. Texeira 1954
Novikov 2010
Dipnoi - Ptychoceratodontidae
Ceratodus donensis Vorobyeva and Minikh 1968
Novikov 2010
recombined as Ptychoceratodus donensis
Dipnomorpha - Arganodontidae
Ceratodus multicristatus Vorobyeva and Minikh 1968
Novikov 2010
recombined as Arganodus multicristatus
subspecies: Ceratodus multicristatus lipovensis
Bystrowianidae
Dromotectum informal new species
Novikov 2010
    = Dromotectum abditum n. sp. Shishkin et al. 2014
Shishkin et al. 2014
PIN 1043/1104 (holotype)
Trilophosauria
Doniceps lipovensis Otschev and Rykov 1968
Sennikov 1999 1 element
Saratov University 104/3106, premaxilla (specimen lost)
Anomodontia - Kannemeyeriidae
Putillosaurus sennikovi n. gen., n. sp. Surkov 2005
Ivakhnenko 2008 1 specimen
PIN 1043/614 (holotype)
unclassified
Vitalia grata n. gen., n. sp. Ivakhnenko 1973
3 specimens
PIN 4173/126 (SGU 104/3105) (type; prevoiusly included in the hypodigm of Coelodontognathus donensis by Ochev, 1967), PIN 1043/627, 1043/628; all are dentaries; current catalogue number of type and catalogue numbers of referred specimens are from Arkhangelskii & Sennikov (2008)
Reptilia - Procolophonidae
Tichvinskia enigmatica n. sp. Chudinov and Vjushkov 1956
Chudinov and Vjushkov 1956 1 element
recombined as Orenburgia enigmaticus
PIN 1043/1 (type)
? Kapes sp. Ivakhnenko 1975
Novikov 2010
Reptilia - Eosauropterygia - Cymatosauridae
Nothosauria indet. Baur 1889
Sennikov 1999 10 elements
    = Tanaisosaurus kalandadzei n. gen., n. sp. Sennikov 2001
Sennikov 2001
PIN 1043/663, left humerus (holotype). PIN nos. 1043/520–522, 661, 662, and 707, quadrates; PIN no. 1043/705, left scapula; and, probably, PIN no. 1043/706 and 750, fragmentary left clavicles
Reptilia
Tsylmosuchus donensis n. sp. Sennikov 1990
Sennikov 1999 1 element
nomen dubium belonging to Crocopoda
PIN 1043/42, cervical vertebra (holotype)
Bystrowisuchus flerovi n. gen., n. sp. Sennikov 2012
Sennikov 2012 7 elements
PIN 1043/1346, anterior thoracic vertebra (holotype). PIN 1043/393, neural spine of epistropheus; PIN 1043/1128, incomplete cervical vertebra; PIN 1043/ 656, 1493, incomplete posterior cervical vertebrae; PIN 1043/147, incomplete anterior thoracic vertebra; PIN 1043/831, right ilium
Scythosuchus basileus n. gen., n. sp. Sennikov 1999
Sennikov 1999 114 elements
PIN 1043/144, fragmentary cervical vertebra (holotype). PIN 1043/537, 538, 540, 607, and 608, fragmentary postorbitals; PIN 1043/574 and 615–617, fragmentary maxillae; PIN 1043/142 and 612, left squamosals; PIN 1043/525, articular part of the left ramus of the lower jaw; PIN 1043/575–584 and 636–639, teeth; PIN 1043/393, spinous process of the epistropheus; PIN 1043/143 and 630, spinous processes of the cervical vertebrae; PIN 1043/506 and 507, spinous processes of the posterior cervical vertebrae; PIN 1043/656, cervical vertebra; PIN 1043/657 and 658, posterior cervical and anterior thoracic vertebrae; PIN 1043/532, 609, and 659, spinous processes of anterior thoracic vertebrae; PIN 1043/533, 621, and 622, spinous processes of posterior thoracic vertebrae; PIN 1043/146–176 and 531, thoracic vertebrae; PIN 1043/660, sacral vertebra; PIN 1043/548–573 and 631–634, caudal vertebrae; PIN 1043/511, proximal fragment of the left humerus; PIN 1043/652, fragmentary right ilium; PIN 1043/517, 518, and 625 fragmentary femurs; PIN 1043/515 right tibia; PIN 1043/516, fragmentary fibula; PIN 1043/547 right calcaneum; PIN 1043/544–546 distal tarsal elements.
Scolotosuchus basileus n. gen., n. sp. Sennikov 2022
Sennikov 2022
Holotype: PIN 1043/833 (neural arch of the mid-cervical vertebra). In addition to the holotype, specimens PIN, nos. 1043/658, 828, posterior cervical vertebrae, specimen PIN, no. 1043/531, incomplete neural arch of the posterior cervical vertebra, specimens PIN, nos. 1043/506, 507, and 834, spinous processes of posterior cervical vertebrae, specimen PIN, no. 1043/657, neural arch of the first trunk vertebra, specimens PIN, nos. 1043/822, 1584, incomplete neural arches of the first trunk vertebrae, specimen PIN, no. 1043/824, neural arch of the second trunk vertebra, specimens PIN, nos. 1043/630, 1323, 1324, 1325, and 1340, neural arches of the anterior thoracic vertebrae, specimens PIN, nos. 1043/143, 751, 759, and 1583, spinous processes of the anterior thoracic vertebrae, specimen PIN, no. 1043/511, proximal fragment of the left humerus, specimen PIN, no. 1043/1117, left ilium, specimen PIN, no. 1043/652, fragment of the right ilium, specimen PIN, no. 1043/1472, left ischium, specimen PIN, no. 1043/961, proximal fragment of the right femur, specimen PIN, no. 1043/962, proximal and distal fragment of the right femur, specimen PIN, no. 1043/515, right tibia, specimen PIN, no. 1043/516, fragment of the fibula, specimen PIN, no. 1043/547, right calcaneum, specimens PIN, nos. 1043/544– 546, and distal tarsal elements.
Coelodontognathus ricovi n. sp. Ochev 1967
Arkhangelskii and Sennikov 2008 2 specimens
PIN 4173/127 (SGU 104/3101) (type), PIN 4173/128 (SGU 104/3102); both are dentaries
Coelodontognathus donensis n. gen., n. sp. Ochev 1967
Arkhangelskii and Sennikov 2008 2 specimens
PIN 4173/129 (SGU 104/3103) (type), PIN 4173/130 (SGU 104/3104); both are dentaries
Reptilia - Tanystropheidae
Augustaburiania vatagini n. gen., n. sp. Sennikov 2011
Sennikov 2011 60 elements
PIN 1043/587, cervical vertebra (holotype). PIN no. 1043/842, epistropheus; PIN, nos. 1043/840, 841, 1141, 1327, 1328, 1392, cervical vertebrae; PIN, nos. 1043/588, 651, fragments of cervical vertebrae; PIN, nos. 1043/585, 586, 843, 1142, posterior cervical vertebrae; PIN, nos. 1043/589–594, 844, 845, 1143, 1144, 1310, pectoral vertebrae; PIN, nos. 1043/397, 398, 598, neural arches of pectoral vertebrae; PIN, no. 1043/595, sacral vertebra; PIN, nos. 1043/596, 597, 847–850, 1145–1147, caudal vertebrae; PIN, no. 1043/648, fragment of right humerus; PIN, no. 1043/771, fragment of left humerus; PIN, nos. 1043/649, 650, 851, left femora; PIN, nos. 1043/603, 671, proximal fragments of left femora; PIN, no. 1043/606, proximal fragment of right femur; PIN, nos. 1043/601, 604, 641, 642, distal fragments of femora; PIN, no. 1043/514, articulated distal fragment of femur and proximal fragment of tibia (left); PIN, nos. 1043/512, 513, 644–646, 770, proximal fragments of tibiae; PIN, no. 1043/643, distal fragment of tibia; PIN, no. 1043/647, proximal fragment of fibla; and PIN, nos. 1043/602, 1393, metatarsals.
Amphibia - Temnospondyli
Parotosuchus panteleevi
Sennikov 1999
Amphibia - Temnospondyli - Trematosauridae
Trematosaurus galae n. sp. Novikov 2010
Novikov 2010 115 elements
PIN 1043/774, preorbital skull fragment without the narial region (holotype). Eight fragments of the postorbital skull region (specimens PIN, nos. 1043/59, 75, 779, 783, 867, 976, 996, 1089); four premaxillae (PIN, nos. 1043/54, 104, 105, 810); one maxillary fragment (PIN, no. 1043/370); one postfrontal (PIN, no. 1043/977); two postorbitals (PIN, nos. 1043/886, 978); five complete and fragmentary jugals (PIN, nos. 1043/880, 912, 1050, 1093, 1097); 11 complete and fragmentary squamosals (PIN, nos. 1043/124, 342, 343, 950, 998, 999, 1084–1088); five complete and fragmentary quadratojugals (PIN, nos. 1043/347, 351, 357, 972, 973); four postparietals (PIN, nos. 1043/195, 869, 951, 1091); 27 complete and fragmentary tabulars (PIN, nos. 1043/52, 92, 94, 95, 96, 115–120, 303, 758, 793, 806–808, 903, 904, 952–955, 957–959, 997); five vomer fragments (PIN, nos. 1043/674, 891, 892, 1004, 1098); six complete and fragmentary palatines (PIN, nos. 1043/123, 775, 877, 878, 915, 1099); nine skull base fragments (PIN, nos. 1043/77, 655, 672, 673, 875, 893, 987, 988, 990); eight complete and fragmentary pterygoids (PIN, nos. 1043/101, 121, 792, 923, 949, 991–993); eight complete and fragmentary parasphenoids (specimen PIN, nos. 1043/107, 109, 390, 675, 872–874, 906); nine complete and fragmentary exoccipitals (PIN, nos. 1043/69, 70, 76, 93, 97, 98, 127, 501, 989); and a fragmentary right stapes (PIN, no. 1043/678).
Amphibia - Temnospondyli - Rhytidosteidae
Rhytidosteidae indet. Huene 1920
Novikov 2010
Amphibia - Temnospondyli - Tupilakosauridae
Batrachosuchoides sp. Shishkin 1967
Shishkin 1967
Parasemionotiformes - Parasemionotidae
Watsonulus sp. Brough 1939
Novikov 2010
Saurichthyiformes - Saurichthyidae
Saurichthys sp. Agassiz 1834
Novikov 2010
Chondrichthyes - Hybodontiformes - Hybodontidae
Hybodus maximi
Novikov 2010
Chondrichthyes - Hybodontiformes
Lissodus triaktis
Novikov 2010
Lissodus angulatus (Stensiö 1921)
Novikov 2010
see common names

Geography
Country:Russian Federation State/province:Volgograd County:Ilovlya
Coordinates: 49.3° North, 43.7° East (view map)
Paleocoordinates:25.8° North, 40.7° East
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Triassic Epoch:Early/Lower Triassic
Stage:Olenekian 10 m.y. bin:Triassic 1
*Period:Early/Lower Triassic *Epoch:Scythian
*Local age/stage:Baskunchak
Key time interval:Olenekian
Age range of interval:251.20000 - 247.20000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Lipovskaya
Stratigraphic resolution:bed
Stratigraphy comments: Lower Triassic, equivalent of Yarengian Gorizont, Lipovskaya Svita (Novikov, 1991, Paleont. Zh. 1991(2)).
Lipovskaya Svita, Gamian Gorizont, Yarengian Supergorizont, Upper Olenekian Substage, Olenekian Stage, Lower Triassic (Arkhangelskii & Sennikov, 2008).
The Yarengian is uppermost Olenekian according to fig. 2 in Tverdokhlebov et al. (2003, Earth Sci. Rev. 60).
Chronostratigraphy originally entered as >>Early/Lower Triassic<< is rendered more precisely according to data given above.
Lithology and environment
Primary lithology: not reported
Environment:terrestrial indet. Tectonic setting:cratonic basin
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Articulated whole bodies:none
Disassociated minor elements:all
Collection methods and comments
Collection methods:field collection
Reason for describing collection:taxonomic analysis
Museum repositories:PIN
Metadata
Also known as:Lipovaya Balka
Database number:27343
Authorizer:J. Alroy, J. Mueller, R. Butler, E. Dunne, P. Mannion Enterer:R. Whatley, E. Dunne, R. Butler, T. Liebrecht, J. Tennant
Modifier:R. Butler Research group:vertebrate
Created:2002-12-02 13:54:08 Last modified:2011-03-10 16:23:08
Access level:the public Released:2002-12-02 13:54:08
Creative Commons license:CC BY
Reference information

Primary reference:
7550.5% 37780M. F. Ivakhnenko. 1973. New Cisuralian cotylosaurs. Paleontological Journal 7(2):247-249 [J. Alroy/R. Whatley/T. Liebrecht]

Secondary references:

28874 M. S. Arkhangelskii and A. G. Sennikov. 2008. Podklass Synaptosauria. In M. F. Ivakhnenko and E. N. Kurotchkin (eds.), Piskopaemye pozvonotchnye Rossii i sopredel'nykh stran: Piskopaemye reptilii i ptitsy. Tchast' 1. [Fossil vertebrates of Russia and adjacent countries: Fossil reptiles and birds. Part 1]. GEOS. Moscow 224-243 [J. Mueller/T. Liebrecht/T. Liebrecht]
31386 P. K. Chudinov and B. P. Vjushkov. 1956. Novye Dannye o Melkikh Kotilosavrakh iz Permi i Triasa SSSR [New Data on Small Cotylosaurs from the Permian and Triassic of the USSR]. Doklady Akademii Nauk SSSR 108(3):547-550 [J. Mueller/T. Liebrecht]
28971 M. F. Ivakhnenko. 2008. Podklass Theromorpha. In M. F. Ivakhnenko and E. N. Kurochkin (eds.), Iskopaemye pozvonotchnye Rossii i sopredel'nykh stran: Iskopaemye reptilii i ptitsy, Tchast' 1 [Fossil vertebrates of Russia and adjacent countries: Fossil reptiles and birds, Part 1], GEOS, Moscow 101-183 [J. Mueller/T. Liebrecht/T. Liebrecht]
45793 I. V. Novikov. 2010. New data on trematosauroid labyrinthodonts of Eastern Europe: 2. Trematosaurus galae sp. nov.: cranial morphology. Paleontological Journal 44(4):94-104 [P. Mannion/J. Tennant]
46393 A. G. Sennikov. 1999. The evolution of the postcranial skeleton in archosaurs in connection with new finds of the Rauisuchidae in the Early Triassic of Russia. Paleontological Journal 33(6):648-659 [R. Butler/R. Butler]
46396 A. G. Sennikov. 2001. Discovery of a primitive sauropterygian from the Lower Triassic of the Donskaya Luka (Don Basin) and the Range of Triassic marine reptiles in Russia. Paleontological Journal 35(3):301-309 [R. Butler/R. Butler]
35501 A. G. Sennikov. 2011. New tanystropheids (Reptilia: Archosauromorpha) from the Triassic of Europe. Paleontological Journal 45(1):90-104 [R. Butler/R. Butler/R. Butler]
45855 A. G. Sennikov. 2012. The first ctenosauriscid (Reptilia: Archosauromorpha) from the Lower Triassic of Eastern Europe. Paleontological Journal 46(5):499-511 [P. Mannion/J. Tennant]
84793 A. G. Sennikov. 2022. A New Pseudosuchian from the Early Triassic of Eastern Europe. Paleontological Journal 56(11):1391-1418 [E. Dunne/E. Dunne]
46394 M. A. Shishkin. 1967. A brachyopid labyrinthodont from the Triassic of the Russian Platform. International Geology Review 9:310-322 [R. Butler/R. Butler]
63910 M. A. Shishkin, I. V. Novikov, and J. Fortuny. 2014. New Bystrowianid Chroniosuchians (Amphibia, Anthracosauromorpha) from the Triassic of Russia and Diversification of Bystrowianidae. Paleontological Journal 48(5):512-522 [R. Butler/R. Butler]