Mahmudia Quarry, Caeracul Mare Hill, sample 114: Bithynian, Romania

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Rhynchonellata - Spiriferinida - Mentzeliidae
Koeveskallina informal sp. A Dagys 1965
Grădinaru and Gaetani 2019 8 specimens
Koeveskallina informal sp. B Dagys 1965
Grădinaru and Gaetani 2019 2 specimens
Ptychomentzelia dobrogeana n. sp. Gaetani 2019
Grădinaru and Gaetani 2019 63 specimens
Ptychomentzelia simionescui n. sp. Gaetani 2019
Grădinaru and Gaetani 2019 10 specimens
Rhynchonellata - Spiriferinida - Pennospiriferinidae
Punctospirella sp.
Grădinaru and Gaetani 2019 1 specimen
Rhynchonellata - Spiriferinida - Balatonospiridae
? Dinarispira sp. Dagys 1974
Grădinaru and Gaetani 2019 4 specimens
Rhynchonellata - Terebratulida - Angustothyrididae
Rhaetina sp. Waagen 1882
Grădinaru and Gaetani 2019 2 specimens
Rhynchonellata - Terebratulida - Dielasmatidae
Adygella sp. Dagys 1959
Grădinaru and Gaetani 2019 2 specimens
Sulcatinella aff. incrassata (Bittner 1890)
Grădinaru and Gaetani 2019 43 specimens
Rhynchonellata - Rhynchonellida - Norellidae
Piarorhynchella kittli n. sp. Gaetani 2019
Grădinaru and Gaetani 2019 12 specimens
Rhynchonellata - Rhynchonellida - Cyclothyrididae
Costirhynchopsis informal sp. B Dagis 1977
Grădinaru and Gaetani 2019 1 specimen
Costirhynchopsis informal sp. C Dagis 1977
Grădinaru and Gaetani 2019 2 specimens
Gastropoda - Cycloneritida - Delphinulopsidae
Marmolatella iordanae n. sp. Nützel et al. 2018
Nützel et al. 2018 10 specimens
Marmolatella gigas (Reis 1926)
Nützel et al. 2018 6 specimens
Gastropoda - Cycloneritida - Scalaneritinidae
Ladinaticella simionescui n. sp. Nützel et al. 2018
Nützel et al. 2018 1 specimen
Gastropoda - Coelostylinidae
? Omphaloptycha sp. Ammon 1878
Nützel et al. 2018 9 specimens
Coelostylina micropunctata n. sp. Nützel et al. 2018
Nützel et al. 2018 2 specimens
Gastropoda - Murchisoniina - Plethospiridae
Murchisonietta acuta n. gen., n. sp. Nützel et al. 2018
Nützel et al. 2018 1 specimen
recombined as Wortheniopsis acuta
Gastropoda - Murchisoniina - Zygopleuridae
? Polygyrina peracuta (Assmann 1937)
Nützel et al. 2018 11 specimens
Gastropoda - Murchisoniina - Wortheniellidae
Worthenia (Humiliworthenia) microstriata n. sp. Nützel et al. 2018
Nützel et al. 2018 6 specimens
recombined as Dimorphotomaria microstriata
Gastropoda - Murchisoniina - Rhaphistomellidae
Sisenna sp. Koken 1896
Nützel et al. 2018 1 specimen
Sisenna inaequistriata n. sp. Nützel et al. 2018
Nützel et al. 2018 4 specimens
Gastropoda - Murchisoniina - Phymatopleuridae
Worthenia (Humiliworthenia) anisica n. sp. Nützel et al. 2018
Nützel et al. 2018 4 specimens
? Worthenia (Worthenia) sp. de Koninck 1883
Nützel et al. 2018 1 specimen
Worthenia (Worthenia) dobrogeana n. sp. Nützel et al. 2018
Nützel et al. 2018 3 specimens
Euryalox applanatus n. sp. Nützel et al. 2018
Nützel et al. 2018 9 specimens
Gastropoda
Vetigastropoda indet. Salvini-Plawen 1980
Nützel et al. 2018 5 specimens
Gastropoda - Pleurotomariidae
Rasatomaria multistriata n. sp. Nützel et al. 2018
Nützel et al. 2018 12 specimens
Gastropoda - Turbinidae
Turbinidae indet. Rafinesque 1815
Nützel et al. 2018 1 specimen
Gastropoda - Naticopsidae
Naticopsis sp. M'Coy 1844
Nützel et al. 2018 32 specimens
Gastropoda - Neritidae
? Protonerita cf. comensis (Hoernes 1856)
Nützel et al. 2018 1 specimen
Bivalvia - Arcida - Parallelodontidae
? Parallelodon sp. Meek and Worthen 1866
Friesenbichler et al. 2019 1 specimen
Bivalvia - Ostreida - Posidoniidae
Bositra wengensis (Wissmann 1841)
Friesenbichler et al. 2019 1 specimen
Bivalvia - Ostreida - Pteriidae
Pteria sp. Scopoli 1777
Friesenbichler et al. 2019 3 specimens
Pteria sturi (Bittner 1895)
Friesenbichler et al. 2019 4 specimens
Pteria cf. cassiana (Bittner 1895)
Friesenbichler et al. 2019 1 specimen
Bivalvia - Ostreida - Bakevelliidae
Bakevelliidae indet. King 1850
Friesenbichler et al. 2019 1 specimen
Cultriopsis sp. Cossmann 1904
Friesenbichler et al. 2019 1 specimen
Gervillaria subelegans (Chen 1976)
Friesenbichler et al. 2019 3 specimens
Bivalvia - Ostreida - Pinnidae
Atrina multicostata n. sp. Friesenbichler et al. 2019
Friesenbichler et al. 2019 1 specimen
LPB IIIL 2005 (holotype), left valve
Pinna cf. raibliana Parona 1889
Friesenbichler et al. 2019 1 specimen
Pinna simionescui n. sp. Friesenbichler et al. 2019
Friesenbichler et al. 2019 1 specimen
LPB IIIL 2006 (holotype), right valve
Bivalvia - Ostreida - Ostreidae
? Umbrostrea sp. Hautmann 2001
Friesenbichler et al. 2019 1 specimen
Bivalvia - Pectinida - Pectinidae
Pectinidae indet. Wilkes 1810
Friesenbichler et al. 2019 2 specimens
Chlamys (Praechlamys) prima n. sp. Friesenbichler et al. 2019
Friesenbichler et al. 2019 16 specimens
LPB IIIL 2062 (holotype), fragmented but well preserved left valve
Avichlamys cf. nicolensis (Ogilvie Gordon 1927)
Friesenbichler et al. 2019 1 specimen
Pleuronectites cf. balatonicus (Bittner 1901)
Friesenbichler et al. 2019 2 specimens
"Pecten" volaris Bittner 1902
Friesenbichler et al. 2019 2 specimens
Bivalvia - Pectinida - Aviculopectinidae
Aviculopectinoidea indet. Meek and Hayden 1864
Friesenbichler et al. 2019 3 specimens
    = Globodiscus kiliani n. gen., n. sp. Hautmann et al. 2021
Hautmann et al. 2021
Ornithopecten interruptus (Bittner 1902)
Friesenbichler et al. 2019 1 specimen
Ornithopecten cf. katzeri (Bittner 1902)
Friesenbichler et al. 2019 1 specimen
Amphijanira aff. landrana (Bittner 1895)
Friesenbichler et al. 2019 3 specimens
Neomorphotis comta (Goldfuss 1863)
Friesenbichler et al. 2019 31 specimens
Bivalvia - Pectinida - Hunanopectinidae
Leptochondria pervulgata (Bittner 1902)
Friesenbichler et al. 2019 17 specimens
Leptochondria viezzenensis (Wilckens 1909)
Friesenbichler et al. 2019 5 specimens
Leptochondria separata (Reis 1926)
Friesenbichler et al. 2019 5 specimens
? Asoella sp. Tokuyama 1959
Friesenbichler et al. 2019 1 specimen
Bivalvia - Pectinida - Limidae
Mysidioptera sp. Salomon 1895
Friesenbichler et al. 2019 1 specimen
Mysidioptera fornicata Bittner 1895
Friesenbichler et al. 2019 17 specimens
Plagiostoma striatum (Schlotheim 1820)
Friesenbichler et al. 2019 88 specimens
Plagiostoma cf. angulatum (Münster 1841)
Friesenbichler et al. 2019 3 specimens
Bivalvia - Pectinida - Monotidae
Monotidae indet. Fischer 1887
Friesenbichler et al. 2019 1 specimen
Bivalvia - Pectinida - Entoliidae
Entolium sp. (Meek 1865)
Friesenbichler et al. 2019 2 specimens
Entolium reticulatum n. sp. Friesenbichler et al. 2019
Friesenbichler et al. 2019 2 specimens
LPB IIIL 2070 (holotype), left valve
Scythentolium anisicum n. sp. Friesenbichler et al. 2019
Friesenbichler et al. 2019 17 specimens
LPB IIIL 2075 (holotype), articulated specimen
Entolioides deeckei (Parona 1889)
Friesenbichler et al. 2019 4 specimens
Bivalvia - Myalinida - Mysidiellidae
Mytiliconcha cf. orobica Tommasi 1911
Friesenbichler et al. 2019 6 specimens
Bivalvia - Mytilida - Mytilidae
Modiolus sp. (Lamarck 1801)
Friesenbichler et al. 2019 3 specimens
Bivalvia - Anomalodesmata
Anomalodesmata indet. (Dall 1889)
Friesenbichler et al. 2019 3 specimens
Bivalvia - Trigoniida - Myophoriidae
Elegantinia elegans (Dunker 1849)
Friesenbichler et al. 2019 1 specimen
? Heminajas sp. Wöhrmann 1891
Friesenbichler et al. 2019 1 specimen
Bivalvia - Lucinida - Lucinidae
Schafhaeutlia ? trigona (Stoppani 1858)
Friesenbichler et al. 2019 2 specimens
Schafhaeutlia astartiformis (Münster 1841)
Friesenbichler et al. 2019 6 specimens
recombined as Fimbria astartiformis
Bivalvia - Lucinida - Mactromyidae
Unicardium schmidii (Geinitz 1842)
Friesenbichler et al. 2019 12 specimens
Bivalvia - Pholadomyida - Megadesmidae
Romaniamya mahmudiaensis n. gen., n. sp. Friesenbichler et al. 2019
Friesenbichler et al. 2019 3 specimens
LPB IIIL 2145 (holotype), left valve
Bivalvia - Cardiida - Kalenteridae
Myoconcha sp. Sowerby 1824
Friesenbichler et al. 2019 1 specimen
Joannina aff. joannae Waagen 1907
Friesenbichler et al. 2019 1 specimen
Bivalvia - Megalodontida - Dicerocardiidae
Praedicerocardium vetulus n. gen., n. sp. Friesenbichler et al. 2019
Friesenbichler et al. 2019 4 specimens
LPB IIIL 2142 (holotype), articulated specimen
Cephalopoda - Phylloceratida - Ussuritidae
Ussurites aff. arthaberi (Welter 1915)
Cephalopoda - Ceratitida - Longobarditidae
Longobarditoides solimani (Toula 1896)
Cephalopoda - Ceratitida - Gymnitidae
Caucasites inflatus Shevyrev 1968
Gymnites aff. tozeri Bucher 1992
Cephalopoda - Ceratitida - Khvalynitidae
Alanites aff. visendus Shevyrev 1968
Cephalopoda - Ceratitida - Megaphyllitidae
Megaphyllites prometheus Shevyrev 1961
Cephalopoda - Ceratitida - Acrochordiceratidae
Acrochordiceras hyatti Meek 1877
Cephalopoda - Ceratitida - Ceratitidae
Hollandites silberlingi Bucher 1992
Cephalopoda - Ceratitida - Balatonitidae
Robinsonites sp. Shevyrev 1995
Ostracoda - Myodocopida - Polycopidae
Polycope sp.
Ostracoda - Myodocopida - Cypridinidae
Palaeocypridina spp.
? Triadogigantocypris spp.
Ostracoda - Halocyprida - Thaumatocyprididae
Thaumatocyprididae indet. Muller 1906
Ostracoda - Podocopida - Cytherissinellidae
? Tscherdynzeviana sp.
Ostracoda - Podocopida - Berounellidae
Paraberounella sp.
Ostracoda - Podocopida - Bythocytheridae
? Pseudomonoceratina sp.
Praebythoceratina sp.
Ostracoda - Podocopida - Cytherideidae
Basslerella sp. Kellett 1935
Ostracoda - Podocopida - Microcheilinellidae
Microcheilinella spp.
Microcheilinella contrariella n. sp. Forel 2017
Microcheilinella cf. egerensis Forel 2013
Ostracoda - Podocopida - Healdiidae
Hungarella sp.
Hungarella cf. ovata Zheng 1976
Hungarella mahmudiaensis n. sp. Forel 2017
Ostracoda - Podocopida - Bairdiidae
Bairdia (Urobairdia) angusta (Kollmann 1963)
ssp. recta
? Petasobairdia sp. Chen 1982
Petasobairdia crassa (Kristan-Tollmann 1970)
Bairdia spp.
Bairdia monostorii Forel 2017
Bairdia mandruella n. sp. Forel 2017
Bairdia liviae n. sp. Forel 2017
Bairdia finalyi Mehes 1911
Bairdiacypris spp.
Bairdiacypris triassica Kozur 1971
Bairdiacypris anisica Kozur 1971
Bairdiacypris cf. mirabilis Guan 1985
Kempfina tineriella n. sp. Forel 2017
Liuzhinia bithynica n. sp. Forel 2017
Liuzhinia parva Wei 1981
Liuzhinia edvardgriegi n. sp. Forel 2017
Liuzhinia larmae Forel 2011
Liuzhinia antalyaformis n. sp. Forel 2017
Ostracoda - Podocopida - Bythocyprididae
? Bythocypris sp. Brady 1880
Ostracoda - Podocopida - Acratiidae
Acratia informal sp. 1 Delo 1930
Acratia caeracella n. sp. Forel 2017
Acratia goemeryi (Kozur 1970)
original and current combination Acratia goemoeryi
Ostracoda - Podocopida - Candonidae
Spinocypris vulgaris Kozur 1971
Paracypris spp.
Paracypris oanaella n. sp. Forel 2017
Paracypris cf. aculeata Guan 1985
Triassocypris sp. Kozur 1970
Ostracoda - Podocopida
Renngartennella sp. ()
original and current combination Renngartenella
Ostracoda - Podocopida - Pontocyprididae
? Pontocypris sp.
see common names

Geography
Country:Romania
Coordinates: 45.1° North, 29.1° East (view map)
Paleocoordinates:17.5° North, 33.4° East
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period:Triassic Epoch:Middle Triassic
Stage:Anisian 10 m.y. bin:Triassic 2
Key time interval:Bithynian
Age range of interval:245.50000 - 244.70000 m.y. ago
Stratigraphy
Formation:Caerace Member:Tubiphytes-Limestone
Stratigraphic resolution:bed
Lithology and environment
Primary lithology: lithified bindstone
Environment:reef, buildup or bioherm
Geology comments: The Tubiphytes-boundstones exposed in the north-eastern part of the Mahmudia Quarry are a massive carbonate buildup, lacking any bedding and showing a mixture of different carbonate facies, with frequent packages of microbially-mediated cement crusts.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,microfossils
Collection methods and comments
Collection excludes:some macrofossils
Collection methods:chemical,mechanical,acetic
Reason for describing collection:taxonomic analysis
Museum repositories:BSP
Collection method comments: Repository: CR2P, Université Pierre et Marie Curie, Paris, France (ostracods), BSPG (gastropods)
Metadata
Also known as:Reef 4291
Database number:197722
Authorizer:M. Clapham, P. Wagner, A. Dunhill Enterer:M. Clapham, B. Allen, P. Wagner
Modifier:M. Krause Research group:marine invertebrate
Created:2018-11-23 14:28:49 Last modified:2020-01-12 05:19:53
Access level:the public Released:2018-11-23 14:28:49
Creative Commons license:CC BY
Reference information

Primary reference:
67419. M. B. Forel and E. Gradinaru. 2018. First report of ostracods (Crustacea) associated with Bithynian (Anisian, Middle Triassic) Tubiphytes-microbial reef in the North Dobrogean Orogen (Romania). Papers in Palaeontology 4:211-244 [M. Clapham/M. Clapham]

Secondary references:

70629 E. Friesenbichler, M. Hautmann, E. Grădinaru and H. Bucher. 2019. A highly diverse bivalve fauna from a Bithynian (Anisian, Middle Triassic) Tubiphytes‐microbial buildup in North Dobrogea (Romania). Papers in Palaeontology 7:447-495 [A. Dunhill/B. Allen/M. Clapham]
67834 E. Grădinaru and M. Gaetani. 2019. Upper Spathian to Bithynian (Lower to Middle Triassic) brachiopods from North Dobrogea (Romania). Rivista Italiana di Paleontologia e Stratigrafia 125:91-123 [M. Clapham/M. Clapham]
86429 M. Hautmann, E. Friesenbichler, E. Grădinaru, R. Jattiot, and H. Bucher. 2021. New Triassic Aviculopectinoidea (Bivalvia), with notes on the taxonomic concept of the superfamily. Journal of Paleontology 95(2):291-297 [P. Wagner/P. Wagner]
67421 A. Nützel, A. Kaim, and E. Gradinaru. 2018. Middle Triassic (Anisian, Bithynian) gastropods from North Dobrogea (Romania) and their significance for gastropod recovery from the end-Permian mass extinction event. Papers in Palaeontology 4:477-512 [M. Clapham/M. Clapham]