Kerassia 4 (K4): Late/Upper Miocene, Greece
collected by G. Theodorou Team 1992

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Reptilia - Struthioniformes - Struthionidae
Aves indet. (Linnaeus 1758)
    = Struthio karatheodoris Forsyth Major 1888
Kampouridis et al. 2020
K4: K4.343.22/1 (C16; Figure 3(I))
Reptilia - Phasianidae
Pavo archiaci (Gaudry 1862)
Michailidis et al. 2010
tibiotarsus
Mammalia - Proboscidea - Gomphotheriidae
Tetralophodon cf. longirostris (Kaup 1832)
    = Konobelodon atticus Wagner 1857
Konidaris et al. 2014
Mammalia - Perissodactyla - Rhinocerotidae
? Dicerorhinus cf. pikermiensis Toula 1906
recombined as Dihoplus pikermiensis
Rhinocerotidae indet. Gray 1821
sp. nov.
    = Acerorhinus neleus n. sp. Athanassiou et al. 2014
Athanassiou et al. 2014
Holotype: K4/119.37, skull with articulated mandible (Plates 1, 2). AMPG collection, currently on display at the local Fossil Mammal Museum of Kerassiá (Northern Euboea)
Mammalia - Perissodactyla - Chalicotheriidae
Ancylotherium sp. Gaudry 1863
Mammalia - Perissodactyla - Equidae
Hipparion sp. de Christol 1832
Mammalia - Giraffidae
Palaeotragus sp. Gaudry 1861
Helladotherium cf. duvernoyi Gaudry and Lartet 1856
Mammalia - Bovidae
Gazella sp. Blaineville 1816
Mammalia - Carnivora - Felidae
Machairodus giganteus Wagner 1848
Roussiakis and Theodorou 2003
Mammalia - Carnivora - Hyaenidae
Adcrocuta eximia Roth and Wagner 1854
Roussiakis and Theodorou 2003
see common names

Geography
Country:Greece State/province:Euboea
Coordinates: 38.9° North, 23.3° East (view map)
Paleocoordinates:38.6° North, 23.0° East
Basis of coordinate:estimated from map
Geographic resolution:outcrop
Time
Period:Neogene Epoch:Miocene
10 m.y. bin:Cenozoic 6
Key time interval:Late/Upper Miocene Mammal zone: MN 12
Age range of interval:11.63000 - 5.33300 m.y. ago
Stratigraphy
Stratigraphy comments: Combining the above given data, an MN12 age is plausible for the fauna of Kerassiá, although MN11 cannot be rejected. A biochronological distinction of the two fossiliferous levels found in Kerassiá is not possible for the moment.
Lithology and environment
Primary lithology:burrows,red or brown lithified silty,conglomeratic sandstone
Includes fossils?Y
Lithology description: They belong to the so-called "reddish-brown fluvial deposits" (Mettos et al. 1991) in the lower layers of the upper sequence of the Límni–Istiéa Basin. The lower sequence, as well as the ophiolithic basement, are exposed NE of the village Kerassiá. The superposed reddish-brown fluvial deposits include clays, conglomerates, sands, and siltstones in succession. Their total thickness is about 250–300 m. (brown silted mud with few scattered rounded to subrounded, polymict grains in a homogeneous matrix), the presence of a red-brown layer above the bone bed, and also the presence of the same trace fossils (roots, insect burrows etc.).
Environment:terrestrial indet.
Geology comments: The numerical dominance of bovids and hipparions in the fauna of Kerassiá indicates an open and rather dry environment. Forest elements as cervids and suids are rare or absent. The giraffids, as well as Tragoportax, may have lived in open woodland environments (Gentry et al. 1999). A paleoecological shift between the lower and the upper fossiliferous level is possible due to the small differences in the faunal content, but such an assumption requires more material from both levels.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils
Articulated whole bodies:none
Disassociated major elements:many
Bioerosion:occasional
Spatial resolution:parautochthonous
Collection methods and comments
Collection excludes:some macrofossils
Collection methods:surface (in situ),field collection,survey of museum collection
Reason for describing collection:taxonomic analysis
Museum repositories:AMPG
Collectors:G. Theodorou Team Collection dates:after 1992
Collection method comments: AMPG, Museum of Palaeontology and Geology of the National and Kapodistrian University of Athens
Taxonomic list comments:In this paper (Theodorou et al. 2003) they describe the herbivore fossils. The carnivores are described in another paper.
Metadata
Database number:195435
Authorizer:E. Vlachos Enterer:E. Vlachos, P. Kampouridis
Modifier:P. Kampouridis
Created:2018-08-08 12:45:18 Last modified:2018-08-13 09:27:45
Access level:the public Released:2018-08-08 12:45:18
Creative Commons license:CC BY
Reference information

Primary reference:

66526. G. Theodorou, A. Athanassiou, S. Roussiakis and G. Iliopoulos. 2003. Preliminary remarks on the Late Miocene herbivores of Kerassiá (Northern Euboea, Greece). In J.W.F. Reumer, W. Wessels (eds.), Deinsea, Distribution and Migration of Tertiary Mammals in Eurasia. A volume in honour of Hans de Bruijn 10:519-530 [E. Vlachos/E. Vlachos]

Secondary references:

66528 A. Athanassiou, S. J. Roussiakis, I. X. Giaourtsakis, G. E. Theodorou, and G. Iliopoulos. 2014. A new hornless rhinoceros of the genus Acerorhinus (Perissodactyla: Rhinocerotidae) from the Upper Miocene of Kerassiá (Euboea, Greece), with a revision of related forms. Palaeontographica Abteilung A 303(1-3):23-59 [E. Vlachos/E. Vlachos]
72832 P. Kampouridis, D. Michailidis, N. Kargopoulos, S. Roussiakis, and G. Theodorou. 2020. First description of an ostrich from the late Miocene of Kerassia (Euboea, Greece): remarks on its cervical anatomy. Historical Biology 1-9 [E. Vlachos/E. Vlachos]
69453 G. E. Konidaris, S. J. Roussiakis, G. E. Theodorou and G. D. Koufos. 2014. The Eurasian occurrence of the shovel-tusker Konobelodon (Mammalia, Proboscidea) as illuminated by its presence in the late Miocene of Pikermi (Greece). Journal of Vertebrate Paleontology 34:1437-1453 [E. Vlachos/E. Vlachos]
66564 D. Michailidis, S. Roussiakis, and G. Theodorou. 2010. Palaeoavian remains from the Late Miocene localities of Pikermi, Chomateri and Kerassiá; palaeoecological implications. In A. Chatzipetros, V. Melfos, P. Marchev, I. Lakova (eds.), XIX Congress of the Carpathian-Balkan Geological Association 39(1-2):250-251 [E. Vlachos/P. Kampouridis]
66538 S. J. Roussiakis and G. E. Theodorou. 2003. Carnivora from the Late Miocene of Kerassiá (Northern Euboea, Greece). In J. W. F. Reumer, W. Wessels (eds.), Deinsea. Distribution and Migration of Tertiary Mammals in Eurasia. A Volume in Honour of Hans de Bruijn 10:469-497 [E. Vlachos/P. Kampouridis/P. Kampouridis]