Arapahovius Paleofaunule: Lysitean - Lostcabinian, Wyoming

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Mammalia - Multituberculata - Neoplagiaulacidae
Parectypodus sp. Jepsen 1930
"O"
Mammalia - Leptictidae
Prodiacodon sp. Matthew 1929
presumably includes "Leptictidae" and "Palaeictops" from "O"
Mammalia - Rodentia - Ischyromyidae
Microparamys sp. Wood 1959
"Rodentia" at "O"
Mammalia - Primates - Microsyopidae
Uintasorex sp. Matthew 1909
"Uintasoricinae" at "O"
Mammalia - Primates - Notharctidae
Notharctus sp. Leidy 1870
"Notharctus or Pelycodus" at "O"
Mammalia - Primates - Paromomyidae
Phenacolemur sp. Matthew 1915
    = Phenacolemurinae indet. Simpson 1955
Alroy 2002
Mammalia - Primates - Omomyidae
Tetonoides sp. Gazin 1962
    = Anemorhysis pearcei Gazin 1962
Bown and Rose 1984
original and current combination Tetonoides pearcei
Tetonius sp. Matthew 1915
Arapahovius gazini n. sp. Savage and Waters 1978
"O," "S," Turtle Graveyard (3 measurements)
Loveina sp. Simpson 1940
Mammalia - Erinaceidae
Entomolestes sp. Matthew 1909
    = Erinaceidae indet. Fischer von Waldheim 1817
Alroy 2002
"O"; "Adapisoricinae" also present at "O"
Mammalia - Nyctitheriidae
Leptacodon sp. Matthew and Granger 1921
may include "Nyctitherium" from "O"
Mammalia - Perissodactyla
cf. Heptodon sp. Cope 1882
may include "cf. Homogalax sp." from Lower Patrick Draw
Mammalia - Perissodactyla - Equidae
Hyracotherium sp. Owen 1841
    = Equidae indet. Gray 1821
Alroy 2002
from "O"; Lower Patrick Draw; Upper Patrick Draw
Mammalia - Phenacodontidae
Meniscotherium cf. tapiacitis Cope 1882
    = Meniscotherium chamense Cope 1874
Williamson and Lucas 1992
replaces ID of Gazin 1965b from Upper Patrick Draw; "? Meniscotherium" from "O"
Mammalia - Homacodontidae
Antiacodon sp. Marsh 1872
Mammalia - Creodonta
Creodonta indet. (Cope 1875)
Savage et al. 1972
"O"
Didelphodus sp. Cope 1882
Mammalia - Carnivora - Miacidae
Miacidae indet. Cope 1880
"O"
Mammalia - Condylarthra - Hyopsodontidae
Hyopsodus cf. miticulus (Cope 1874)
    = Hyopsodus sp. Leidy 1870
Gazin 1965
Upper Patrick Draw; includes "H. sp." from "O"
Mammalia - Apatemyidae
Apatemys sp. Marsh 1872
Savage et al. 1972
"O"
Mammalia - Coryphodontidae
Coryphodon sp. (Owen 1845)
original and current combination Lophiodon (Coryphodon)
this might be C. anthracoideus if UCMP V-77011 belongs to the "faunule": Lucas and Schoch 1987; lower Patrick Draw
Mammalia - Groeberida - Nototheriidae
Peradectes sp. Matthew and Granger 1921
presumably includes "Peratherium" from "O"
Reptilia - Testudines
Psilosemys wyomingensis Hutchison 2013
Hutchison 2013
Reptilia - Squamata
Squamata indet. Oppel 1811
Smith and Gauthier 2013 3 specimens
V74022: UCMP 214134 (jaw fragment), 214154, 214157 (?maxilla fragments).
Reptilia
Iguania indet. Cuvier 1817
Smith and Gauthier 2013 2 specimens
V77001: UCMP 165233 (interclavicle fragment), 167742 (partial right prefrontal).
Reptilia - Xantusiidae
Palepidophyma lilliputiana n. sp. Smith and Gauthier 2013
Smith and Gauthier 2013 2 specimens
V74023: UCMP 399755 (holotype), partial right maxilla; Referred specimen: V77001: UCMP 167934 (right maxilla fragment)
Palaeoxantusia amara n. sp. Smith and Gauthier 2013
Smith and Gauthier 2013 67 specimens
V74022: UCMP 165315 (holotype), right spleniodentary; Paratypes: V74022: UCMP 165313 (left dentary), 165317 (left dentary fragment), 167815 (left maxilla fragment), 167816 (right dentary fragment), ?167817 (left maxilla fragment), 214227 (partial right dentary), 214229 (left dentary fragment), 214230 (right dentary fragment), 214244 (left dentary fragment), 214332 (partial left dentary), 214333 (dentary fragment), 214349 (left dentary fragment), 214441 (partial right dentary), 214450 (right dentary fragment), 214569 (right maxilla fragment), 399781 (partial premaxilla). Referred specimens: V70246: UCMP 100068–100099, 100176–100179, 100182–100190 (jaw fragments), 115128 (left maxilla fragment), 165484 (partial left postorbital; Figure 13D), 165485 (left frontal fragment; Figure 13B, C), ?165488 (right pterygoid fragment); V71235: UCMP 214688 (partial left dentary); V77001: UCMP 165265 (left dentary fragment), 167927 (right maxilla fragment; Figure 13A). Most jaw fragments from Level O (100068–100099, 100176– 100179, 100182–100190) were not examined personally, but had been identified prior to this study.
Reptilia
Scincoideus grassator n. sp. Smith and Gauthier 2013
Smith and Gauthier 2013 106 specimens
V74022: UCMP 214231 (holotype), nearly complete right dentary; Paratypes: V74022: UCMP 165316 (partial left maxilla; Figure 15B, C), 165320, 165321 (left dentary fragments), 165323 (right dentary fragment), 165324–165326 (left maxilla fragments), 165327 (right maxilla fragment), 165328, 165329 (maxilla fragments), 165330, 165331 (jaw fragments), 165371, 165376, 165493 (partial premaxillae), 167677 (partial left dentary), 167679 (partial left maxilla), 167712 (nearly complete parietal; Figure 15H, I), 167722, 167724 (left frontal fragments), 167725 (left frontal fragment), 167726 (right frontal fragment), 167807 (right dentary fragment), 167808 (left maxilla fragment), 167818 (right maxilla fragment), 167819 (left maxilla fragment), 167820 (right maxilla fragment), 167821, 167822 (left dentary fragments), 214155 (right dentary fragment), 214194 (partial right frontal), 214220 (right maxilla fragment), 214221, 214228 (left maxilla fragments), 214243 (right dentary fragment), 165245 (left dentary fragment), 214254 (left jugal; Figure 15F, G), 214299, 214330 (partial premaxillae), 214307 (partial right maxilla), 214308 (right maxilla fragment), 214343, 214344 (partial left dentaries), 214345 (right dentary fragment), 214346, 214347 (left dentary fragments), 214348 (right dentary fragment), 214367 (partial right frontal; Figure 15D, E), 214371 (partial premaxilla), 214387, 214388 (left dentary fragments), 214389 (right dentary fragment), 214391 (jaw fragment), 214432 (parietal fragment; Figure 15J), 214436 (partial premaxilla), 214456 (right dentary fragment), ?214472 (jaw fragment), 214482 (left maxilla fragment), 214484–214489 (right maxilla fragments), 214490 (left maxilla fragment), 214564 (right maxilla fragment), 214591 (right dentary fragment), 214592 (left dentary fragment), 214626 (frontal fragment); V74023: UCMP ?165402, ?165403 (jaw fragments), 214652 (partial premaxilla), 399774 (partial premaxilla; Figure 15A); V74024: UCMP 165457 (right dentary fragment), 165458 (jaw fragment). Referred specimens: V70246: UCMP 100051 (right frontal fragment), 100220 (right and left dentary fragments), 100224 (right dentary fragment), 100225 (two left dentary fragments), 100226 (left dentary fragment), 115169, 115170 (left dentary fragments), 165482 (left frontal fragment), 165483 (parietal fragment), 215654 (right dentary fragment); V71232: UCMP 165291 (partial premaxilla), 165292 (partial right maxilla), 165293 (partial right dentary), 165294 (right dentary fragment), 165305 (left dentary fragment); V71235: UCMP 214684 (right maxilla fragment), 214691 (right dentary fragment), 214693 (partial left dentary); V77001: UCMP 165264 (right dentary fragment), 165268 (partial left maxilla), 165269 (left maxilla fragment), 165270 (left dentary fragment), 165272 (right maxilla fragment), 167748 (left jugal fragment), 167917, 167918 (right frontal fragments), 167923 (left frontal fragment), 167933 (partial right dentary), 167973 (partial left maxilla), 167974 (right maxilla fragment), 167976 (right dentary fragment).
Reptilia - Rhineuridae
cf. Jepsibaena sp. Vanzolini 1951
Smith and Gauthier 2013 24 specimens
V70246: UCMP 99855, 215759 (vertebrae); V70251: UCMP 151908 (right dentary; Figure 16A); V74022: UCMP 165364 (right maxilla fragment), 165494 (left maxilla fragment), 167834 (anterior left maxilla fragment), 214151 (right maxilla fragment), 214283, 214284 (three vertebra fragments each), 214285 (two vertebra fragments), 214286 (vertebra fragment), 214290 (partial vertebra), 214414 (vertebra fragment), 214424 (right dentary fragment), 214515, 215530, 215531 (vertebra fragments), 214584 (right maxilla fragment); V74023: UCMP 165405 (partial right dentary), 165406 (left dentary fragment), 165407 (two partial vertebrae), 399768 (left dentary); V74024: UCMP 165470 (four partial vertebrae); V77001: UCMP 165262 (four partial vertebrae), 167967 (three partial vertebrae).
Reptilia - Agamidae
Tinosaurus sp. (Marsh 1872)
Smith and Gauthier 2013 6 specimens
synonym of Saniwa
V70246: UCMP 98237 (right dentary fragment; Figure 9E); V74022: UCMP 165358 (left dentary fragment), ?165416 (frontal fragment; Figure 9C, D), ?165417 (frontal fragment), 214170 (left dentary fragment), 214422 (partial left maxilla; Figure 9A, B).
Reptilia - Iguanidae
Parasauromalus sp. Gilmore 1928
Smith and Gauthier 2013 3 specimens
V70246: UCMP 115106 (partial right dentary, in three pieces; Figure 8B, C), 115108 (jaw fragment); V77001: UCMP 167915 (partial left jugal; Figure 8A).
Paranolis sp. Smith 2011
Smith and Gauthier 2013 7 specimens
V70246: UCMP 115177 (right dentary fragment; Figure 7F), 165473 (partial left postorbital; Figure 7D), 215717 (parietal fragment), 215749 (partial right jugal; Figure 7C); V71235: UCMP 214689 (left dentary fragment); V77001: UCMP 167737 (partial parietal; Figure 7E); V74022: UCMP 167700 (partial frontal; Figure 7A, B).
Suzanniwana sp. Smith 2009
Smith and Gauthier 2013 91 specimens
V70246: UCMP 85683 (jaw fragment), 85687 (right dentary fragment, jaw fragment), 85695 (left dentary fragment), 85697 (partial right maxilla; Figure 3B, C), 85704 (left maxilla fragment), 85716 (right dentary fragment), 85747 (nearly complete right dentary; Figure 3J), 98227 (partial left dentary), 98228 (nearly complete left dentary; Figure 3I), 99806 (right dentary fragment, maxilla fragment), 100050 (frontal fragment; cf. Gauthier 1982), 100222 (left dentary fragment), 100228 (right dentary fragment), 115031, 115032 (left dentary fragments), 100036, 100038 (right dentary fragments), 115044 (left dentary fragment), 115046 (left dentary fragment), 115049 (right maxilla fragment), 115051, 115055 (right dentary fragments), 115058 (left dentary fragment), 115059 (right dentary fragment), 115068 (partial right mandible), 115072 (right dentary fragment), 115076 (partial left dentary), 115079 (right dentary fragment), 115080 (left dentary fragment), ?115081, 115083 (right dentary fragments), 115085, 115087, 115088 (left dentary fragments), 115089 (right dentary fragment), 115090 (left dentary fragment), 115092 (right dentary fragment), 115095, ?115096, 115097, 115098 (frontal fragments), 115100 (frontal fragment; Figure 3E), 115102 (frontal fragment), 115103 (frontal fragment; Figure 3F), ?115104, 115105 (frontal fragments), 115143 (left maxilla fragment), 115146 (right maxilla fragment), ?115147, 115148 (left maxilla fragments), 115151 (right maxilla fragment), 115152 (jaw fragment), 115154 (partial right maxilla), 115155, 115156 (right maxilla fragments), 115158 (partial left maxilla), 115160 (left maxilla fragment; Figure 3A), 115162 (two left maxilla fragments), 115165 (left maxilla fragment), ?115167 (maxilla fragment), 115171 (left maxilla fragment), 115172, 115173 (right maxilla fragments), 115176 (two maxilla fragments), 115179 (left maxilla fragment), 115181 (right maxilla fragment), 115182 (left maxilla fragment), 115187 (right maxilla fragment), 115189 (two maxilla fragments), 115190, 115191 (partial right dentaries), ?115192 (partial left dentary), 115193 (jaw fragment), 115195 (partial right maxilla), ?115198 (left maxilla fragment), 115203 (two maxilla fragments), ?115204, 115206 (right maxilla fragments), 165480 (partial right postorbital; Figure 3H), 165481 (partial premaxilla), 215657 (left maxilla fragment), ?215681 (right dentary fragment), 215747 (left jugal fragment), 215750 (right jugal fragment), 215752 (left jugal fragment), 215757 (left pterygoid fragment), 215766 (right compound bone fragment), 215767 (left compound bone fragment), 215795 (partial left jugal; Figure 3G), 399769 (left maxilla fragment), 399773 (partial premaxilla).
Suzanniwana revenanta n. sp. Smith and Gauthier 2013
Smith and Gauthier 2013 116 specimens
V74022: UCMP 167682 (holotype), partial right jugal; Paratypes: V74022: UCMP 165309 (partial frontal), 165310 (left dentary fragment), 165332 (partial left maxilla; Figure 4C, D), 165333 (partial left maxilla), 165334 (right maxilla fragment), 165337 (jaw fragment), 165338 (partial right dentary), 165339 (right dentary fragment), 165340 (dentary fragment), 165344 (left maxilla fragment), ?165349 (jaw fragment), ?165378 (right squamosal; Figure 4J), ?167689 (partial left ectopterygoid; Figure 4E), 167692 (partial frontal), 167696 (partial right compound bone), 167699 (left compound bone fragment), 167727 (right pterygoid fragment), 167810 (left maxilla fragment), 167811, 167812 (partial right dentaries), 214131 (partial premaxilla), ?214142 (left maxilla fragment), 214145 (right maxilla fragment), 214164 (right dentary fragment), ?214179 (five caudal vertebrae), 214202 (left maxilla fragment), 214203 (jaw fragment), 214205 (partial left dentary), 214206 (dentary fragment), 214207 (maxilla fragment), 214208 (left dentary fragment), 214209 (maxilla fragment), 214212 (partial right maxilla), 214215 (right maxilla fragment), 214225 (right dentary fragment), 214226, 214234 (partial left dentaries), 214248 (anterior frontal fragment; Figure 4G), 214251 (frontal fragment; Figure 4H), 214295 (partial premaxilla), ?214313 (right maxilla fragment), 214322 (partial left maxilla), 214323 (left maxilla fragment), 214324 (right maxilla fragment), 214330, 214339 (right dentary fragment), 214341 (left dentary fragment), 214352 (right dentary fragment), 214353, 214354 (jaw fragments), 214361 (left dentary fragment), 214383 (right dentary fragment), ?214392 (jaw fragment), 214397–214399 (partial left dentaries), 214439 (partial premaxilla), 214447 (left dentary fragment), 214449, 214451, 214464 (right dentary fragments), 214465 (partial left dentary), 214468 (left dentary fragment), 214494, 214495, 214500, 214501 (right maxilla fragments), 214568 (right maxilla fragment), 214572 (right maxilla fragment), 214573 (partial right maxilla), 214599 (left dentary fragment), 214613 (partial right dentary), 214630 (frontal fragment), 217909 (partial premaxilla; Figure 4B) 399778 (left mandible fragment), 399780 (partial premaxilla); V74023: UCMP 165382 (partial left compound bone; Figure 4K), 165390 (right maxilla fragment), 165394 (partial premaxilla), 165395 (left mandible fragment), 165396 (left maxilla fragment), 214644, ?214645 (left dentary fragments), 214665 (dentary fragment), 214666 (partial right dentary), ?214667 (right maxilla fragment), 399758 (partial frontal), 399759 (partial left dentary), 399762 (left maxilla fragment), 399763 (left dentary fragment), 399765 (right dentary fragment), 399766 (partial premaxilla); V74024: UCMP 165424 (left jugal fragment), ?165451 (left dentary fragment), 165452 (right maxilla fragment), 165459 (partial left maxilla), ?165460 (dentary fragment), 165461 (partial left dentary), 165462 (partial right dentary), 165463 (right dentary fragment), 165467 (right maxilla fragment). Referred specimens: V71232: UCMP 165287 (partial left maxilla); V71234: UCMP 214679 (right dentary fragment), ?214682 (caudal vertebra); V71235: UCMP 214690 (right dentary fragment), 214692 (dentary fragment); V77001: UCMP 165245 (partial premaxilla), 165246 (partial premaxilla; Figure 4A), 165251 (frontal fragment), ?165252 (caudal vertebra; Figure 4L), 167741 (partial left prefrontal; Figure 4F), 167749 (left pterygoid fragment), 167916 (right jugal fragment), 167946 (right dentary fragment), ?167962 (caudal vertebra).
Anolbanolis sp. Smith 2009
Smith and Gauthier 2013 12 specimens
V70246: UCMP 115050 (partial left dentary), 165478 (partial parietal; Figure 5E), 215748 (partial right jugal), 399772 (left dentary fragment; Figure 5F); V74022: UCMP 165309 (partial frontal; Figure 5A), 167683 (partial left jugal; Figure 5B), ?167813 (right maxilla fragment), 214249, 214250 (frontal fragments), 214310 (right maxilla fragment), 214400 (left dentary fragment); V77001: UCMP 167743 (partial left postorbital; Figure 5C, D)
Reptilia - Helodermatidae
Helodermatidae indet. Gray 1837
Smith and Gauthier 2013 1 specimen
V77001: UCMP 167922 (osteoderm)
Reptilia
cf. Saniwa informal sp. A Leidy 1870
Smith and Gauthier 2013 1 specimen
V74022: UCMP 214172 (jaw fragment).
Reptilia - Anguidae
Anguidae indet. Gray 1825
Smith and Gauthier 2013 18 specimens
"small anguid specimens" V70246: UCMP 165475 (partial parietal), 165490 (right jugal fragment), 214255 (parietal fragment), 214432 (partial parietal), 215753 (partial left jugal), 215796 (partial caudal vertebra; Figure 18F); V74022: UCMP 167681 (partial right jugal), 167707–167711 (parietal fragments), 167890 (partial proximal caudal vertebra), 214193, 214195 (parietal fragments), 399779 (partial premaxilla; Figure 18A); V74023: UCMP 165381 (partial right jugal; Figure 18B); V77001: UCMP 167736 (parietal fragment).
Apodosauriscus minutus n. gen., n. sp. Gauthier 1982
Gauthier 1982 38 specimens
UCMP 100125 (holotype), posterior half of left dentary; UCMP 100056, 100057, 100059, isolated body osteoderms; UCMP 100103, posterior one-third of right frontal; UCMP 1001 12, fragment of right dentary; UCMP 100118, posterior twothirds of left dentary; UCMP 100126, fragment of left dentary; UCMP 100128, fragment of posterior end of right dentary; UCMP 100138, fragment from anterior end of right maxilla; UCMP 100139, posterior process of left maxilla; UCMP 100145, fragment from posterior end of left dentary; UCMP 100149, fragment from anterior end of right dentary; UCMP 100154, right maxilla broken anteriorly and posteriorly; UCMP 100155, anterior onehalf of left dentary; UCMP 100157, fragment of left dentary; UCMP 100158, fragment from anterior and left dentary; UCMP 116548, isolated body osteoderm. Newly referred specimens from ref. 60296: V70246: UCMP ?100113 (right maxilla fragment), ?100136 (jaw fragment), 100147 (left maxilla fragment), 100151 (left maxilla fragment), 165489 (partial right compound bone; Figure 17E, F), 215660 (left dentary fragment), 215671 (right dentary fragment), 399750 (partial left dentary). V74022: UCMP 165365 (maxilla fragment), 167713 (right frontal fragment), 167721 (left frontal fragment), 214213 (premaxilla; Figure 17A, B), ?214406 (frontal fragment), ?214458 (jaw fragment), 214554 (right maxilla fragment); V74023: UCMP 399760 (left maxilla fragment); V74024: UCMP 165456 (partial right maxilla); V77001: UCMP 167286 (partial premaxilla), 167936 (partial right dentary), 399761 (left coronoid).
Eodiploglossus borealis n. gen., n. sp. Gauthier 1982
Gauthier 1982 48 specimens
UCMP 100100 (holotype), posterior two-thirds of right dentary; UCMP 100044, fragment from anterior half of left maxilla; UCMP 100058, isolated body osteoderms; UCMP 100104, right frontal missing parts from anterior and posterior ends; UCMP 100105, right frontal missing parts from anteromedial and posterolateral ends; UCMP 100106, posterior half of right frontal; UCMP 100123, posterior process of right maxilla; UCMP 100124, fragment left maxilla; UCMP 100129, posterior process of right maxilla; UCMP 100134, fragment right maxilla; UCMP 100135, posterior process of right maxilla; UCMP 100136, tooth-bearing fragment; UCMP 100143, fragment right maxilla; UCMP 100153, fragment left maxilla; UCMP 100156, toothbearing fragment; UCMP 116547, body osteoderm; UCMP 116549, body osteoderm; UCMP 116550, body osteoderm. New referred specimens from ref. 60926: V70246: UCMP 100132 (jaw fragment), 100140 (left maxilla fragment), 100141 (right maxilla fragment), 100148 (left maxilla fragment), 100152 (jaw fragment), ?215668 (right dentary fragment). V74022: UCMP 167717 (right frontal fragment), 167718 (left frontal fragment), ?167719 (anterior left frontal fragment), 167723 (left frontal fragment), 167823 (right dentary fragment), ?214218 (right maxilla fragment), 214316, 214380 (jaw fragment), 214430 (osteoderms), 214444 (anterior left dentary fragment), 214460 (left maxilla fragment), 214461 (jaw fragment), 214463 (left dentary fragment), 214508 (osteoderms), 214607 (right maxilla fragment), 399798 (left maxilla fragment), 399799 (right maxilla fragment); V74023: UCMP 165401 (left dentary fragment), 214656 (left maxilla fragment); V77001: UCMP 165275 (left dentary fragment), 165429 (osteoderm), 167938 (partial left maxilla), 167939 (right maxilla fragment), 167953 (osteoderm), 167975 (partial right dentary).
Glyptosaurinae informal indet A
Gauthier 1982 4 specimens
UCMP 100161, posterior process of right maxilla; UCMP 100168, tooth-bearing fragment; UCMP 100172, posterior end of left dentary. Newly referred by ref. 60296: UCMP 116450 (two isolated teeth).
Glyptosaurinae informal indet B
Gauthier 1982 7 specimens
UCMP 100162, fragment from posterior end of right dentary; UCMP 100174, anterior two-thirds of right dentary; UCMP 100327, fragment from anterior end of right maxilla. Newly referred from ref. 60296: V70246: UCMP 115127 (right maxilla fragment), 116451 (two isolated teeth); V71232: UCMP 165299 (right maxilla fragment); V71236: UCMP ?151943 (skull fragment).
Glyptosaurinae informal osteoderms
Gauthier 1982
Isolated osteoderms, UCMP 85719, 85737, 85981, 96614, 99829, 99844, 99845, 99857, 99873, 100060, 100061, 100062, 100063, 100064, 100065, 100067. "Approximately 900 isolated osteoderms are clearly referrable to the Glyptosaurinae".
Xestops savagei Smith and Gauthier 2013
Smith and Gauthier 2013 26 specimens
V70246 (cf. Gauthier 1982): UCMP 100101 (right frontal fragment), 100102 (partial left frontal; Gauthier 1982, fig. 16a, b), 100110 (left pterygoid fragment), 100166 (left dentary fragment), 100169 (right maxilla fragment), ?100170 (jaw fragment), 100171 (partial right maxilla; Gauthier 1982, fig. 22; Figure 23A), ?100172 (left dentary fragment; cf. Gauthier 1982), 100173 (right maxilla fragment), 100175 (right maxilla fragment), 100165 (jaw fragment), 116440–116445 (osteoderms; see Gauthier 1982), 116446 (two jaw fragments), 116447–116449 (osteoderms), ?215715 (right jugal fragment); V71232: UCMP ?165302 (osteoderm); V74022: UCMP ?165492 (left frontal fragment), ?214211 (right maxilla fragment), 214628 (partial right frontal; Figure 19B, C).
cf. Proxestops sp. Gauthier 1982
ElShafie 2024
Glyptosaurus agmodon n. sp. Smith and Gauthier 2013
Smith and Gauthier 2013 4 specimens
V74022: UCMP 179384 (holotype), partial right maxilla; Paratypes: V74022: 214338 (jaw fragment), 214627 (partial right frontal; Figure 22D–F), 399782 (left maxilla fragment).
Arpadosaurus sepulchralis n. sp. Smith and Gauthier 2013
Smith and Gauthier 2013 4 specimens
V74022: UCMP 214431 (holotype), partial right frontal; Paratypes: V74022: UCMP 214586 (partial dentary), 214675 (left dentary; Figures 20C–E, 22C); V74024: UCMP ?151761 (partial ?left ?maxilla).
Gerrhonotinae indet. (Cope 1864)
Gauthier 1982 24 specimens
UCMP 100115, right dentary missing anterior and posterior ends; UCMP 100119 and 100131, fragments of two right dentaries. New referrals from ref. 60296: V70246: UCMP 100117 (right dentary fragment), ?100120, 100121 (right maxilla fragments), 100122 (left maxilla fragment), 100127 (right dentary fragment), 100138 (right maxilla fragment); V71232: UCMP 165304 (partial right dentary); V74022: UCMP 165359, 165360, 165361 (left dentary fragments), ?165495, ?214457 (right dentary fragments), 214602 (left dentary fragment); V74023: UCMP 214650 (right dentary fragment); V74024: UCMP 165454, ?165455 (left maxilla fragments); V77001: UCMP 165273 (partial left dentary), 165274 (left dentary fragment), 165276 (right dentary fragment), 165277 (left maxilla fragment), 167941 (left dentary fragment).
Reptilia
cf. Restes sp. Gauthier 1982
Smith and Gauthier 2013 1 specimen
V77001: UCMP 167940 (right maxilla fragment)
Entomophontes incrustatus n. gen., n. sp. Smith and Gauthier 2013
Smith and Gauthier 2013 65 specimens
V70246: UCMP 98223 (holotype), right dentary; Paratypes: V70246: UCMP 85783 (partial premaxilla; Figure 24A, B), 100031 (skull fragment), 100032 (partial parietal; Figure 25B, C), 100033 (left frontal fragment; Figure 24G), 100034 (right frontal fragment; Figure 24H, I), 100035 (frontal fragment), 100036 (15 osteoderms; example: Figure 25J), 100037 (13 miscellaneous teeth and fragments), 100038 (right dentary fragment), 100040 (right dentary fragment), 100041–100043, 100045, 100046 (left maxilla fragments), 100047, 100048 (right maxilla fragments), 100055 (two cranial osteoderms), 100107, 100108 (parietal fragments). Referred specimens: V70251: UCMP 151808 (heavily corroded right frontal; Figure 24J); V71232: 165296 (premaxilla fragment), 165297 (jaw fragment), 165300 (partial parietal; Figure 25D, E); V71235: UCMP 214694 (jaw fragment); V74022: UCMP 165369 (left dentary fragment), 167705 (partial right maxilla; Figure 24D), 167706 (left maxilla fragment), 167734 (right dentary fragment; Figure 25I), 167780 (left maxilla fragment), 167825 (left dentary fragment), 167827 (right maxilla fragment), 167829 (left dentary fragment), 167896 (?cranial osteoderm), 214135 (right maxilla fragment), 214137 (left maxilla fragment), 214141 (right maxilla fragment), 214200 (partial premaxilla), 214222, 214257 (right maxilla fragments), 214319 (left maxilla fragment), ?214370 (partial premaxilla), 214421 (right maxilla fragment), 214433 (partial ?nasal), 214434 (cranial fragment), 214435 (frontal fragment), 214436 (left dentary fragment), 214440 (right dentary fragment), 214478 (left maxilla fragment), 214609 (left dentary fragment); V74023: UCMP 214649 (left maxilla fragment), 214655 (right dentary fragment); V74024: UCMP 165439 (partial premaxilla), 165443 (partial right maxilla), 165444 (left dentary fragment); V77001: UCMP 165279 (right maxilla fragment), 167735 (parietal fragment), 167738 (partial right frontal), 167921 (parietal fragment), 167924 (maxilla fragment), 167925 (right maxilla fragment), 167926 (partial left maxilla), 167969 (partial left frontal), 399771 (left maxilla fragment; Figure 24C).
Reptilia - Necrosauridae
Provaranosaurus fatuus n. sp. Smith and Gauthier 2013
Smith and Gauthier 2013 80 specimens
V70246: UCMP 98234 (holotype), left dentary; Paratypes: V70246: UCMP 85701 (jaw fragment), 98229 (partial frontal), 99863 (parietal; Figure 29A, B), 100049, 100051, 100053, 100054 (partial frontals; cf. Gauthier 1982), 100120 (right maxilla fragments), 115109 (left dentary fragment), 115110 (right dentary fragment), 115111 (left maxilla fragment), 115112 (eight teeth and jaw fragments), 115113 (partial left dentary; Figure 29G), 115114 (left dentary fragment), 115115 (right maxilla fragment), 115116 (partial left dentary), 115117 (left dentary fragment), 115118 (tooth fragment; Figure 29H), 115119, 115120 (left dentary fragments), 115121, 115122 (right dentary fragments), 115123 (right dentary fragment), 115126 (three maxilla and dentary fragments), 115130 (right maxilla fragment), 115131 (right dentary fragment), 115136 (jaw fragment), 115137 (left dentary fragment), 165474 (partial left frontal), 165476 (three osteoderms; two examples: Figure 29I, J and K, L, respectively), 215716 (partial right prefrontal; Figure 28E), 215791 (parietal fragment), 399752 (right jugal fragment; Figure 28F). Referred specimens: V71232: UCMP 165298 (right dentary fragment); V74022: UCMP 165308 (partial left frontal), 165314 (three osteoderms; example: Figure 29L), 165318 (anterior right maxilla fragment), 165370 (premaxilla; Figure 28A, B), ?165496 (partial left jugal), ?167687, ?167688 (partial right palatines), 167693 (partial left frontal), 167694 (parietal fragment), 214127 (partial premaxilla), 214167 (jaw fragment), 214232 (jaw fragment), 214252 (anterior frontal fragment), ?214253 (partial right jugal), ?214282 (partial vertebra), 214314 (left maxilla fragment), 214318 (jaw fragment), 214334 (left dentary fragment), 214366 (partial left frontal), ?214368 (left frontal fragment), 214425 (two jaw fragments), 214445 (left dentary fragment), 214555 (right maxilla fragment), 214605 (right dentary fragment), 214638 (partial left frontal), 399770 (right maxilla fragment; Figure 28C), 399796 (left maxilla fragment), 399797 (jaw fragment); V74023: UCMP ?165379 (left jugal fragment), 165404 (jaw fragment), 399756 (partial parietal), 399757 (partial frontal); V74024: UCMP 165418 (right frontal fragment), 165419 (left frontal fragment), 165445– 165447 (jaw fragments); V77001: UCMP 165244 (nearly complete frontal; Figure 28I–K), 165278 (left dentary fragment), 165280 (left maxilla fragment), 165281, 165282, ?165283 (jaw fragments), ?167447 (right jugal fragment), 167913 (partial left ectopterygoid; Figure 28D), 167942 (left maxilla fragment).
see common names

Geography
Country:United States State/province:Wyoming County:Sweetwater
Coordinates: 41.7° North, 109.0° West (view map)
Paleocoordinates:46.6° North, 92.6° West
Basis of coordinate:based on political unit
Time
Period:Paleogene Epoch:Eocene
Stage:Ypresian 10 m.y. bin:Cenozoic 2
*Period:Tertiary *Local age/stage:Lostcabinian - Lysitean
Key time interval:Lysitean - Lostcabinian
Age range of interval:55.80000 - 50.30000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Wasatch Member:Main Body
Local section:Washa Local bed:1
Local order:bottom to top
Stratigraphy comments: said to be Lysitean to Lostcabinian
Lithology and environment
Primary lithology: not reported
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:bulk,surface (float),sieve
Reason for describing collection:biostratigraphic analysis
Museum repositories:UCM,UCMP
Collection method comments: "New fossils" collected by the UCM "in 2001-2002" were "recovered through surface collection and screen washing" (Cuozzo 2008)
Taxonomic list comments:records for "Zonule O" of Savage et al. 1972 and for the upper and lower Patrick Draw sites of Gazin 1965b are indicated below
list is preliminary and possibly incomplete
Metadata
Also known as:UCMP V-70246, 70251, 71232 - 236, 77001, 74022 - 024; Oh! Locality; Patrick Draw Zonules "O" and "S"; Turtle Graveyard; UCM L88040
Database number:15451
Authorizer:J. Alroy, P. Holroyd, P. Barrett Enterer:J. Alroy, P. Holroyd, T. Cleary
Modifier:J. Alroy Research group:vertebrate
Created:1993-02-18 00:00:00 Last modified:2010-02-28 16:37:52
Access level:the public Released:1993-02-18 00:00:00
Creative Commons license:CC BY
Reference information

Primary reference:

3018. D. E. Savage and B. T. Waters. 1978. A New Omomyid Primate from the Wasatch Formation of Southern Wyoming. Folia Primatologica 30 [J. Alroy/J. Alroy/J. Ju]

Secondary references:

6294 J. Alroy. 2002. Synonymies and reidentifications of North American fossil mammals. [J. Alroy/J. Alroy/M. Kouvari]
1122 T. M. Bown and K. D. Rose. 1984. Reassessment of Some Early Eocene Omomyidae, with Description of a New Genus and Three New Species. Folia Primatologica 43 [J. Alroy/J. Alroy/J. Ju]
32177 F. P. Cuozzo. 2008. Using extant patterns of dental variation to identify species in the primate fossil record: a case study of middle Eocene Omomys from the Bridger Basin, southwestern Wyoming. Primates 49(2):101-115 [J. Alroy/J. Alroy]
87093 S. J. ElShafie. 2024. Body size estimation from isolated fossil bones reveals deep time evolutionary trends in North American lizards. PLoS ONE 19(1:e0296318):1-25 [P. Holroyd/P. Holroyd/P. Holroyd]
14650 J. A. Gauthier. 1982. Fossil xenosaurid and anguid lizards from the early Eocene Wasatch Formation, southeast Wyoming, and a revision of the Anguidea. Contributions to Geology, University of Wyoming 21(1):7-54 [R. Whatley/R. Whatley/R. Whatley]
1618 C. L. Gazin. 1965. Early Eocene mammalian faunas and their environment in the vicinity of the Rock Springs Uplift, Wyoming. Wyoming Geological Association Guidebook 19:171-180 [J. Alroy/J. Alroy/J. Marcot]
55096 J. H. Hutchison. 2013. New turtles from the Paleogene of North America. In D. B. Brinkman, P. A. Holroyd, J. D. Gardner (eds.), Morphology and Evolution of Turtles 477-497 [P. Holroyd/P. Holroyd]
3017 D. E. Savage, B. T. Waters, and J. H. Hutchison. 1972. in West guidebook [J. Alroy/J. Alroy]
60296 K. T. Smith and J. A. Gauthier. 2013. Early Eocene lizards of the Wasatch Formation near Bitter Creek, Wyoming: diversity and paleoenvironment during an interval of global warming. Bulletin of the Peabody Museum of Natural History 54(2):135-230 [P. Barrett/T. Cleary]
3647 T. E. Williamson and S. G. Lucas. 1992. Meniscotherium (Mammalia, "Condylarthra) from the Paleocene-Eocene of Western North America). New Mex Museum of Natural History Science Bulletin 1:1-75 [J. Alroy/J. Alroy/M. Uhen]