Cochrane 2: Ti1, Canada
collected by G. P. Youzwyshyn 1988

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Amphibia - Temnospondyli - Albanerpetontidae
Albanerpeton informal sp. nov. Estes and Hoffstetter 1976
Gardner and DeMar 2013
Mammalia - Multituberculata - Microcosmodontidae
Acheronodon sp. Archibald 1982
said to be new
Pentacosmodon sp. Jepsen 1940
Mammalia - Multituberculata - Cimolodontidae
Anconodon sp. Jepsen 1940
Anconodon cochranensis (Russell 1929)
Anconodon gidleyi (Simpson 1935)
Mammalia - Multituberculata - Neoplagiaulacidae
Neoplagiaulacidae indet. Ameghino 1890
Parectypodus sp. Jepsen 1930
said to be new
Parectypodus sinclairi (Simpson 1935)
Parectypodus cf. sylviae (Rigby 1980)
Ectypodus sp. Matthew and Granger 1921
Ectypodus cf. szalayi Sloan 1981
Neoplagiaulax donaldorum Scott and Krause 2006
Scott and Krause 2006
Neoplagiaulax nelsoni Sloan 1987
nomen nudum belonging to Neoplagiaulax
Neoplagiaulax cf. hunteri (Simpson 1936)
Mesodma pygmaea Sloan 1987
Mimetodon silberlingi (Simpson 1935)
Mammalia - Multituberculata - Ptilodontidae
Baiotomeus sp. Krause 1987
said to be new (1 measurement)
    = Baiotomeus russelli n. sp. Scott et al. 2002
Scott et al. 2002
Prochetodon speirsae n. sp. Scott 2004
Scott 2004
Ptilodus sp. Cope 1881
two forms "T" and "C," both said to be new
Ptilodus gnomus n. sp. Scott et al. 2002
Scott et al. 2002
(9 measurements)
Mammalia - Multituberculata - Eucosmodontidae
Eucosmodontinae indet. Jepsen 1940
Mammalia - Leptictidae
Prodiacodon concordiarcensis Simpson 1935
Prodiacodon furor Novacek 1977
Prodiacodon cf. puercensis (Matthew 1918)
Myrmecoboides montanensis Gidley 1915
Mammalia - Mixodectidae
Eudaemonema cf. cuspidata Simpson 1935
Mammalia - Mimotonidae
cf. Mimotonidae indet. Li 1977
Mammalia - Plagiomenidae
cf. Thylacaelurus sp. Russell 1954
Mammalia - Primates - Paromomyidae
Paromomys depressidens Gidley 1923
Gunnell 1989
presumably includes "? Palaechthon sp."
Ignacius fremontensis (Gazin 1971)
    = Ignacius glenbowensis Scott et al. 2023
Scott et al. 2023
UALVP specimens
Ignacius cf. frugivorus Matthew and Granger 1921
    = Ignacius glenbowensis Scott et al. 2023
Scott et al. 2023
UALVP specimens
Mammalia - Primates - Palaechthonidae
Phoxomylus puncticuspis n. gen., n. sp. Fox 2011
Fox 2011
(1 measurement)
Plesiolestes sirokyi Szalay 1973
    = Torrejonia cf. sirokyi Szalay 1973
Alroy 2002
Mammalia - Primates - Plesiadapidae
Nannodectes intermedius (Gazin 1971)
? Pronothodectes sp. Gidley 1923
regarded as N. intermedius by Gingerich 1982a, but described by Fox 1990b
Plesiadapis praecursor Gingerich 1975
not clear if this is really present in light of Fox 1990b
Plesiadapis cf. anceps Simpson 1936
Mammalia - Primates - Carpolestidae
Elphidotarsius russelli n. sp. Krause 1978
(5 measurements)
Carpodaptes cf. hazelae Simpson 1936
Mammalia - Primates - Picrodontidae
Picrodus silberlingi Douglass 1908
(7 measurements)
    = Picrodus canpacius n. sp. Scott and Fox 2005
Scott and Fox 2005
Mammalia - Dermoptera
Elpidophorus cf. elegans Simpson 1927
Mammalia - Nyctitheriidae
Nyctitheriidae indet. Simpson 1928
Limaconyssus sp. Gingerich 1987
Leptacodon cf. tener Matthew and Granger 1921
"Leptacodon" munusculum Simpson 1935
Leptacodon cf. packi Jepsen 1930
cf. Nyctitherium sp. Marsh 1872
said to be new
Mammalia - Periptychidae
? Periptychidae indet. Cope 1882
Mammalia - Phenacodontidae
Ectocion collinus Russell 1929
Mammalia - Procreodi - Arctocyonidae
Claenodon montanensis (Gidley 1919)
    = Arctocyon cf. montanensis Gidley 1919
Alroy 2002
synonym of Arctocyon corrugatus
? Mimotricentes sp. Simpson 1937
Colpoclaenus cf. keeferi Patterson and McGrew 1962
? Oxyprimus sp. Van Valen 1978
Thryptacodon orthogonius (Russell 1929)
    = Chriacus orthogonius Russell 1929
Alroy 2002
    = Thryptacodon orthogonius Russell 1929
Scott et al. 2002
Chriacus cf. baldwini (Cope 1882)
Chriacus pelvidens (Cope 1881)
Mammalia - Creodonta
Creodonta indet. (Cope 1875)
Mammalia - Viverravidae
Simpsonictis cf. tenuis (Simpson 1935)
Simpsonictis pegus Gingerich and Winkler 1985
Mammalia - Carnivora
Pristinictis connata n. sp. Fox and Youzwyshyn 1994
Fox and Youzwyshyn 1994
presumably includes new viverravid of Fox (3 measurements)
Mammalia - Acreodi - Oxyclaenidae
? Oxyclaeninae indet. (Scott 1892)
invalid subgroup of Acreodi
said to be new
Mammalia - Acreodi - Mesonychidae
Mesonychidae indet. Cope 1875
Mammalia - Condylarthra
Claenodon sp. Scott 1892
    = Condylarthra indet. Cope 1881
Alroy 2002
Desmatoclaenus sp. Gazin 1941
Mammalia - Macroscelidea - Apheliscidae
Apheliscinae indet. Matthew 1918
said to be new
    = Gingerichia hystrix n. sp. Zack et al. 2005
Zack et al. 2005
UALVP 43082 - type (left p4); numerous referred UALVP material
Litomylus sp. Simpson 1935
said to be new (7 measurements)
    = Litomylus grandaletes n. sp. Scott et al. 2002
Scott et al. 2002
Litomylus dissentaneus Simpson 1935
Mammalia - Macroscelidea
Litocherus sp. Gingerich 1983
Litocherus cf. lacunatus (Gazin 1956)
Mammalia - Proteutheria
Palaeoryctoidea indet. Winge 1917
said to be new
Mammalia - Proteutheria - Palaeoryctidae
Palaeoryctidae indet. Winge 1917
three forms
Palaeoryctes cf. punctatus Van Valen 1966
Didelphodontinae indet. Simpson 1927
Pararyctes sp. Van Valen 1966
said to be new (3 measurements)
    = Pararyctes rutherfordi n. sp. Scott et al. 2002
Scott et al. 2002
Pararyctes pattersoni Van Valen 1966
Mammalia - Apatemyidae
Jepsenella cf. praepropera Simpson 1940
Mammalia - Cimolestidae
Paleotomus sp. Van Valen 1967
said to be new (5 measurements)
    = Paleotomus junior Scott et al. 2002
Scott et al. 2002
Paleotomus senior (Simpson 1937)
Mammalia - Pantolestidae
Propalaeosinopa septentrionalis (Russell 1929)
    = Bessoecetor septentrionalis Russell 1929
Scott et al. 2002
Mammalia - Pentacodontidae
Pentacodontidae indet. Simpson 1937
said to be new
Bisonalveus browni Gazin 1956
Mammalia - Pantodonta
Pantodonta indet. Cope 1873
Mammalia
Peradectes sp. Matthew and Granger 1921
Chronoperatidae
Chronoperates paradoxus n. gen., n. sp. Fox et al. 1992
Fox et al. 1992
see common names

Geography
Country:Canada State/province:Alberta
Coordinates: 51.2° North, 114.4° West (view map)
Paleocoordinates:57.7° North, 92.1° West
Basis of coordinate:stated in text
Altitude:1059 meters
Geographic resolution:outcrop
Time
Period:Paleogene Epoch:Paleocene
10 m.y. bin:Cenozoic 1
*Period:Tertiary *Local age/stage:Early/Lower Tiffanian
Key time interval:Ti1
Age range of interval:61.70000 - 56.80000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Paskapoo
Stratigraphic resolution:group of beds
Stratigraphy comments: reassigned from the Porcupine Hills Formation to the Paskapoo Formation by Fox (2011)
fossils are from "within an interval ranging from a few centimeters to a meter or so in thickness" (Fox 2011)
Lithology and environment
Primary lithology:pebbly,shelly/skeletal,green siltstone
Lithology description: "poorly bedded grayish−green siltstones, rich in broken bivalve and gastropod shells that are accompanied by bits of coalified wood and occasional well−rounded pebbles" (Fox 2011)
Environment:fluvial-lacustrine indet.
Geology comments: "Paskapoo sediments were deposited in the channels and floodplains of small to moderate-sized rivers, and in ponds, lakes, and swamps" (Fox 2011)
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:bulk,chemical,sieve
Collection size:1700 specimens
Reason for describing collection:biostratigraphic analysis
Collectors:G. P. Youzwyshyn Collection dates:1988
Collection method comments: see Scott et al. 2002
fossils were presumably collected over numerous field seasons
Taxonomic list comments:same as the list given by Fox 1990a
Metadata
Also known as:Cochrane 11; Cochrane II
Database number:14939
Authorizer:J. Alroy, P. Mannion, K. Beard, P. Holroyd Enterer:J. Alroy, K. Beard, P. Holroyd, P. Mannion
Modifier:J. Alroy Research group:vertebrate
Created:1993-02-18 00:00:00 Last modified:2011-02-24 02:53:46
Access level:the public Released:1993-02-18 00:00:00
Creative Commons license:CC BY
Reference information

Primary reference:

1534. R. C. Fox. 1988. Late Cretaceous and Paleocene mammal localities of southern Alberta. Occasional Paper of the Tyrrell Museum of Palaeontology 6:1-38 [J. Alroy/J. Alroy/P. Mannion]

Secondary references:

6294 J. Alroy. 2002. Synonymies and reidentifications of North American fossil mammals. [J. Alroy/J. Alroy/M. Kouvari]
35300 R. C. Fox. 2011. An unusual early primate from the Paleocene Paskapoo Formation, Alberta, Canada. Acta Palaeontologica Polonica 56(1):1-10 [J. Alroy/J. Alroy]
828 R. C. Fox, G. P. Youzwyshyn, and D. W. Krause. 1992. Post-Jurassic mammal-like reptile from the Palaeocene. Nature 358:233-235 [J. Alroy/M. Sommers/J. Alroy]
1542 R. C. Fox and G. P. Youzwyshyn. 1994. New primitive carnivorans (Mammalia) from the Paleocene of western Canada, and their bearing on relationships of the order. Journal of Vertebrate Paleontology 14(3):382-404 [J. Alroy/J. Alroy/J. Marcot]
65027 J. D. Gardner and D. G. DeMar. 2013. Mesozoic and Palaeocene lissamphibian assemblages of North America: a comprehensive review. Palaeobiodiversity and Palaeoenvironments 93:459-515 [P. Holroyd/P. Holroyd/M. Carrano]
1730 G. F. Gunnell. 1989. Evolutionary History of Microsyopoidea (Mammalia, ?Primates) and the Relationship Between Plesiadapiformes and Primates. University of Michigan Papers on Paleontology 27:1-157 [J. Alroy/J. Alroy/P. Mannion]
10095 C. S. Scott. 2004. A new species of the ptilodontoiid multituberculate Prochetodon (Mammalia, Allotheria) from the Paleocene Paskapoo Formation of Alberta, Canada. Canadian Journal of Earth Sciences 41(2):237-246 [J. Alroy/J. Alroy/J. Alroy]
7614 C. S. Scott, R. C. Fox, and G. P. Youzwyshyn. 2002. New earliest Tiffanian (late Paleocene) mammals from Cochrane 2, southwestern Alberta, Canada. Acta Palaeontologica Polonica 47(4):691-704 [J. Alroy/J. Alroy/J. Alroy]
14679 C. S. Scott and R. C. Fox. 2005. Windows on the evolution of Picrodus (Plesiadapiformes: Primates): morphology and relationships of a species complex from the Paleocene of Alberta. Journal of Paleontology 79(4):635-657 [J. Alroy/J. Alroy]
33356 C. S. Scott and D. W. Krause. 2006. Multituberculates (Mammalia, Allotheria) from the earliest Tiffanian (late Paleocene) Douglass Quarry, eastern Crazy Mountains Basin, Montana. Contributions from the Museum of Paleontology, University of Michigan 31(10):211-243 [K. Beard/K. Beard]
84657 C. S. Scott, S. López-Torres, M. T. Silcox and R. C. Fox. 2023. New paromomyids (Mammalia, Primates) from the Paleocene of southwestern Alberta, Canada, and an analysis of paromomyid interrelationships. Journal of Paleontology 97(2):477-498 [P. Mannion/P. Mannion]
19168 C. S. Scott, M. W. Webb, and R. C. Fox. 2006. Horolodectes sunae, an enigmatic mammal from the late Paleocene of Alberta, Canada. Journal of Paleontology 80(5):1009-1025 [J. Alroy/J. Alroy]
42892 S. P. Zack, T. A. Penkrot, D. W. Krause and M. C. Maas. 2005. A new apheliscine "condylarth" mammal from the late Paleocene of Montana and Alberta and the phylogeny of "hyopsodontids". Acta Palaeontologica Polonica 50(4):809-830 [P. Mannion/P. Mannion]