Swain Quarry: Torrejonian, Wyoming

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Amphibia - Temnospondyli - Batrachosauroididae
Opisthotriton kayi Auffenberg 1961
Gardner and DeMar 2013
Amphibia - Temnospondyli - Urodela
Habrosaurus sp. Gilmore 1928
Gardner and DeMar 2013
Scapherpeton tectum Cope 1876
Gardner and DeMar 2013
Mammalia - Multituberculata - Neoplagiaulacidae
Ectypodus sylviae n. sp. Rigby 1980
Rigby 1980
recombined as Parectypodus sylviae
Xanclomys mcgrewi n. sp. Rigby 1980
Rigby 1980
Parectypodus clemensi Sloan 1981
Sloan 1981
recombined as Krauseia clemensi
may include Rigby's new "Ectypodus sp.," said to be the same as the Bangtail species by Gingerich et al. 1983, who do not cite Sloan (1 measurement)
Ectypodus szalayi Sloan 1981
Sloan 1981
Neoplagiaulax nelsoni Sloan 1987
Sloan 1987
nomen nudum belonging to Neoplagiaulax
N. "cf. hunteri"
Xyronomys swainae n. sp. Rigby 1980
Rigby 1980
Mammalia - Multituberculata - Ptilodontidae
Baiotomeus douglassi (Simpson 1935)
Krause 1987
one specimen of "Ptilodus mediaevus"
Ptilodus mediaevus Cope 1881
Rigby 1980
Mammalia
Psittacotherium multifragum Cope 1882
Rigby 1980
Mammalia - Leptictidae
Diacodon concordiarcensis (Simpson 1935)
Rigby 1980
original and current combination Prodiacodon concordiarcensis
Prodiacodon cf. puercensis (Matthew 1918)
Rigby 1980
Myrmecoboides montanensis Gidley 1915
Rigby 1980
Mammalia - Mixodectidae
Mixodectes sp. Cope 1883
Rigby 1980
Mixodectes malaris (Cope 1884)
Rigby 1980
confirmed by Gunnell 1989
Mammalia - Primates - Paromomyidae
Paromomys maturus Gidley 1923
Rigby 1980
confirmed by Gunnell 1989
Paromomys depressidens Gidley 1923
Rigby 1980
confirmed by Gunnell 1989
Mammalia - Primates - Palaechthonidae
Palenochtha cf. minor (Gidley 1923)
Rigby 1980
    = Palenochtha minor Gidley 1923
Gunnell 1989
Palenochtha weissae n. sp. Rigby 1980
Rigby 1980
(1 measurement)
Palaechthon problematicus (Jepsen 1930)
Rigby 1980
original and current combination Plesiolestes problematicus
(7 measurements)
Mammalia - Primates - Picrodontidae
Picrodus silberlingi Douglass 1908
Rigby 1980
Mammalia - Nyctitheriidae
Leptacodon cf. tener Matthew and Granger 1921
Rigby 1980
Mammalia - Periptychidae
Periptychus carinidens Cope 1881
Rigby 1980
Haploconus sp. Cope 1882
Rigby 1980
Anisonchus sectorius (Cope 1881)
Rigby 1980
Anisonchus willeyi n. sp. Rigby 1980
Rigby 1980
Mammalia - Phenacodontidae
Tetraclaenodon puercensis (Cope 1881)
Rigby 1980
Mammalia - Procreodi - Arctocyonidae
Arctocyon ferox (Cope 1883)
Rigby 1980
Mimotricentes subtrigonus (Cope 1881)
Rigby 1980
recombined as Tricentes subtrigonus
(2 measurements)
Chriacus pelvidens (Cope 1881)
Rigby 1980
Protogonodon criswelli n. sp. Rigby 1980
Rigby 1980
recombined as Loxolophus criswelli
(1 measurement)
Mammalia - Viverravidae
Bryanictis vanvaleni (MacIntyre 1966)
Rigby 1980
recombined as Intyrictis vanvaleni
Protictis haydenianus (Cope 1882)
Rigby 1980
Prolimnocyon macfaddeni n. sp. Rigby 1980
Rigby 1980
synonym of Protictis haydenianus
macfaddeni
Simpsonictis jaynanneae n. sp. Rigby 1980
Rigby 1980
Mammalia - Acreodi - Triisodontidae
Goniacodon levisanus (Cope 1883)
Rigby 1980
Mammalia - Acreodi - Mesonychidae
Dissacus navajovius (Cope 1881)
Rigby 1980
Mammalia - Deltatheridia
Deltatheridia indet. Van Valen 1965
Rigby 1980
Mammalia - Condylarthra - Chriacidae
Prothryptacodon hilli n. sp. (Rigby 1980)
Rigby 1980
Prothryptacodon cf. furens Simpson 1935
Rigby 1980
Mammalia - Condylarthra - Hyopsodontidae
Litaletes sp. Simpson 1935
Robison 1986
said to be new and not the same as "L. mantiensis", a jun. syn. of L. disjunctus
Promioclaenus acolytus (Cope 1882)
Rigby 1980
Mammalia - Macroscelidea - Apheliscidae
Haplaletes disceptatrix Simpson 1935
Rigby 1980
Litomylus dissentaneus Simpson 1935
Rigby 1980
Mammalia - Macroscelidea
Mckennatherium fredericki n. sp. Rigby 1980
Rigby 1980
recombined as Adunator fredericki
Mckennatherium martinezi n. sp. Rigby 1980
Rigby 1980
recombined as Adunator martinezi
Mammalia - Proteutheria - Palaeoryctidae
Palaeoryctes puercensis Matthew 1913
Rigby 1980
Mammalia - Insectivora
Diacodon sp. Cope 1875
Rigby 1980
    = Insectivora indet. Bowdich 1821
Alroy 2002
Mammalia - Apatemyidae
Jepsenella praepropera Simpson 1940
Rigby 1980
(1 measurement)
Mammalia - Cimolestidae
Gelastops joni n. sp. Rigby 1980
Rigby 1980
Paleotomus milleri n. sp. Rigby 1980
Rigby 1980
Acmeodon hyoni n. sp. Rigby 1980
Rigby 1980
synonym of Acmeodon secans
Mammalia - Pantolestidae
Propalaeosinopa diluculi (Simpson 1935)
Rigby 1980
    = Bessoecetor septentrionalis Russell 1929
Scott et al. 2002
Pantolestinae indet. Simpson 1937
Rigby 1980
Mammalia - Groeberida
Peradectes sp. Matthew and Granger 1921
Rigby 1980
    = Swaindelphys cifellii n. sp. Johanson 1996
Johanson 1996
said to be new by Rigby
Reptilia - Baenidae
Plesiobaena putorius Gaffney 1972
Gaffney 1972 2 specimens
recombined as Cedrobaena putorius
AMNH 4950, anterior portion of skull roof; 4949, posterior portion of skull roof
Reptilia - Squamata
Squamata indet. Oppel 1811
Sullivan 1982 76 specimens
"indeterminate Sauria" AMNH 12041, ventral part of basicranium; AMNH 12046, 14301, 14322, 14323, 14324, 14325, 14326, 15895, 15896, 15897, 15903, 15904, 15906, 15921, 15922, 15924, 15927, 15930, 15941, 15942, 15944, 15948, 15968, 15997, 15998, 16005, 16009, 16013, 16016, 16018, 16020, 16154, 16158, 16160, 16163, 16166, 16167, 16168, 16171, 16172, 16173, 16174, 16177, 16178, 16179, 16180, 16181, 16183, 16188, 16189, 16190, 16192, 16193, (?) 16194, 16196, 16197, 16198, 16216, 16217, 16218, partial dentaries; AMNH 15932, 15934, 15939, (?) 15946, 15947, 15950, 15964, 16025, partial maxillae; AMNH 15943, 15954, 16195, tooth-bearing fragments; AMNH 15958, proximal end of femur; AMNH 15962, trunk vertebra; AMNH 16056, distal end of humerus; AMNH 16137, caudal vertebra.
Reptilia - Xantusiidae
cf. Contogenys sloani Estes 1969
Sullivan 1982
scincid
    = Palaeoxantusia fera Hecht 1956
Gao and Fox 1996
'indistinguishable in both size and tooth form from Palaeoxantusia from the same locality'
Palaeoxantusia fera Hecht 1956
Sullivan 1982 43 specimens
AMNH 12028, posterior part of right maxilla; AMNH 12036, posterior part of left spleniodentary; AMNH 12038, posterior half of left spleniodentary; AMNH 12039, anterior part of right maxilla; AMNH 12040, left spleniodentary; AMNH 12041, posterior part of right spleniodentary; AMNH 12049, medial part of left spleniodentary; AMNH 12064, nearly complete left spleniodentary; AMNH 12071, anterior part of right spleniodentary; AMNH 12072, anterior part of right spleniodentary; AMNH 12073, posterior part of right spleniodentary; AMNH 12074, posterior part of right spleniodentary; AMNH 12075, left spleniodentary; AMNH 12076, nearly complete right spleniodentary; AMNH 12077, posterior part of right spleniodentary; AMNH 12079, anterior part of right spleniodentary; AMNH 12080, posterior part of right spleniodentary; AMNH 12083 (Text-fig. 6), posterior part of left maxilla; AMNH 14302, medial part of right spleniodentary; AMNH 14312, 4 teeth; AMNH 14313, medial part of right spleniodentary; AMNH 14314, posterior part of right maxilla; AMNH 15907, medial part of left spleniodentary; AMNH 15917, medial part of left spleniodentary; AMNH 15920, greater anterior part of left spleniodentary; AMNH 15926, posterior part of left maxilla; AMNH 15928, medial part of left spleniodentary; AMNH 15 929, greater anterior part of right spleniodentary; AMNH 15938, posterior part of left maxilla; AMNH 15949, posterior part of right spleniodentary; AMNH 15952, posterior part of right maxilla; AMNH 15953, medial part of left spleniodentary; AMNH 16007, nearly complete left spleniodentary; AMNH 16156, medial part of left spleniodentary; AMNH 16157, anterior part of left spleniodentary; AMNH 16170, anterior part of left spleniodentary. Also referred tentatively: AMNH 12043, medial part of right spleniodentary; AMNH 12059, medial part of right spleniodentary; AMNH 15925, nearly complete left maxilla; AMNH 15955, incomplete medial part of right spleniodentary; AMNH 16049, spleniodentary fragment; AMNH 16159, medial part of left spleniodentary; AMNH 16182, incomplete posterior part of right maxilla.
Reptilia - Iguanidae
Swainiguanoides milleri n. gen., n. sp. Sullivan 1982
Sullivan 1982 5 specimens
AMNH 12082 (holotype), medial part of right dentary; 12081, medial part of right dentary; 15902, medial part of left dentary; 15933, medial part of left dentary; 12048, medial part of left maxilla
Reptilia
cf. Saniwa sp. Leidy 1870
Sullivan 1982 2 specimens
AMNH 15960, trunk vertebra; AMNH 16003, incomplete trunk vertebra.
cf. Palaeosaniwa canadensis Gilmore 1928
Sullivan 1982 1 specimen
AMNH 15957, medial part of left dentary with 3 tooth bases.
    = cf. Saniwa sp. Leidy 1870
Gao and Fox 1996
Reptilia - Anguidae
Anguidae indet. Gray 1825
Sullivan 1982 6 specimens
AMNH 12052, 12053, 12056, 12057, 12058, 12093, partial dentaries.
Machaerosaurus torrejonensis Gilmore 1928
Sullivan 1982 32 specimens
AMNH 12047, medial part of left dentary; AMNH 12063, posterior part of right maxilla; AMNH 12066, medial part of right dentary; AMNH 12068, anterior part of right maxilla; AMNH 12087, left frontal; AMNH 12094, medial part of left dentary; AMNH 12095, posterior part of right maxilla; AMNH 12097, anterior part of left maxilla; AMNH 12099, nearly complete right dentary; AMNH 14311, posterior part of right dentary; AMNH 15911, left dentary; AMNH 15914, posterior part of left dentary; AMNH 16100, nearly complete right frontal. Also tentatively referred: AMNH 12034, posterior part of left maxilla; AMNH 12050, greater part of left dentary; AMNH 12051, posterior part of right dentary; AMNH 12055, medial part of left dentary; AMNH 12092, posterior part of right dentary; AMNH 14303, greater part of right dentary; AMNH 14327, posterior part of right maxilla; AMNH 15898, anterior part of left dentary; AMNH 15899, left maxilla; AMNH 15908, posterior part of a left maxilla; AMNH 15909, greater part of right maxilla; AMNH 15913, posterior part of left maxilla; AMNH 15931, posterior part of left dentary; AMNH 15935, anterior part of right maxilla; AMNH 15936, maxilla fragment; AMNH 15951, posterior part of right maxilla; AMNH 16008, posterior part of left dentary; AMNH 16010, posterior part of right maxilla.
Odaxosaurus sp. Gilmore 1928
ElShafie 2024 1 specimen
CM 87168
Odaxosaurus piger (Gilmore 1928)
Sullivan 1982 14 specimens
AMNH 12030, posterior part of right dentary; AMNH 12032, right dentary; AMNH 12033, medial part of right dentary; AMNH 12035, posterior part of right dentary; AMNH 12060, medial part of left dentary; AMNH 12084, medial part of left dentary; AMNH 12085, posterior part of left dentary; AMNH 12086, posterior part of left maxilla; AMNH 14310, medial part of left dentary; AMNH 15900, posterior part of left maxilla; AMNH 15901, posterior part of right maxilla; AMNH 15940, posterior part of right dentary. Also tentatively referred: AMNH 15905, posterior part of right dentary; AMNH 15918, medial part of right dentary.
Odaxosaurinae indet. Sullivan 1982
Sullivan 1982 13 specimens
AMNH 12089, 21 body osteoderms; AMNH 15910, posterior part of right maxilla; AMNH 15916, right maxilla fragment with tooth; AMNH 15937, posterior part of right maxilla; AMNH 15956, posterior part of left maxilla; AMNH 16004, medial part of right dentary; AMNH 16011, medial part of right dentary; AMNH 16012, anterior part of right dentary; AMNH 16109, frontal fragment; AMNH 16124 and 16125, body osteoderms, associated with frontal fragments TEXT-FIG. 11-Glyptosaurinae, genus and species indeterminate (AMNH 12088), right pterygoid (palatal view), x10. and garscales; AMNH 16176, maxilla fragment; AMNH 16209, (?) cheek osteoderm.
Glyptosaurinae indet. Marsh 1872
Sullivan 1982 8 specimens
AMNH 12031, posterior part of right maxilla; AMNH 12042, medial part of left dentary; AMNH 12044, premaxilla; AMNH 12045, anterior part of left dentary; AMNH 12065, medial part of right dentary; AMNH 12070, anterior part of right maxilla; AMNH 12088, tooth bearing region of right pterygoid; AMNH 12091, medial part of (?) right maxilla
Reptilia - Xenosauridae
Exostinus cf. lancensis Gilmore 1928
Sullivan 1982 4 specimens
AMNH 14315, medial part of right dentary; AMNH 14316, medial part of left dentary; AMNH 14317, medial part of left dentary; AMNH 14318, anterior part of left dentary.
Reptilia - Necrosauridae
cf. Provaranosaurus sp. Gilmore 1942
Sullivan 1982 3 specimens
AMNH 14306, medial part of left dentary; AMNH 14307, medial part of left dentary; AMNH 14309, medial part of right dentary
see common names

Geography
Country:United States State/province:Wyoming County:Carbon
Coordinates: 41.3° North, 107.7° West (view map)
Paleocoordinates:47.7° North, 86.2° West
Basis of coordinate:stated in text
Geographic resolution:small collection
Time
Period:Paleogene Epoch:Paleocene
Stage:Danian 10 m.y. bin:Cenozoic 1
*Period:Early/Lower Tertiary *Epoch:Early/Lower Paleocene
*Local age/stage:Torrejonian
Key time interval:Torrejonian
Age range of interval:63.30000 - 61.70000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Fort Union
Stratigraphic resolution:group of beds
Lithology and environment
Primary lithology: not reported
Environment:terrestrial indet.
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Collection methods and comments
Collection methods:surface (in situ),field collection
Collection size:28000 specimens
Reason for describing collection:taxonomic analysis
Taxonomic list comments:there are seven other localities in the same section, but none of the material is described
Mesodma pygmaea, i.e. sp. P, not listed, but mentioned by Krause 1977
Metadata
Also known as:CM loc 217
Database number:14886
Authorizer:J. Alroy, R. Benson, P. Holroyd, P. Barrett Enterer:J. Alroy, T. Cleary, P. Holroyd, R. Benson
Modifier:P. Holroyd Research group:vertebrate
Created:1993-02-18 00:00:00 Last modified:2024-01-07 15:18:33
Access level:the public Released:1993-02-18 00:00:00
Creative Commons license:CC BY
Reference information

Primary reference:

3591. R. M. West. 1970. Tetraclaenodon puercensis (Mammalia: Phenacodontidae), Goler Formation, Paleocene of California, and distribution of the genus. Journal of Paleontology 44(5):851-857 [J. Alroy/J. Alroy/M. Carrano]

Secondary references:

6294 J. Alroy. 2002. Synonymies and reidentifications of North American fossil mammals. [J. Alroy/J. Alroy/M. Kouvari]
87093 S. J. ElShafie. 2024. Body size estimation from isolated fossil bones reveals deep time evolutionary trends in North American lizards. PLoS ONE 19(1:e0296318):1-25 [P. Holroyd/P. Holroyd/P. Holroyd]
704 E. S. Gaffney. 1972. The systematics of the North American family Baenidae (Reptilia, Cryptodira). Bulletin of the American Museum of Natural History 147(5):245-312 [J. Alroy/M. Sommers/R. Benson]
37933 K. -Q. Gao and R. C. Fox. 1996. Taxonomy and evolution of Late Cretaceous lizards (Reptilia: Squamata) from western Canada. Bulletin of the Carnegie Museum of natural History 33:1-107 [R. Benson/R. Benson]
65027 J. D. Gardner and D. G. DeMar. 2013. Mesozoic and Palaeocene lissamphibian assemblages of North America: a comprehensive review. Palaeobiodiversity and Palaeoenvironments 93:459-515 [P. Holroyd/P. Holroyd/M. Carrano]
1730 G. F. Gunnell. 1989. Evolutionary History of Microsyopoidea (Mammalia, ?Primates) and the Relationship Between Plesiadapiformes and Primates. University of Michigan Papers on Paleontology 27:1-157 [J. Alroy/J. Alroy/P. Mannion]
2035 Z. Johanson. 1996. New marsupial from the Fort Union Formation, Swain Quarry, Wyoming. Journal of Paleontology 70(6):1023-1031 [J. Alroy/J. Alroy/J. Alroy]
2160 D. W. Krause. 1987. Baiotomeus, a New Ptilodontid Multituberculate (Mammalia) from the Middle Paleocene of Western North America . Journal of Paleontology 61(3):595-603 [J. Alroy/J. Alroy/J. Ju]
2905 J. K. Rigby, Jr. 1980. Swain Quarry of the Fort Union Formation, Middle Paleocene (Torrejonian), Carbon County, Wyoming: geologic setting and mammalian fauna. Evolutionary Monographs 3:1-178 [J. Alroy/J. Alroy/J. Marcot]
2930 S. F. Robison. 1986. Paleocene (Puercan-Torrejonian) mammalian faunas of the North Horn Formation, central Utah. Brigham Young University Geology Studies 33(1):87-133 [J. Alroy/J. Alroy]
3195 R. E. Sloan. 1981. Systematics of Paleocene multituberculates from the San Juan Basin, NewMexico. In S. G. Lucas, J. K. Rigby Jr., B. S. Kues (eds.), Advances in San Juan Basin Paleontology 127-160 [J. Alroy/J. Alroy/M. Uhen]
3196 R. E. Sloan. 1987. Paleocene and latest Cretaceous mammal ages, biozones, magnetozones, rates of sedimentation, and evolution. Geological Society of America Special Paper 209:165-200 [J. Alroy/J. Alroy/M. Uhen]
7573 R. M. Sullivan. 1982. Fossil lizards from Swain Quarry "Fort Union Formation," middle Paleocene (Torrejonian), Carbon County, Wyoming. Journal of Paleontology 56(4):996-1010 [J. Alroy/J. Alroy/J. Alroy]