Angeac-Charente: Berriasian, France
collected by Néraudeau et al. 2008–2018

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
unclassified
Ammocycloloculina sp.
Eoparacypris sp.
Microcarpolithes hexagonalis
Rozada et al. 2020
Lobionella hirsute
Rozada et al. 2020
Amphibia - Temnospondyli
Anura indet. (Fischer von Waldheim 1813)
Allain et al. 2022
? Caudata indet. Scopoli 1777
Allain et al. 2022
Amphibia - Temnospondyli - Albanerpetontidae
Albanerpetontidae indet. Fox and Naylor 1982
Allain et al. 2022
Reptilia
Ornithischia indet. (Seeley 1888)
Rozada et al. 2020 9 specimens
Ankylosauria indet. (Osborn 1923)
Allain et al. 2022
Stegosauria indet. (Marsh 1877)
Rozada et al. 2020 84 specimens
Deltapodus sp. Whyte and Romano 1995
Rozada et al. 2020 100 specimens
tracks, ANG 16 Ep60
Reptilia - Stegosauridae
Dacentrurus sp. Lucas 1902
Allain et al. 2022
Reptilia - Hypsilophodontidae
Hypsilophodon sp. Huxley 1870
6 specimens
ANG 10-153, teeth
    = Hypsilophodontidae indet. Dollo 1882
Rozada et al. 2020
Reptilia - Camptosauridae
Iguanodontidae indet. Bonaparte 1850
2 specimens
2 teeth, 1 femur
    = Camptosauridae indet. Marsh 1885
Rozada et al. 2020
Reptilia - Heterodontosauridae
Heterodontosauridae indet. Romer 1966
Rozada et al. 2020 2 specimens
teeth
Echinoidea indet. (Leske 1778)
Rozada et al. 2020
debris
    = Echinodon sp. Owen 1861
Allain et al. 2022
Reptilia
Ornithomimosauria indet. (Barsbold 1976)
3800 specimens
ANG 10-175
Reptilia - Dromaeosauridae
cf. Nuthetes sp. Owen 1854
Allain et al. 2022
Reptilia - Archaeopterygidae
Archaeopterygidae indet. Huxley 1871
Allain et al. 2022
invalid subgroup of Avialae
Reptilia
Tyrannosauroidea indet. Walker 1964
Rozada et al. 2020 6 specimens
teeth
cf. Neovenator sp. Hutt et al. 1996
133 specimens
ANG 10-51, 10-262; teeth
    = Megalosauroidea indet. Nopcsa 1928
Rozada et al. 2020
Sauropoda indet. (Marsh 1878)
Rozada et al. 2020 58627 specimens
tracks, ANG 18‐6529 and many bone fragments
Turiasauria indet. Royo-Torres et al. 2006
Rozada et al. 2020 929 specimens
ANG 10-400, femur; ANG 10-10, 10-27, teeth
Macronaria indet. Sereno 1997
Rozada et al. 2020 1 specimen
tooth
Pterosauria indet. (Kaup 1834)
Allain et al. 2022
Pterodactyloidea indet. (Plieninger 1901)
Allain et al. 2022
Crocodylomorpha indet. (Hay 1930)
Rozada et al. 2020 116 specimens
Reptilia - Bernissartiidae
Bernissartiidae indet. (Dollo 1883)
7 specimens
ANG 10-268
Reptilia - Goniopholididae
Goniopholididae indet. Cope 1875
885 specimens
ANG 10-263
    = Goniopholis sp. Owen 1842
Allain et al. 2022
Reptilia
Pholidosauridae indet. von Zittel and Eastman 1902
267 specimens
    = Pholidosaurus sp. Meyer 1841
Allain et al. 2022
Reptilia - Atoposauridae
Atoposauridae indet. (Gervais 1871)
7 specimens
ANG 10-267
Reptilia
Theriosuchus sp. Owen 1878
Reptilia - Plesiosauria
Pliosauroidea indet. Welles 1943
1 specimen
ANG 10-581, left tibia
Reptilia - Cteniogenyidae
cf. Cteniogenys sp. Gilmore 1928
Allain et al. 2022
Reptilia
cf. Opisthias sp. Gilmore 1909
Allain et al. 2022
Reptilia - Paramacellodidae
Paramacellodus sp. Hoffstetter 1967
Allain et al. 2022
Reptilia - Testudines
Chelonii indet. Latreille 1800
105 specimens
synonym of Testudines
Reptilia
Hylaeochelys ? belli (Mantell 1844)
Allain et al. 2022
Reptilia - Plesiochelyidae
Plesiochelyidae indet. Rütimeyer 1873
Rozada et al. 2020 14 specimens
Reptilia
28 specimens
ANG 10-409
    = Helochelydra nopcsai Lapparent de Broin and Murelaga 1999
Joyce 2017
    = Helochelydridae indet. Nopcsa 1928
Rozada et al. 2020
Reptilia - Pleurosternidae
Pleurosternidae indet. Cope 1868
730 specimens
ANG 10-71, carapace (ANG14R-454), the plastron (ANG14R- 474) and right peripheral plates 1 and 2 (ANG 15 R-1082a and b) were collected separately in the field; the material presents evidence of tooth marks
    = Pleurosternon bullockii Owen 1842
Gonet et al. 2019
Mammalia
Mammalia indet. Linnaeus 1758
Allain et al. 2022
Mammalia - Multituberculata - Paulchoffatiidae
Allain et al. 2022
Mammalia - Multituberculata - Pinheirodontidae
Pinheirodontidae indet. Hahn and Hahn 1999
Allain et al. 2022
Mammalia - Spalacotheriidae
Spalacotherium evansae Ensom and Sigogneau-Russell 2000
Allain et al. 2022
Mammalia - Dryolestidae
Dryolestidae indet. Marsh 1879
Allain et al. 2022
Mammalia - Peramura - Peramuridae
Peramus sp. Owen 1871
Allain et al. 2022
Mammalia - Eutriconodonta - Gobiconodontidae
? Gobiconodon sp. Trofimov 1978
Allain et al. 2022
Mammalia - Eutriconodonta - Triconodontidae
? Triconodon sp. Owen 1859
Allain et al. 2022
Mammalia - Thereuodontidae
Thereuodon cf. taraktes Sigogneau-Russell and Ensom 1998
Allain et al. 2022
Ionoscopiformes
Ionoscopiformes indet. Grande and Bemis 1998
Actinopteri - Amiiformes
Amiiformes indet. Hay 1929
Allain et al. 2022
Actinopteri - Amiiformes - Caturidae
Caturidae indet. Owen 1860
Actinopteri
Ginglymodi indet. (Cope 1871)
Allain et al. 2022
Actinopteri - Lepisosteiformes
Lepidotes sp. Agassiz 1832
Actinopteri - Semionotiformes - Semionotidae
Semionotidae indet. Woodward 1890
Actinopteri - Pycnodontiformes - Mesturidae
cf. Micropycnodon sp. Hibbard and Graffham 1945
Allain et al. 2022
Actinopteri - Pycnodontiformes - Pycnodontidae
Pycnodontiformes indet. Berg 1937
    = Pycnodontidae indet. Agassiz 1833
Allain et al. 2022
Chondrichthyes - Hybodontiformes - Lonchidiidae
Parvodus celsucuspus Rees et al. 2013
Allain et al. 2022
Chondrichthyes - Hybodontiformes - Hybodontidae
Hybodontidae indet. (Agassiz 1834)
2 specimens
ANG 10-126, incomplete tooth and a worn, fragmentary dorsal fin-spine
unclassified
Bryozoa indet. (Ehrenberg 1831)
Rozada et al. 2020
debris
Ostracoda - Podocopida - Cyprididae
Cypridea cf. tuberculata
Cypridea laevigata
Cypridea sp. Bosquet 1852
Two species
Ostracoda - Podocopida - Limnocytheridae
Theriosynoecum forbesii
Ostracoda - Podocopida
Fabanella boloniensis
Rozada et al. 2020
Equisetopsida - Araucariaceae
Agathoxylon sp. Hartig 1848
Algae - Zygnemataceae
Ovoidites parvus Nakoman 1966
Rozada et al. 2020
Lecaniella sp. Cookson and Eisenack 1962
Rozada et al. 2020
Coniferales - Cupressaceae
Protocupressinoxylon sp. Eckhold 1921
Charales - Characeae
Sphaerochara aff. verticillata (Peck 1937)
recombined as Peckisphaera verticillata
Charales - Porocharaceae
Porochara sp. Maedler 1955
Charales
Clavator grovesii
Charales - Characea
Mesochara sp. Grambast 1962
unclassified
Nodosoclavator bradleyi
Trocholinidae
Trocholina cf. odukpaniensis
    = Trocholina odukpaniensis
Rozada et al. 2020
Dinophyceae - Gonyaulacales - Ceratiaceae
Odontochitina imparilis Jain and Khowaja-Ateequzzaman 1984
Rozada et al. 2020
see common names

Geography
Country:France State/province:Poitou-Charentes County:Charente
Coordinates: 45.6° North, 0.1° East (view map)
Paleocoordinates:36.5° North, 9.8° East
Basis of coordinate:estimated from map
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Early/Lower Cretaceous
Stage:Berriasian 10 m.y. bin:Cretaceous 1
Key time interval:Berriasian
Age range of interval:145.00000 - 139.80000 m.y. ago
Stratigraphy
Stratigraphic resolution:group of beds
Stratigraphy comments: "Gardonian". Original stratigraphy used layers An1 to An5, top to bottom: An 1 (fine sand), An 2 (calcareous), An 3 (calcareous conglomerate), An 4 (lignitic clay), and An 5 (green clay). These were redefined in 2020 with a new system, bottom to top, so that: An 5 = Unit 1; An 4 is split into Unit 2 (sandy clay) and Unit 3 (lignitic clay with lenses); An 2 + An 3 = Unit 4. An 5 is no longer present. The bonebed represents units 2, 3, and 4.
From Allain et al. 2022: The recently proposed earliest Cretaceous (Berriasian) age of the Angeac-Charente locality, based on both charophytes and palynomorphs (Benoit et al. 2017; Polette et al. 2018), is here confirmed by the vertebrate fauna.
Lithology and environment
Primary lithology:gray,blue sandy,carbonaceous claystone
Secondary lithology:intraclastic,gray,blue calcareous conglomerate
Includes fossils?Y
Lithology description: An1: "fine white to greyish sand"; An2: "beige to yellow and grey calcareous deposits with irregular stratification and both lateral and vertical transitions between marls and limestones"; An3: calcareous conglomerate with a blue-grey to beige grainstone matrix and whitish limestone clasts embedded in it"; An4: blue-grey lignitic clay"
Environment:mire/swamp
Geology comments: "deposition in subtropical to tropical conditions" "accumulated mainly in freshwater" - progressive flooding of a swamp.
Taphonomy
Modes of preservation:body
Lagerst�tten type:concentrate
Degree of concentration:concentrated
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:excellent
Associated major elements:some
Disassociated major elements:many
Disassociated minor elements:many
Fragmentation:occasional
Feeding/predation traces:tooth marks
Temporal resolution:time-averaged
Spatial resolution:parautochthonous
Collection methods and comments
Collection methods:bulk,selective quarrying,mechanical,sieve,field collection
Reason for describing collection:general faunal/floral analysis
Collectors:Néraudeau et al. Collection dates:2008–2018
Metadata
Database number:142093
Authorizer:P. Mannion, E. Vlachos, M. Carrano Enterer:J. Tennant, E. Vlachos, P. Mannion, M. Carrano
Modifier:M. Carrano Research group:micropaleontology,paleobotany,vertebrate
Created:2013-04-05 02:17:53 Last modified:2022-12-08 14:45:36
Access level:the public Released:2013-04-05 02:17:53
Creative Commons license:CC BY
Reference information

Primary reference:
46154. D. Néraudeau, R. Allain, M. Ballèvre, D. J. Batten, E. Buffetaut, J. P. Colin, M. P. Dabard, V. Daviero-Gomez, A. El Albani, B. Gomez, D. Grosheny, J. Le Loeuff, A. Leprince, C. Martín-Closas, E. Masure, J.-M. Mazin, M. Phillipe, J. Pouech, H. Tong, J. F. Tournepiche, and R. Vullo. 2012. The Hautevarian–Barremian lignitic bone bed of Angeac (Charente, south-west France): stratigraphical, palaeobiological and palaeogeographical implications. Cretaceous Research 37(1):1-14 [P. Mannion/J. Tennant/P. Mannion]

Secondary references:

81286 R. Allain, L. Rozada, and R. Vullo. 2013. A cross-perspective into the Early Cretaceous dinosaur faunas from Europe and Asia: new data on the dinosaurs from the Savannakhet Bassin (Laos) and from the lignitic bonebed of Angeac-Charente (France). In F. Torcida Fernández-Baldor & P. Huerta (ed.), VI Jornadas Internacionales sobre Paleontología de Dinosaurios y su Entorno. Libro de Resúmenes 17-18 [M. Carrano/M. Carrano]
51312 R. Allain, R. Vullo, J. Le Loeuff and J.-F. Tournepiche. 2014. European ornithomimosaurs (Dinosauria, Theropoda): an undetected record. Geological Acta 12(2):127-135 [P. Mannion/J. Tennant/M. Carrano]
82353 R. Allain, R. Vullo, L. Rozada, J. Anquetin, R. Bourgeais, J. Goedert, M. Lasseron, J. E. Martin, A. Pérez-García, C. Peyre De Fabrègues, R. Royo-Torres, D. Augier, G. Bailly, L. Cazes, Y. Despres, A. Gailliègue, B. Gomez, F. Goussard, T. Lenglet, R. Vacant, Mazan and J.-F. Tour. 2022. Vertebrate paleobiodiversity of the Early Cretaceous (Berriasian) Angeac-Charente Lagerstätte (southwestern France): implications for continental faunal turnover at the J/K boundary. Geodiversitas 44(25):683-752 [E. Vlachos/E. Vlachos]
67101 J. Gonet, L. Rozada, R. Bourgeais and R. Allain. 2019. Taphonomic study of a pleurosternid turtle shell from the Early Cretaceous of Angeac-Charente, southwest France. Lethaia 55(2):232-243 [E. Vlachos/E. Vlachos/E. Vlachos]
62124 W. G. Joyce. 2017. A Review of the Fossil Record of Basal Mesozoic Turtles. Bulletin of the Peabody Museum of Natural History 58(1):65-113 [P. Holroyd/P. Holroyd/P. Mannion]
83324 J. Le Loeuff, D. Néraudeau, R. Vullo, A. Leprince, R. Allain and E. Buffetaut. 2011. A giant sauropod from the Barremian of France. In A. van der Geer & A. Athanassiou (ed.), European Association of Vertebrate Palaeontologists, 9th Annual Meeting, Program and Abstracts 36 [M. Carrano/M. Carrano]
76586 L. Rozada, R. Allain, R. Vullo, J. Goedert, D. Augier, A. Jean, J. Marchal, C. Peyre Fabrègues, M. Qvarnström and R. Royo-Torres. 2020. A Lower Cretaceous Lagerstätte from France: a taphonomic overview of the Angeac‐Charente vertebrate assemblage. Lethaia 54:141-165 [M. Carrano/M. Carrano]