Bug Creek Anthills: Late/Upper Maastrichtian, Montana
collected by Univ. of Minnesota 1962-1964

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Amphibia - Temnospondyli
Anura indet. (Fischer von Waldheim 1813)
Estes et al. 1969 3 specimens
Amphibia - Temnospondyli - Pelobatidae
? Eopelobates sp. Parker 1929
Estes et al. 1969 1 specimen
    = cf. Eopelobates sp. Parker 1929
Gardner 2008
UALVP 40164
Amphibia - Temnospondyli - Discoglossidae
Discoglossidae indet. Günther 1859
Estes et al. 1969 2 specimens
cf. Barbourula sp. Taylor and Noble 1924
Estes et al. 1969 1 specimen
Amphibia - Temnospondyli
Scotiophryne pustulosa n. gen., n. sp. Estes 1969
Estes et al. 1969 11 specimens
MCZ 3623-3626; AMNH 26422; UALVP 40155-60
Amphibia - Temnospondyli - Palaeobatrachidae
Palaeobatrachus occidentalis Estes and Sanchíz 1982
Gardner 2008 1 specimen
UALVP 40163
Amphibia - Temnospondyli - Urodela
Proamphiuma cretacea Estes 1969
Estes et al. 1969 2 specimens
Habrosaurus dilatus Gilmore 1928
Estes et al. 1969 9 specimens
cf. Piceoerpeton sp. Meszoely 1967
Naylor 1983 2 specimens
UA 14918 (atlas)
    = Piceoerpeton naylori n. sp. Gardner 2012
Gardner 2012
UALVP 14918 - type (atlantal centrum); referred specimen: UALVP 40122, trunk vertebra
Lisserpeton bairdi n. gen., n. sp. Estes 1965
20 specimens
some specimens were uncatalogued
Scapherpeton tectum Cope 1876
Estes et al. 1969 61 specimens
Amphibia - Temnospondyli - Batrachosauroididae
Opisthotriton kayi Auffenberg 1961
Estes et al. 1969 107 specimens
Prodesmodon copei Estes 1964
Estes et al. 1969 2 specimens
Cuttysarkus mcnallyi Estes 1964
Estes et al. 1969 2 specimens
synonym of Prodesmodon copei
Amphibia - Temnospondyli - Albanerpetontidae
Albanerpetontidae indet. Fox and Naylor 1982
Gardner 2000 3 specimens
Albanerpeton nexuosus Estes 1981
Gardner 2000 1 specimen
Reptilia - Ceratopsidae
Triceratops sp. Marsh 1889
Estes et al. 1969
Reptilia - Pachycephalosauridae
? Pachycephalosauridae indet. Sternberg 1945
Estes et al. 1969
Reptilia
Thescelosaurus neglectus Gilmore 1913
Estes et al. 1969
    = Orodromeus makelai Horner and Weishampel 1988
Galton 1995
Reptilia - Hadrosauridae
Anatosaurus sp. Lull and Wright 1942
Estes et al. 1969
synonym of Edmontosaurus
Reptilia - Theropoda
Theropoda indet. Marsh 1881
Estes et al. 1969
Reptilia
Paronychodon lacustris Cope 1876
Estes et al. 1969
nomen dubium belonging to Paronychodon
Reptilia - Dromaeosauridae
? Dromaeosauridae indet. Colbert and Russell 1969
Estes et al. 1969
Reptilia - Coeluridae
? Coeluridae indet. Marsh 1881
Estes et al. 1969
Reptilia
Brachychampsa montana Gilmore 1911
Estes et al. 1969 1 specimen
Leidyosuchus sternbergi (Gilmore 1910)
Estes et al. 1969 1 specimen
recombined as Borealosuchus sternbergii
Champsosaurus sp. Cope 1876
Estes et al. 1969 2 specimens
Coniophis precedens Marsh 1892
Estes et al. 1969 1 specimen
nomen dubium belonging to Serpentes
Reptilia - Boidae
? Erycinae indet. Bonaparte 1825
Estes et al. 1969 1 specimen
Reptilia - Xantusiidae
Xantusiidae indet. Baird 1858
Gao and Fox 1996 1 specimen
UALVP 29840 (gen. et sp. nov.)
Reptilia - Cordylidae
? Cordylidae informal indet. B Gray 1837
Gao and Fox 1996 1 specimen
UALVP 29722 (dentary)
Reptilia - Globauridae
Contogenys sloani n. gen., n. sp. Estes 1969
Estes 1969 5 specimens
MCZ 3682-3684; OMNH 59221; UALVP 29839
Reptilia - Chamopsiidae
Chamops segnis Marsh 1892
Estes et al. 1969 12 specimens
Additional specimens from Gao & Fox (1996): UALVP 29724-29726, 29730, 29737-29738 (maxilla), and 29721, 29723, 29815 (dentaries)
Meniscognathus altmani Estes 1964
Gao and Fox 1996 4 specimens
UALVP 29775, 29779 (maxilla), and 29776, 29831 (dentaries)
Haptosphenus placodon Estes 1964
Estes et al. 1969 1 specimen
Leptochamops denticulatus (Gilmore 1928)
Estes et al. 1969 1 specimen
Peneteius aquilonius Estes 1969
Estes et al. 1969 1 specimen
Peneteius aquilonius n. gen., n. sp. Estes 1969
Estes 1969 1 specimen
MCZ 3612 (holotype dentary)
Reptilia - Xenosauridae
Exostinus lancensis Gilmore 1928
Estes et al. 1969 4 specimens
plus specimen reported by Gao & Fox (1996), UALVP 29919 (frontal)
Reptilia
Paraderma bogerti Estes 1964
Estes et al. 1969 1 specimen
Colpodontosaurus cracens Estes 1964
Estes et al. 1969 3 specimens
Gao & Fox (1996): UALVP 29785-29786
Reptilia - Varanidae
Palaeosaniwa cf. canadensis Gilmore 1928
Estes et al. 1969 1 specimen
MCZ 3665 (large vertebra)
    = Varanidae indet. Hardwicke and Gray 1824
Gao and Fox 1996
'lack of information on the morphologies of the condylar cup, precondylar constriction, and neural spine prevents a definite referral of this specimen to Palaeosaniwa'
Reptilia
Parasaniwa wyomingensis Gilmore 1928
Estes et al. 1969 11 specimens
plus Gao & Fox (1996) specimens: UALVP 33357-33360 (maxilla), 33361 and several unnumbered (dentaries), 33362, 33363 (pterygoids), 33364, 33365 (parietals)
Reptilia - Anguidae
Pancelosaurus piger (Gilmore 1928)
Estes et al. 1969 15 specimens
recombined as Odaxosaurus piger
Reptilia - Testudines - Emydidae
? Emydinae indet. Rafinesque 1815
Estes et al. 1969 3 specimens
Reptilia - Testudines - Trionychidae
Estes et al. 1969 1 specimen
    = Trionychidae indet. Gray 1825
Alroy 2006
Reptilia - Testudines - Adocidae
Adocus sp. Cope 1868
Estes et al. 1969 1 specimen
Reptilia
Compsemys victa (Leidy 1856)
Estes et al. 1969 1 specimen
Reptilia - Baenidae
Baenidae informal indet. undescribed genus and species 1 Cope 1882
Estes et al. 1969
Baenidae informal indet. 2 Cope 1882
Estes et al. 1969
"undescribed genus and species"
Palatobaena bairdi Gaffney 1972
Gaffney 1972 2 specimens
MCZ 3515 (right maxilla), and a specimen referred by Lyson & Joyce (2009): AMNH 8277 (complete lower jaws)
Eubaena cephalica (Hay 1904)
Gaffney 1972 6 specimens
MCZ 3530 (both parietals), AMNH 4948 (skull), MCZ 3510, 3511, 3512, 3519 (maxillae), MCZ 3525 (right parietal)
Acipenseriformes - Acipenseridae
Acipenser eruciferus (Cope 1876)
Estes et al. 1969 7 elements
Acipenseriformes - Polyodontidae
Paleopsephurus wilsoni MacAlpin 1947
Estes et al. 1969 3 elements
Actinopteri
Teleostei indet. Müller 1846
Estes et al. 1969
numerous vertebrae and fragments
Actinopteri - Elopiformes - Elopidae
Elopidae indet. Bonaparte 1846
Estes et al. 1969 1 element
Actinopteri - Elopiformes - Phyllodontidae
cf. Paralbula casei Estes 1969
Estes et al. 1969 1 element
Actinopteri - Albuliformes - Albulidae
Coriops amnicolus Estes 1969
Estes et al. 1969 3 elements
Actinopteri - Sciaenidae
Platacodon nanus Marsh 1889
Estes et al. 1969 2 elements
Actinopteri - Aspidorhynchiformes - Aspidorhynchidae
Belonostomus longirostris (Lambe 1902)
Estes et al. 1969 1 element
Actinopteri - Amiiformes - Palaeolabridae
Palaeolabrus montanensis Estes 1969
Estes et al. 1969 7 elements
Actinopteri - Amiiformes - Amiidae
Amia sp. Linnaeus 1766
Estes et al. 1969 1 element
not sure. perhaps Protamia
Amia fragosa Jordan 1927
Estes et al. 1969 94 elements
Actinopteri - Lepisosteiformes - Lepisosteidae
Lepisosteus occidentalis (Leidy 1856)
Estes et al. 1969 25 elements
original and current combination Lepidotus occidentalis
Chondrichthyes - Rajiformes - Rhinobatidae
Myledaphus bipartitus Cope 1876
Estes et al. 1969 1 element
Chondrichthyes - Hybodontiformes - Lonchidiidae
Lonchidion selachos Estes 1964
Estes et al. 1969 1 element
see common names

Geography
Country:United States State/province:Montana County:McCone County
Coordinates: 47.8° North, 106.1° West (view map)
Paleocoordinates:54.4° North, 79.0° West
Basis of coordinate:based on nearby landmark
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Maastrichtian 10 m.y. bin:Cretaceous 8
*Period:Cretaceous - Cretaceous *Epoch:Late/Upper Cretaceous - Late/Upper Cretaceous
*International age/stage:Maastrichtian - Maastrichtian
Key time interval:Late/Upper Maastrichtian
Age range of interval:72.10000 - 66.00000 m.y. ago
* legacy (obsolete) database fields
Stratigraphy
Formation:Hell Creek
Stratigraphic resolution:group of beds
Stratigraphy comments: upper 1/3 of formation, 80 ft below coal bed
Lithology and environment
Primary lithology:current ripples sandstone
Includes fossils?Y
Lithology description: "a cross-bedded stram channel"
Environment:coarse channel fill
Geology comments: 'Coastal plain deposits'
Taphonomy
Modes of preservation:body
Size of fossils:macrofossils,mesofossils
Preservation of anatomical detail:good
Disassociated minor elements:some
Collection methods and comments
Collection excludes:some macrofossils,some microfossils
Collection methods:surface (float),anthill,field collection
Reason for describing collection:taxonomic analysis
Collectors:Univ. of Minnesota Collection dates:1962-1964
Taxonomic list comments:mammal fauna said to be present
Metadata
Also known as:Bug Creek, BCA, OMNH V1173
Database number:13142
Authorizer:E. Fara, R. Benson, P. Mannion, M. Carrano, J. Alroy Enterer:E. Fara, M. Carrano, P. Mannion, R. Benson, J. Alroy
Modifier:G. Varnham Research group:vertebrate
Created:2002-01-23 07:05:23 Last modified:2022-02-01 07:35:55
Access level:group members Released:2002-04-23 07:05:23
Creative Commons license:CC BY
Reference information

Primary reference:

4377.5% 41840R. Estes. 1965. A new fossil salamander from Montana and Wyoming. Copeia 1965(1):90-95 [E. Fara/E. Fara/J. Alroy]

Secondary references:

18120 J. Alroy. 2006. Synonymies and reidentifications of North American fossil vertebrates. [J. Alroy/J. Alroy/J. Alroy]
38165 J. D. Archibald and J. H. Hutchison. 1979. Revision of the genus Palatobaena (Testudines, Baenidae), with the description of a new species. Postilla 177:1-19 [P. Mannion/P. Mannion/P. Holroyd]
37920 R. Estes. 1969. Relationships of two Cretaceous lizards (Sauria, Teiidae). Breviora 317:1-8 [R. Benson/R. Benson]
12167ETE R. Estes. 1969. A scincoid lizard from the Cretaceous and Paleocene of Montana. Breviora 331:1-9 [M. Carrano/M. Carrano/R. Benson]
43815% 40240R. Estes, P. Berberian, and C. A. M. Meszoely. 1969. Lower vertebrates from the Late Cretaceous Hell Creek Formation, McCone County, Montana. Breviora 337:1-33 [E. Fara/E. Fara/M. Carrano]
704 E. S. Gaffney. 1972. The systematics of the North American family Baenidae (Reptilia, Cryptodira). Bulletin of the American Museum of Natural History 147(5):245-312 [J. Alroy/M. Sommers/R. Benson]
12971ETE P. M. Galton. 1995. The species of the basal hypsilophodontid dinosaur Thescelosaurus Gilmore (Ornithischia: Ornithopoda) from the Late Cretaceous of North America. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 198(3):297-311 [M. Carrano/M. Carrano/M. Carrano]
37933 K. -Q. Gao and R. C. Fox. 1996. Taxonomy and evolution of Late Cretaceous lizards (Reptilia: Squamata) from western Canada. Bulletin of the Carnegie Museum of natural History 33:1-107 [R. Benson/R. Benson]
42675 J. D. Gardner. 2000. Albanerpetontid amphibians from the Upper Cretaceous (Campanian and Maastrichtian) of North America. Geodiversitas 22(3):349-388 [M. Carrano/M. Carrano/M. Carrano]
52044 J. D. Gardner. 2008. New information on frogs (Lissamphibia: Anura) from the Lance Formation (Maastrichtian) and Bug Creek Anthills (late Maastrichtian and early Paleocene), Hell Creek Formation, USA. In J. T. Sankey & S. Baszio (ed.), Vertebrate Microfossil Assemblages: Their Role in Paleoecology and Paleobiogeography 219-249 [M. Carrano/M. Carrano/M. Carrano]
44340 J. D. Gardner. 2012. Revision of Piceoerpeton MESZOELY (Caudata: Scapherpetontidae) and description of a new species from the late Maastrichtian and ? early Paleocene of western North America. Bulletin de la Société Géologique de France 183(6):611-620 [P. Mannion/P. Mannion]
52039 J. D. Gardner and M. Böhme. 2008. Review of the Albanerpetontidae (Lissamphibia), with comments on the paleoecological preferences of European Tertiary albanerpetontids. In J. T. Sankey & S. Baszio (ed.), Vertebrate Microfossil Assemblages: Their Role in Paleoecology and Paleobiogeography 178-218 [M. Carrano/M. Carrano]
38940 B. G. Naylor. 1983. New salamander (Amphibia: Caudata) atlantes from the Upper Cretaceous of North America. Journal of Paleontology 57(1):48-52 [R. Benson/R. Benson]
31563ETE R. L. Nydam and B. M. Fitzpatrick. 2009. The occurrence of Contogenys-like lizards in the Late Cretaceous and Early Tertiary of the Western Interior of the U.S.A. Journal of Vertebrate Paleontology 29(3):677-701 [M. Carrano/M. Carrano/P. Mannion]
53515 R. E. Sloan. 1976. The ecology of dinosaur extinction. In C. S. Churcher (ed.), Athlon: Essays on Palaeontology in Honor of Loris Shano Russell. Royal Ontario Museum, Life Sciences Miscellaneous Publications 134-154 [M. Carrano/M. Carrano]
53471 J. Smit and S. van der Kaars. 1984. Terminal Cretaceous extinctions in the Hell Creek area, Montana: compatible with catastrophic extinction. Science 223(4641):1177-1179 [M. Carrano/M. Carrano]