Zhinst Tolgoi, Udyn Sayr [HMNS-GIN]: Late/Upper Campanian, Mongolia
collected by HMNS-GIN 1995, 1998, 2000, 2005, 2007

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Mammalia - Multituberculata
Multituberculata indet. (Cope 1884)
Suzuki and Watabe 2000 1 individual
At outcrop no. 41 (S of no. 39), and lower jaw and postcranial elements of a Multituberculata, 980817 US Mammal LKH
Mammalia
? Eutheria indet. (Huxley 1880)
Suzuki and Watabe 2000 8 specimens
980818-21 US
Reptilia
Testudinata indet. (Oppel 1811)
Watabe and Tsogtbaatar 2004
turtle carapaces were found at southern outcrops (outcrop no. 37)
Reptilia - Testudines - Nanhsiungchelyidae
Nanhsiungchelyidae indet. Yeh 1966
Suzuki and Narmandakh 2004 2 specimens
fragmentary remains
cf. Basilemys sp. Hay 1902
Watabe and Tsogtbaatar 2004 1 individual
carapace
Reptilia
Crocodylia indet. (Owen 1842)
Watabe and Tsogtbaatar 2004 1 individual
000722 US SZK Croco; "an infant crocodile"
Dinosauria indet. (Owen 1842)
Suzuki and Watabe 2000 3 individuals
980819 US Egg Nest B-O, 000719 US FJ, 070626 US SZK; At outcrop no. 41 (S of no. 39), eggshells
Theropoda indet. (Marsh 1881)
Suzuki and Watabe 2000 1 individual
000722 US BON; "small theropod pelvis-hind limb specimen"; isolated bones - theropod digits
Reptilia - Elongatoolithidae
Nanhsiungoolithus sp. Zhao 1975
Reptilia - Dromaeosauridae
Velociraptor sp. Osborn 1924
Suzuki and Watabe 2000 1 specimen
forelimbs; teeth of Velociraptor: outcrops in the SE area of Udyn Sayr
Reptilia - Prismatoolithidae
Protoceratopsidovum sp. Mikhailov 1994
Suzuki and Watabe 2000 1 individual
950704 Zhinst SZK; nest with eggs
Reptilia - Ankylosauridae
Ankylosauridae indet. Brown 1908
Watabe and Tsogtbaatar 2004 3 individuals
000716 US ENKH-ANKLYO, 000717 US ENKH-ANKYLO, 000718 US BON; ankylosaur scutes and vertebrae, and a small skull of ankylosaurwere found at southern outcrops (outcrop no. 37)
? Pinacosaurus sp. Gilmore 1933
Suzuki and Watabe 2000 1 individual
partial skeleton, only incompletely excavated
Pinacosaurus sp. Gilmore 1933
Watabe and Tsogtbaatar 2004 1 individual
On July 16 the skeleton of a small individual ankylosaur (Pinacosaurus) found by Mongolian preparator Enkhbat in 1998 was excavated by the monolith method at the southern small outcrop of eolian deposits (no. 34)
Reptilia - Protoceratopsidae
Protoceratopsidae indet. Granger and Gregory 1923
Watabe and Tsogtbaatar 2004 1 specimen
000723 US BON; "a robust jaw of a protoceratopsid"; At outcrop no. 41 (S of no. 39), teeth and partial skeletons of a protoceratopsid
Bagaceratops sp. Maryanska and Osmólska 1975
Watabe and Tsogtbaatar 2004 1 individual
MPC-D 100/551B (= 000719 US FJM Baga)
    = cf. Bagaceratops sp. Maryanska and Osmólska 1975
Czepiński 2020
Protoceratops sp. Granger and Gregory 1923
Watabe and Suzuki 2000 10 specimens
MPC-D 100/540a-d (= 070626 US KWR); 950704 Udyn Sayr MTS-I & MTS-II, 980819 US Proto NAR; 980821 US SZK 9:05; 000720 US SZK Proto; 000720 US TSGT small Proto
Protoceratops andrewsi Granger and Gregory 1923
Handa et al. 2012 3 specimens
MPC-D 100/537, 100/539 (= 980821 US Proto ENKH?); MPC-D 100/551 (= 000720 US-Zhinst, GI SPS 100/588, and 100/528; formerly Bagaceratops)
Protoceratops cf. andrewsi Granger and Gregory 1923
Handa et al. 2012 1 specimen
MPC-D 100/538
Reptilia - Ovaloolithidae
Ovaloolithidae indet. Mikhailov 1991
Suzuki and Watabe 2000
eggs
Reptilia
Varanoidea indet. Gray 1827
Watabe and Tsogtbaatar 2004 1 individual
outcrop no. 52, isolated bones from the skeleton of a varanoid lizard
Lacertilia indet. (Owen 1842)
Watabe and Suzuki 2000 10 specimens
000725 US OTG LIZARD-1, LIZARD-2; skull of a lizard; a lizard skeleton with scales; 980818 US Lizard ENKH, 980818-21 US
see common names

Geography
Country:Mongolia State/province:Omnogov
Coordinates: 44.1° North, 102.9° East (view map)
Paleocoordinates:41.6° North, 92.0° East
Basis of coordinate:stated in text
Altitude:1327 meters
Geographic resolution:outcrop
Time
Period:Cretaceous Epoch:Late/Upper Cretaceous
Stage:Campanian 10 m.y. bin:Cretaceous 7
Key time interval:Late/Upper Campanian
Age range of interval:83.60000 - 72.10000 m.y. ago
Stratigraphy
Formation:Djadokhta
Stratigraphic resolution:group of beds
Stratigraphy comments: -upper horizon
-possibly Baruungoyot Fm. (Czepinksi 2020)
-Dashzeveg et al. 2005: The quick stratigraphic succession of normal and reversed magnetozones suggests, but does not clearly establish, that the sediments may have been deposited during the rapid sequence of polarity changes in the late part of the Campanian between about 75 to 71 Ma.
Lithology and environment
Primary lithology:"cross stratification",fine,nodular,red sandstone
Includes fossils?Y
Lithology description: "fine sandstone…characterized by large-scaled, steeply-dipping cross-stratification". "Fossiliferous beds are made of fine sandstone including many calcarous nodules"
Environment:dune
Taphonomy
Modes of preservation:body,concretion
Size of fossils:macrofossils,mesofossils
Spatial resolution:parautochthonous
Collection methods and comments
Collection methods:surface (in situ),mechanical,field collection
Reason for describing collection:general faunal/floral analysis
Collectors:HMNS-GIN Collection dates:1995, 1998, 2000, 2005, 2007
Collection method comments: HMNS-Hayashibara Museum of Natural Sciences
Metadata
Also known as:Udan Sayr,
Database number:127678
Authorizer:R. Benson, M. Carrano Enterer:R. Benson, M. Carrano
Modifier:M. Carrano Research group:vertebrate
Created:2012-05-22 00:42:44 Last modified:2022-07-29 14:46:55
Access level:the public Released:2012-05-22 00:42:44
Creative Commons license:CC BY
Reference information

Primary reference:
76473. Y. Ariunchimeg. 1997. Results of studies of dinosaur eggs. Abstract of Report Meeting, Mongolia-Japan Joint Paleontological Expedition 13 [M. Carrano/M. Carrano]

Secondary references:

82280 Ł Czepiński. 2020. New protoceratopsid specimens improve the age correlation of the Upper Cretaceous Gobi Desert strata. Acta Palaeontologica Polonica 65(3):481-497 [M. Carrano/M. Carrano/M. Carrano]
15459ETE D. Dashzeveg, L. Dingus, D.B. Loope, C.C. Swisher, III, T. Dulam and M.R. Sweeney. 2005. New stratigraphic subdivision, depositional environment, and age estimate for the Upper Cretaceous Djadokhta Formation, southern Ulan Nur Basin, Mongolia. American Museum Novitates (3498)1-31 [M. Carrano/K. Maguire/G. Varnham]
50510 N. Handa, M. Watabe, and K. Tsogtbaatar. 2012. New specimens of Protoceratops (Dinosauria: Neoceratopsia) from the Upper Cretaceous in Udyn Sayr, southern Gobi area, Mongolia. Paleontological Research 16(3):179-198 [M. Carrano/M. Carrano]
82301 Y. Matsumoto, R. Hashimoto, and T. Sonoda. 2000. Report of preparation works for Mongolian specimens in HMNS from March 1994 to December 1998. Hayashibara Museum of Natural Sciences Research Bulletin 1:113-127 [M. Carrano/M. Carrano]
82304 Y. Matsumoto, R. Hashimoto, T. Sonoda, Y. Fujiyama, B. Mifune, Y. Kawahara, and M. Saneyoshi. 2010. Report of the preparation works for Mongolian specimens in Hayashibara Museum of Natural Sciences: 1999–2008. Hayashibara Museum of Natural Sciences Research Bulletin 3:167-185 [M. Carrano/M. Carrano/M. Carrano]
55067 M. Saneyoshi, M. Watabe, T. Tsubamoto, K. Tsogtbaatar, T. Chinzorig and S. Suzuki. 2010. Report of the HMNS-MPC Joint Paleontological Expedition in 2007. Hayashibara Museum of Natural Sciences Research Bulletin 3:19-28 [M. Carrano/M. Carrano]
41670 S. Suzuki and P. Narmandakh. 2004. Change of the Cretaceous turtle faunas in Mongolia. Hayashibara Museum of Natural Sciences Research Bulletin 2:7-14 [R. Benson/R. Benson]
42292 S. Suzuki and M. Watabe. 2000. Report on the Japan-Mongolia Joint Paleontological Expedition to the Gobi desert, 1995. Hayashibara Museum of Natural Sciences Research Bulletin 1:45-57 [R. Benson/R. Benson/M. Carrano]
42293 S. Suzuki and M. Watabe. 2000. Report on the Japan–Mongolia Joint Paleontological Expedition to the Gobi desert, 1998. Hayashibara Museum of Natural Sciences Research Bulletin 1:83-98 [R. Benson/R. Benson/M. Carrano]
42154 M. Watabe and S. Suzuki. 2000. Cretaceous fossil localities and a list of fossils collected by the Hayashibara Museum of Natural Sciences and Mongolian Paleontological Center Joint Paleontological Expedition (JMJPE) from 1993 through 1998. Hayashibara Museum of Natural Sciences Research Bulletin 1:99-108 [R. Benson/R. Benson]
42294 M. Watabe and K. Tsogtbaatar. 2004. Report on the Japan–Mongolia Joint Paleontological Expedition to the Gobi desert, 2000. Hayashibara Museum of Natural Sciences Research Bulletin 2:45-67 [R. Benson/R. Benson/M. Carrano]