Ben Guerir, couche 6: Early/Lower Maastrichtian, Morocco
List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
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unclassified
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Duwibatis sp.
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Cappetta et al. 2014 | |||||||||
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Actinopteri
- Pycnodontiformes
- Pycnodontidae
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Pycnodontidae indet.
(Agassiz 1833)
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Cappetta et al. 2014 | |||||||||
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Actinopteri
- Aulopiformes
- Enchodontidae
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Enchodus elegans
Dartevelle and Casier 1949
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Cappetta et al. 2014 | |||||||||
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Actinopteri
- Aulopiformes
- Dercetidae
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Stratodus apicalis
Cope 1872
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Cappetta et al. 2014 | |||||||||
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Actinopteri
- Tetraodontiformes
- Eotrigonodontidae
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Stephanodus libycus
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Cappetta et al. 2014 | |||||||||
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Actinopteri
- Elopiformes
- Phyllodontidae
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Pseudoegertonia sp.
Dartevelle and Casier 1949
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Cappetta et al. 2014 | |||||||||
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Reptilia
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Eusuchia indet.
Huxley 1875
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Cappetta et al. 2014 | |||||||||
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Reptilia
- Dyrosauridae
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Dyrosauridae indet.
Stefano 1903
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Cappetta et al. 2014 | |||||||||
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Reptilia
- Plesiosauria
- Elasmosauridae
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Elasmosauridae indet.
Cope 1869
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Cappetta et al. 2014 | |||||||||
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Reptilia
- Pachyvaranidae
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Pachyvaranus crassispondylus
Arambourg 1952
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Houssaye et al. 2011 | 1 specimen | ||||||||
| OCP.DEK/GE (vertebrae) | ||||||||||
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Reptilia
- Mosasauridae
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"Platecarpus" ptychodon
Arambourg 1952
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Cappetta et al. 2014 | |||||||||
| nomen dubium belonging to Plioplatecarpinae | ||||||||||
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Eremiasaurus heterodontus
LeBlanc et al. 2012
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Cappetta et al. 2014 | |||||||||
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Globidens phosphaticus
Bardet et al. 2005
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Bardet et al. 2005 | 7 specimens | ||||||||
| OCP.DEK/GE 347 (median tooth); OCP.DEK/GE unnumbered (6 teeth) | ||||||||||
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Prognathodon currii
Christiansen and Bonde 2002
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Bardet et al. 2005 | 1 specimen | ||||||||
| OCP.DEK/GE (one tooth) | ||||||||||
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Prognathodon giganteus
Dollo 1889
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Cappetta et al. 2014 | |||||||||
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Halisaurus arambourgi
Bardet et al. 2005
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Cappetta et al. 2014 | |||||||||
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Reptilia
- Testudines
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Chelonioidea indet.
Baur 1893
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Cappetta et al. 2014 | |||||||||
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Reptilia
- Testudines
- Bothremydidae
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Bothremydidae indet.
Baur 1891
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Cappetta et al. 2014 | |||||||||
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Chondrichthyes
- Myliobatiformes
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Myliobatoidea informal indet. 1
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Noubhani and Cappetta 1997 | |||||||||
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Myliobatoidea informal indet. 2
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Noubhani and Cappetta 1997 | |||||||||
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Chondrichthyes
- Myliobatiformes
- Rhombodontidae
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Rhombodus meridionalis
Arambourg 1952
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Cappetta et al. 2014 | |||||||||
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Rhombodus meridionalis
Arambourg 1952
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Rhombodus binkhorsti
Dames 1881
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Rhombodus microdon
Arambourg 1952
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Cappetta et al. 2014 | |||||||||
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Chondrichthyes
- Myliobatiformes
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Dasyatoidea indet.
Whitley 1940
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Cappetta et al. 2014 | |||||||||
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Coupatezia sp.
Cappetta 1982
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Cappetta et al. 2014 | |||||||||
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Coupatezia ambroggii
Noubhani and Cappetta 1997
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Noubhani and Cappetta 1997 | |||||||||
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Chondrichthyes
- Rajiformes
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Rhinobatoidei indet.
Fowler 1941
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Cappetta et al. 2014 | |||||||||
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Chondrichthyes
- Rajiformes
- Rhinobatidae
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"Rhinobatos" informal sp. 1
Link 1790
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Cappetta et al. 2014 | |||||||||
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"Rhinobatos" informal sp. 5
Link 1790
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Cappetta et al. 2014 | |||||||||
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"Rhinobatos" aff. primarmatus
Woodward 1889
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Cappetta et al. 2014 | |||||||||
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Chondrichthyes
- Rajiformes
- Parapalaeobatidae
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Parapalaeobates atlanticus
Arambourg 1952
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Cappetta et al. 2014 | |||||||||
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Chondrichthyes
- Rajiformes
- Sclerorhynchidae
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Ganopristis leptodon
Arambourg 1935
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Cappetta et al. 2014 | |||||||||
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Dalpiazia stromeri
Checchia-Rispoli 1933
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Cappetta et al. 2014 | |||||||||
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Ctenopristis nougareti
Arambourg 1940
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Cappetta et al. 2014 | |||||||||
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Chondrichthyes
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Hypsobatis weileri
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Cappetta et al. 2014 | |||||||||
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Youssoubatis sp.
Cappetta 1992
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Cappetta et al. 2014 | |||||||||
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Chondrichthyes
- Orectolobiformes
- Hemiscylliidae
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Chiloscyllium informal sp. 1
Müller and Henle 1837
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Cappetta et al. 2014 | |||||||||
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Chiloscyllium informal sp. 2
Müller and Henle 1837
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Cappetta et al. 2014 | |||||||||
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Chondrichthyes
- Orectolobiformes
- Ginglymostomatidae
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Plicatoscyllium gharbii n. sp.
Noubhani and Cappetta 1997
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Noubhani and Cappetta 1997 | |||||||||
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Chondrichthyes
- Carcharhiniformes
- Scyliorhinidae
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Pteroscyllium sp.
Cappetta 1980
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Cappetta et al. 2014 | |||||||||
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Chondrichthyes
- Carcharhiniformes
- Triakidae
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Squatigaleus atlasi
Cappetta 1989
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Noubhani and Cappetta 1997 | |||||||||
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Chondrichthyes
- Lamniformes
- Mitsukurinidae
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Scapanorhynchus rapax
Quaas 1902
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Cappetta et al. 2014 | |||||||||
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Chondrichthyes
- Lamniformes
- Otodontidae
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Cretolamna aff. maroccana
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Cappetta et al. 2014 | |||||||||
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Cretolamna sp.
Glikman 1958
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Cappetta et al. 2014 | |||||||||
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Cretolamna biauriculata
(Wanner 1902)
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Cappetta et al. 2014 | |||||||||
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Chondrichthyes
- Lamniformes
- Aquilolamnidae
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Cretomanta sp.
Case et al. 1990
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Cappetta et al. 2014 | |||||||||
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Chondrichthyes
- Lamniformes
- Serratolamnidae
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Serratolamna khderii
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Cappetta et al. 2014 | |||||||||
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Chondrichthyes
- Lamniformes
- Anacoracidae
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Squalicorax africanus
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Cappetta et al. 2014 | |||||||||
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Squalicorax bassanii
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Cappetta et al. 2014 | |||||||||
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Squalicorax informal nov. sp. 2
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Cappetta et al. 2014 | |||||||||
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= Squalicorax benguerirensis
Cappetta et al. 2014
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Cappetta et al. 2014 | |||||||||
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Squalicorax pristodontus
(Agassiz 1835)
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Cappetta et al. 2014 | |||||||||
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Squalicorax informal nov. sp. 1
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Cappetta et al. 2014 | |||||||||
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= Squalicorax microserratus
Cappetta et al. 2014
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Cappetta et al. 2014 | |||||||||
| see common names | ||||||||||
Geography
| Country: | Morocco |
| Coordinates: | 32.3° North, 7.8° West (view map) |
| Paleocoordinates: | 24.4° North, 8.4° West |
| Basis of coordinate: | estimated from map |
| Geographic resolution: | outcrop |
Time
| Period: | Cretaceous | Epoch: | Late/Upper Cretaceous |
| Stage: | Maastrichtian | 10 m.y. bin: | Cretaceous 8 |
| Key time interval: | Early/Lower Maastrichtian | ||
| Age range of interval: | 72.10000 - 66.00000 m.y. ago |
Stratigraphy
| Stratigraphic resolution: | group of beds | ||||
| Stratigraphy comments: Couche VI (or level C6). According to Cappetta et al. (2014): The use of dissimilarity indices and agglomerative method underscores two distinct associations: a lower one including the levels L6 and L5, and an upper one comprising the levels L4 to L2. These two associations allow to separate a lower and an upper Maastrichtian level. Moreover, the special morphology of the teeth of S. pristodontus occurring in L2, compared to what is observed in lower levels, permits us to distinguish a lower Upper Maastrichtian and an upper Upper Maastrichtian faunal assemblage. | |||||
Lithology and environment
| Primary lithology: | phosphatic poorly lithified marl |
| Includes fossils? | Y |
| Lithology description: soft phosphatic beds | |
| Environment: | carbonate indet. |
Taphonomy
| Modes of preservation: | body |
| Size of fossils: | macrofossils |
| Preservation of anatomical detail: | good |
| Spatial resolution: | parautochthonous |
Collection methods and comments
| Collection methods: | field collection |
| Reason for describing collection: | general faunal/floral analysis |
| Museum repositories: | MNHN |
| Collection method comments: Repository: University of Montpellier 2; Office Chérifien des Phosphates, Khouribga, Morocco (OCP) | |
Metadata
| Database number: | 116903 | ||
| Authorizer: | P. Mannion, M. Clapham | Enterer: | P. Mannion, M. Clapham |
| Modifier: | M. Clapham | Research group: | vertebrate |
| Created: | 2011-09-14 00:28:34 | Last modified: | 2020-10-17 18:07:54 |
| Access level: | the public | Released: | 2011-09-14 00:28:34 |
| Creative Commons license: | CC BY |
Reference information
Primary reference:
| 74121. | A. Noubhani and H. Cappetta. 1994. Révision des Rhombodontidae (Neoselachii, Batomorphi) des bassins à phosphate du Maroc. Palaeovertebrata 23:1-49 [M. Clapham/M. Clapham] |
Secondary references:
| 37608 | N. Bardet, X. Pereda Suberbiola, M. Iarochène, M. Amalik, and B. Bouya. 2005. Durophagous Mosasauridae (Squamata) from the Upper Cretaceous phosphates of Morocco, with description of a new species of Globidens. Netherlands Journal of Geosciences — Geologie en Mijnbouw 84(3):167-175 [P. Mannion/P. Mannion] | |
| 73775 | H. Cappetta, S. Adnet, D. Akkrim and M. Amalik. 2014. New Squalicorax species (Neoselachii: Lamniformes) from the Lower Maastrichtian of Ganntour phosphate deposit, Morocco. Palaeovertebrata 38(3):e3 [M. Clapham/M. Clapham] | |
| 73776 | H. Cappetta, N. Bardet, X. Pereda Suberbiola, S. Adnet, D. Akkrim, M. Amalik, and A. Benabdallah. 2014. Marine vertebrate faunas from the Maastrichtian phosphates of Benguérir (Ganntour Basin, Morocco): Biostratigraphy, palaeobiogeography and palaeoecology. Palaeogeography, Palaeoclimatology, Palaeoecology 409:217-238 [M. Clapham/M. Clapham] | |
| 38134 | A. Houssaye, N. Bardet, J.-C. Rage, X. Pereda-Suberbiola, B. Bouya, M. Amaghzaz, and M. Amalik. 2011. A review of Pachyvaranus crassispondylus Arambourg, 1952, a pachyostotic marine squamate from the latest Cretaceous phosphates of Morocco and Syria. Geological Magazine 148(2):237-249 [P. Mannion/P. Mannion] | |
| 73972 | A. Noubhani and H. Cappetta. 1997. Les Orectolobiformes, Carcharhiniformes et Myliobatiformes (Elasmobranchii, Neoselachii) des bassins à phosphate du Maroc (Maastrichtien-Lutétien basal). Systématique, biostratigraphie, évolution et dynamique des faunes. Palaeo Ichthyologica 8:1-327 [M. Clapham/M. Clapham] |