PBDB Collection

Bentiaba (layer above Bench 19): Early/Lower Maastrichtian, Angola
collected 2005 -

List of taxa

Where & when

Geology

Taphonomy & methods

Metadata & references

Taxonomic list

unclassified

	Mollusca spp.	Mateus et al. 2012
Osteichthyes

	Osteichthyes spp.	Mateus et al. 2012
Reptilia - Mosasauridae

	Globidens phosphaticus Bardet et al. 2005	Polcyn et al. 2010
	PA24 (a partial skeleton including many elements of the skull and some postcrania) and PA23 (a fragmentary skull and posterior dorsal vertebra)

	Prognathodon kianda n. sp. Schulp et al. 2008	Schulp et al. 2008
	MGUAN-PA129 (holotype), MGUAN-PA128, MGUAN-PA149, MGUAN-PA150, MGUAN-PA151 - several specimens of different ontogenetic ages; shed teeth, isolated bones, partial skulls and skeletons

	Liodon sp. (Owen 1842)	Mateus et al. 2012
	original and current combination Leiodon

	Carinodens belgicus (Woodward 1891)	Schulp et al. 2013
	recombined as Compressidens belgicus
	MGUAN-PA 171

	cf. Platecarpus ptychodon Arambourg 1952	Mateus et al. 2012
	nomen dubium belonging to Plioplatecarpinae
	MGUAN-PA160, two partial skulls and skeletons including limb material, isolated tooth crowns [new genus not erected but required]

	Phosphorosaurus sp. Dollo 1889	Mateus et al. 2012
	MGUAN-PA52, isolated partial frontal

	Halisaurus sp. Marsh 1869	Mateus et al. 2012
	MGUAN-PA18, two partial skeletons including many elements of the skulls, vertebrae, ribs and limbs
Reptilia - Testudines - Protostegidae

	Protostega sp. Cope 1871	Mateus et al. 2012
	MGUAN-PA158, two humeri, three costal plates, one xiphiplastron, one peripheral plate (single animal)

	cf. Calcarichelys sp. Zangerl 1953	Mateus et al. 2012
	MGUAN-PA167, neural scute
Reptilia - Testudines

	Toxochelys sp. Cope 1873	Mateus et al. 2012
	MGUAN-PA168, hyoplastron, peripherals and costal plates

	Euclastes sp. Cope 1870	Mateus et al. 2012
	MGUAN-PA157, complete skull and mandible, cervical vertebrae, peripheral plates, forelimb
Reptilia - Plesiosauria - Elasmosauridae

	Elasmosauridae indet. Cope 1869	Mateus et al. 2012	2 specimens
	MGUAN-PA113, basicranium, and MGUAN-PA126, symphyseal region of the mandibl, various unnumbered specimens, teeth

	cf. Tuarangisaurus sp. Wiffen and Moisley 1986	Mateus et al. 2012	1 specimen
	MGUAN-PA106
Reptilia

	Dinosauria indet. (Owen 1842)	Mateus et al. 2012
	MGUAN-PA175, distal part of humerus

	Hadrosauroidea indet. (Huene 1954)	Mateus et al. 2012
	MGUAN-PA176, a single phalanx

	Pterosauria indet. (Kaup 1834)	Fernandes et al. 2022
	MGUAN-PA652, 653, 656-659

	Pterodactyloidea indet. (Plieninger 1901)	Fernandes et al. 2022
	MGUAN-PA660

	Ornithocheiroidea indet. (Seeley 1870)	Mateus et al. 2012
	MGUN-PA163, left femur
	= Pteranodontia indet. Marsh 1876	Fernandes et al. 2022
	MGUAN-PA163, MGUAN-PA654

	Epapatelo otyikokolo n. gen., n. sp. Fernandes et al. 2022	Fernandes et al. 2022
	MGUAN-PA650 - holotype; referred material: MGUAN-PA661
Chondrichthyes

	Chondrichthyes spp.	Mateus et al. 2012
see common names

Geography

Country:	Angola	State/province:	Namibe
Coordinates:	14.3° South, 12.4° East (view map)
Paleocoordinates:	25.0° South, 1.7° East
Basis of coordinate:	based on nearby landmark
Geographic resolution:	small collection

Time

Period:	Cretaceous	Epoch:	Late/Upper Cretaceous
Stage:	Maastrichtian	10 m.y. bin:	Cretaceous 8
Key time interval:	Early/Lower Maastrichtian
Age range of interval:	72.10000 - 66.00000 m.y. ago

Stratigraphy

Formation:	Mocuio
Stratigraphic resolution:	group of beds

Lithology and environment

Primary lithology:	fine siltstone
Includes fossils?	Y
Lithology description: fine-grained sandstone
Environment:	marginal marine indet.

Taphonomy

Modes of preservation:	body
Size of fossils:	macrofossils

Collection methods and comments

Collection methods:	field collection
Reason for describing collection:	taxonomic analysis
Collection dates:	2005 - present
Taxonomic list comments:not the same as a previous Bentiaba = San Nicolau collection reported by Antunes 1964

Metadata

Also known as:	Sao Nicolau
Database number:	106705
Authorizer:	J. Alroy, P. Mannion	Enterer:	J. Alroy, P. Mannion, J. Tennant
Modifier:	P. Mannion	Research group:	vertebrate
Created:	2011-03-28 14:03:24	Last modified:	2023-05-09 05:11:30
Access level:	the public	Released:	2011-03-28 14:03:24
Creative Commons license:	CC BY

Reference information

Primary reference:

48111.

M. T. Antunes. 1964. O Neocretácico e o Cenozoico do litoral de Angola. Junta de Investigações do Ultramar 1-254 [M. Uhen/M. Uhen/M. Uhen]

Secondary references:

84671		A. E. Fernandes, O. Mateus, B. Andres, M. J. Polcyn, A. S. Schulp, A. O. Gonçalves, and L. L. Jacobs. 2022. Pterosaurs from the Late Cretaceous of Angola. Diversity 14:741:1-27 [P. Mannion/P. Mannion]
45398		O. Mateus, M. J. Polcyn, L. L. Jacobs, R. Arujo, A. S. Schulp, J. Marinheiro, B. Pereira and D. P. Vineyard. 2012. Cretaceous amniotes from Angola: dinosaurs, pterosaurs, mosasaurs, plesiosaurs, turtles. Actas de V Jornadas Internacionales sobre Paleontologia de Dinosaurios y su Entorno, Salas de los Infantes, Burgos 71-105 [P. Mannion/J. Tennant/O. Alfaro]
37606		M. J. Polcyn, L. L. Jacobs, A. S. Schulp and O. Mateus. 2010. The North African Mosasaur Globidens phosphaticus from the Maastrichtian of Angola. Historical Biology 22(1-3):175-185 [P. Mannion/P. Mannion/P. Mannion]
35655		A. S. Schulp, M. J. Polcyn, O. Mateus, L. L. Jacobs, and M. L. Morais. 2008. A new species of Prognathodon (Squamata, Mosasauridae) from the Maastrichtian of Angola, and the affinities of the mosasaur genus Liodon. Proceedings of the Second Mosasaur Meeting 1-12 [J. Alroy/J. Alroy]
76002		A. S. Schulp, M. J. Polcyn, O. Mateus and L. L. Jacobs. 2013. Two rare mosasaurs from the Maastrichtian of Angola and the Netherlands. Netherlands Journal of Geosciences 92(1):3-10 [P. Mannion/P. Mannion]