Kupferzell bonebed: Longobardian, Germany
collected by J. G. Wegele 1977

List of taxa
Where & when
Geology
Taphonomy & methods
Metadata & references
Taxonomic list
Coelacanthimorpha
Actinistia indet. (Cope 1870)
Schoch 1999
Dipnoi - Ptychoceratodontidae
Ptychoceratodus serratus (Agassiz 1838)
Schoch 1999
Bystrowianidae
Chroniosuchia indet. Tatarinov 1972
Schoch 1999 1 element
SMNS 81698
    = Bystrowiella schumanni Witzmann et al. 2008
Witzmann et al. 2008
Gomphodontidae
Gomphodontidae indet.
Schoch 1999
Reptilia - Owenettidae
Owenettidae indet. Broom 1939
Martinelli et al. 2016
SMNS 92101, humerus; Barasaurus?
Reptilia - Eosauropterygia - Nothosauridae
Nothosaurus sp. Münster 1834
    = Nothosaurus informal new species
Schoch 1999
Reptilia - Eosauropterygia - Pachypleurosauridae
Neusticosaurus cf. pusillus (Fraas 1881)
Schoch 1999
Reptilia
Rauisuchia indet. Huene 1942
    = Batrachotomus kupferzellensis n. gen., n. sp. Gower 1999
Gower 1999
SMNS 52970 (holotype), partial skeleton. Isolated cranial and postcranial remains of multiple additional individuals
Jaxtasuchus salomoni Schoch and Sues 2014
Schoch and Sues 2014 2 elements
SMNS 90067, 90068, paramedian osteoderms
Reptilia - Tanystropheidae
Tanystropheus sp. von Meyer 1852
    = Tanystropheus cf. conspicuus Meyer 1855
Schoch 1999
Reptilia
Lepidosauria indet. (Haeckel 1866)
Schoch 1999
Amphibia - Temnospondyli
Mastodonsaurus sp. Jaeger 1828
    = Mastodonsaurus giganteus Jaeger 1828
Schoch 1999
SMNS 54675, 54677, 54678, 80249, 80704, 80878, 80887, 80889, 80890, 80905, 80913, 81310, 81368, numerous additional specimens
Amphibia - Temnospondyli - Mastodonsauridae
Kupferzellia wildi n. gen., n. sp. Schoch 1997
Schoch 1997 5 specimens
recombined as Tatrasuchus wildi
SMNS 54670, complete skull. SMNS 54671-54674
Amphibia - Temnospondyli
Almasauridae indet. Dutuit 1972
Schoch 1999 7 elements
"new species"; SMNS 81713, isolated articular; SMNS 84115–84118, 90521, isolated intercentra; SMNS 84119, atlas
    = Callistomordax kugleri Schoch 2008
Schoch 2008
Amphibia - Temnospondyli - Trematosauridae
Bukobaja cf. enigmatica Ochev 1966
Schoch 1999
    = Trematolestes hagdorni Schoch 2006
Schoch 2006
Amphibia - Temnospondyli - Plagiosauridae
Plagiosuchus pustuliferus (Fraas 1896)
Schoch 1999
Gerrothorax informal new species
Schoch 1999
Actinopteri - Semionotiformes
Serrolepis sp. Quenstedt 1852
Schoch 1999
Palaeonisciformes - Palaeoniscidae
Gyrolepis sp. Agassiz 1833
Schoch 1999
Chondrichthyes - Hybodontiformes - Hybodontidae
Hybodus sp. Agassiz 1834
Schoch 1999
Chondrichthyes - Hybodontiformes - Acrodontidae
Acrodus sp. Agassiz 1834
Schoch 1999
see common names

Geography
Country:Germany State/province:Baden-Württemberg County:Hohenlohekreis
Coordinates: 49.2° North, 9.7° East (view map)
Paleocoordinates:16.3° North, 17.3° East
Basis of coordinate:stated in text
Geographic resolution:outcrop
Time
Period:Triassic Epoch:Middle Triassic
Stage:Ladinian 10 m.y. bin:Triassic 2
Key time interval:Longobardian
Age range of interval:239.70000 - 237.00000 m.y. ago
Stratigraphy
Geological group:Keuper Formation:Erfurt Member:Upper Lettenkeuper
Stratigraphic resolution:group of beds
Stratigraphy comments: Collected from the "Untere Graue Mergel" of the Upper Lettenkeuper, now formally the Erfurt Formation of the Lower Keuper (see Deutsche Stratigraphische Kommission 2005: Cour. Forsch.-Inst. Senckenberg 253:1-296). This locality is Longobardian (late Ladinian) in age. Fossils collected from a 30 cm thick unit. Schoch (1999: fig. 5) provides a stratigraphic section through the Kupferzell section.
Lithology and environment
Primary lithology:green,yellow marl
Secondary lithology: muddy dolomite
Includes fossils?Y
Includes fossils?Y
Lithology description: Lower layer of the quarry consists of greenish marls (most fossils). This is overlain by yellow to grey marls with carbonate nodules and then dolomites with marl and clay intercalations (Schoch 1999)
Environment:estuary/bay
Taphonomy
Modes of preservation:body
Lagerst�tten type:concentrate
Degree of concentration:-bonebed
Size of fossils:macrofossils
Preservation of anatomical detail:good
Abundance in sediment:abundant
Fragmentation:occasional
Temporal resolution:snapshot
Spatial resolution:parautochthonous
Collection methods and comments
Collection methods:selective quarrying,mechanical,survey of museum collection
Reason for describing collection:taxonomic analysis
Museum repositories:SMNS
Collectors:J. G. Wegele Collection dates:1977
Metadata
Also known as:Kupferzell-Bauersbach
Database number:101873
Authorizer:R. Butler, A. Dunhill, M. Carrano Enterer:R. Butler, M. Carrano, B. Allen
Modifier:M. Carrano Research group:vertebrate
Created:2011-01-04 19:44:20 Last modified:2019-04-10 14:03:48
Access level:the public Released:2011-01-04 19:44:20
Creative Commons license:CC BY
Reference information

Primary reference:
68549. R. Wild. 1980. Die Saurierfunde von Kupferzell [The reptile finds of Kupferzell]. Schwäbische Heimat 31:110-117 [M. Carrano/M. Carrano]

Secondary references:

34639 D. J. Gower. 1999. The cranial and mandibular osteology of a new rauisuchian archosaur from the Middle Triassic of southern Germany. Stuttgarter Beiträge zur Naturkunde Serie B (Geologie und Paläontologie) 280:1-49 [R. Butler/R. Butler]
34641 D. J. Gower and R. R. Schoch. 2009. Postcranial anatomy of the rauisuchian archosaur Batrachotomus kupferzellensis. Journal of Vertebrate Paleontology 29:103-122 [R. Butler/R. Butler]
66459 A. G. Martinelli, M. Bento Soares, and R. R. Schoch. 2016. Owenettids and procolophonids from the lower Keuper shed new light on the diversity of parareptiles in the German Middle Triassic. Journal of Paleontology 90(1):92-101 [A. Dunhill/B. Allen]
35492 R. R. Schoch. 1997. A new capitosaur amphibian from the Upper Lettenkeuper (Triassic: Ladinian) of Kupferzell (Southern Germany). Neues Jahrbuch für Geologie und Paläontologie 203(2):239-272 [R. Butler/R. Butler]
33823 R. R. Schoch. 1999. Comparative osteology of Mastodonsaurus giganteus (Jaeger, 1828) from the Middle Triassic (Lettenkeuper: Longobardian) of Germany (Baden-Württemberg, Bayern, Thüringen). Stuttgarter Beiträge zur Naturkunde Serie B (Geologie und Paläontologie) 278:1-170 [R. Butler/R. Butler]
34642 R. R. Schoch. 2008. A new stereospondyl from the German Middle Triassic, and the origin of the Metoposauridae. Zoological Journal of the Linnean Society 152:79-113 [R. Butler/R. Butler]
47030 R. R. Schoch and H.-D. Sues. 2014. A new archosauriform reptile from the Middle Triassic (Ladinian) of Germany. Journal of Systematic Palaeontology 12(1):113-131 [R. Butler/R. Butler/M. Carrano]
35488 R. S. Schoch. 2006. A complete trematosaurid amphibian from the Middle Triassic of Germany. Journal of Vertebrate Paleontology 26(1):29-43 [R. Butler/R. Butler]
34640 F. Witzmann, R. R. Schoch, and M. W. Maisch. 2008. A relict basal tetrapod from Germany: first evidence of a Triassic chroniosuchian outside Russia. Naturwissenschaften 95:67-72 [R. Butler/R. Butler]