Classification of Milstead 1956

G. Terrapene Merrem 1820 [box turtle]
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Invalid names: Terrapene antipex Hay 1916 [synonym], Terrapene bulverda Hay 1920 [synonym], Terrapene formosa Hay 1916 [synonym], Terrapene innoxia Hay 1916 [synonym], Terrapene llanensis Oelrich 1953 [synonym], Terrapene singletoni Gilmore 1927 [synonym]
Terrapene carolina Linnaeus 1758 [common box turtle]
Terrapene coahuila Schmidt and Owens 1944
†Terrapene eurypygia Cope 1870
Terrapene klauberi Bogert 1943
†Terrapene marnochi Cope 1877
Terrapene nelsoni Stejneger 1925
Terrapene ornata Agassiz 1857 [Western box turtle]
†Terrapene putnami Hay 1906 [box turtle]
†Terrapene whitneyi Hay 1916
G. Testudo Linnaeus 1758
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Subg. †Testudo (Hesperotestudo) Williams 1950
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†Testudo wilsoni Milstead 1956
Testudo carolina Linnaeus 1758 [common box turtle]
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Testudo carolina bauri Taylor 1894 [Florida box turtle]
Testudo carolina carolina Linnaeus 1758 [Eastern box turtle]
Testudo carolina major Agassiz 1857 [Gulf Coast box turtle]
Testudo carolina triunguis Agassiz 1857 [three-toed box turtle]