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Hesperotestudo

Reptilia - Testudines - Testudinidae

Synonyms
Synonymy list
YearName and author
1877Eupachemys Leidy
1902Eupachemys Hay p. 448
1950Testudo (Hesperotestudo) Williams
1953Testudo tedwhitei Williams p. 537
1953Testudo (Hesperotestudo) Williams p. 542
1956Testudo (Hesperotestudo) Milstead p. 168
1957Testudo tedwhitei Ray
1962Geochelone (Hesperotestudo) Auffenberg p. 628
1963Geochelone (Caudochelys) Auffenberg p. 69
1963Geochelone (Caudochelys) tedwhitei Auffenberg p. 80
1963Geochelone (Hesperotestudo) Auffenberg p. 81
1971Geochelone (Hesperotestudo) Auffenberg p. 115
1992Hesperotestudo Hutchison
1995Geochelone (Hesperotestudo) Parris and Daeschler p. 567
1996Hesperotestudo Hutchison p. 345
2000Hesperotestudo Meylan and Sterrer
2005Hesperotestudo Cisneros p. 241
2014Hesperotestudo de la Fuente et al.
2018Hesperotestudo Vlachos
2019Hesperotestudo May

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
RankNameAuthor
Batrachosauria()
Cotylosauria()
Amniota
Sauropsida
classReptilia
Testudinata(Oppel 1811)
orderTestudinesBatsch 1788
suborderCryptodira
Pantestudinoidea
superfamilyTestudinoidea
Pantestudinidae
familyTestudinidaeBatsch 1788
genusHesperotestudo(Williams 1950)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
P. A. Meylan and W. Sterrer 2000North American land tortoises with very well developed dermal armour, with those ossicles over the tail united and, in combination with enlarged transverse processes of the caudal vertebrae, forming a caudal buckler (Auffenberg, 1963, 1974) or a tail club (DMNH 1600 and 1601); nuchal scute longer than wide; posteriorly excavated epiplastra; medially narrowed pectoral scutes; large inguinal scute in broad contact with femoral scute; transverse ridge (=commissural ridge) on premaxilla, median maxillary ridge and lingual ridge extending onto premaxilla; foramen praepalatinum obsolete; acromion process forming very large angle (110°) to body of scapula.
Measurements
partNmean
carapace height1131.0

Composition: phosphaticsubp
Environment: terrestrialf
Locomotion: actively mobilec
Life habit: ground dwellingf
Diet: herbivoref
Diet 2: herbivoref
Reproduction: oviparousf
Created: 2005-08-26 07:16:55
Modified: 2005-09-09 16:16:46
Source: f = family, c = class, subp = subphylum
References: Carroll 1988, Ernst and Barbour 1989, Hendy et al. 2009

Age range: base of the Whitneyan to the top of the Rancholabrean or 33.30000 to 0.01200 Ma

Collections (93 total)


Time interval Ma Country or state Original ID and collection number
Whitneyan33.3 - 30.8USA (South Dakota) H. sp. (185698)
Arikareean30.8 - 20.43USA (South Dakota) H. sp. (17517)
Late/Upper Oligocene28.4 - 23.03USA (Virginia) H. wilsoni (167203)
Harrisonian24.8 - 20.43USA (Nebraska) Testudo arenivaga (17735 195408)
Miocene23.03 - 5.333USA (Florida) Geochelone alleni (18545)
Miocene23.03 - 5.333USA (Texas) Testudo crassiscutata (190386)
Miocene - Pleistocene23.03 - 0.0117USA (Texas) Testudo turgida (99818)
Early/Lower Hemingfordian20.43 - 15.97USA (Delaware) H. wilsoni (18579)
Hemingfordian20.43 - 15.97USA (Texas) Testudinata indet. (18626)
Hemingfordian20.43 - 15.97USA (Nebraska) Testudo angusticeps (18001)
Hemingfordian20.43 - 15.97USA (Florida) H. sp. (18561) Testudo tedwhitei (18597)
Hemingfordian - Barstovian20.43 - 13.6USA (Colorado) Testudo osborniana (191836)
Langhian15.97 - 13.82USA (Maryland) Testudo ducateli (100213)
Barstovian15.97 - 13.6USA (Nebraska) Geochelone orthopygia (18141 18165 18275 18442) Geochelone orthopygia, Geochelone nordensis (18223 18284)
Barstovian15.97 - 13.6USA (Montana) Testudo farri (191916)
Early/Lower Barstovian - early Late Barstovian15.97 - 13.6USA (California) H. sp. (156738)
Clarendonian13.6 - 10.3USA (Nebraska) H. orthopygia (188779) Testudinidae indet. (18324)
Clarendonian13.6 - 10.3USA (Texas) H. sp. (18610 18611 18623)
Clarendonian13.6 - 10.3USA (Kansas) Geochelone orthopygia (18450)
Early/Lower Clarendonian13.6 - 10.3USA (Florida) H. sp. (161374)
Clarendonian - Hemphillian13.6 - 4.9USA (Oklahoma) H. sp. (70119)
Clarendonian - Pliocene13.6 - 2.588USA (Oklahoma) H. sp. (170193)
Tortonian - Zanclean11.62 - 3.6USA (Kansas) Xerobates undata (191555)
Late/Upper Miocene11.608 - 5.333Canada (Yukon) H. sp. (206138)
Late/Upper Miocene - Early/Lower Pliocene11.608 - 3.6USA (California) H. orthopygia (183467 183468 183469 183470 183471)
Hemphillian10.3 - 4.9USA (Montana) Testudo impensa (191918)
Hemphillian10.3 - 4.9USA (Florida) Geochelone alleni (18560) Geochelone sp. (18601) Geochelone turgida (18556)
Late/Upper Hemphillian10.3 - 4.9USA (Oklahoma) Testudo riggsi (18076)
Late/Upper Hemphillian10.3 - 4.9USA (Texas) Geochelone sp. (18036)
Late/Upper Hemphillian - Blancan10.3 - 1.8USA (Tennessee) H. sp. (26574)
Late/Upper Hemphillian - Blancan10.3 - 1.8USA (Indiana) H. sp. (18320)
Pliocene5.333 - 2.588USA (California) H. orthopygia (67489)
Blancan4.9 - 1.8USA (New Mexico) H. sp. (19968)
Blancan4.9 - 1.8USA (Nebraska) Geochelone oelrichi (19954) Geochelone oelrichi, Geochelone sp. (19963)
Blancan4.9 - 1.8USA (Texas) Geochelone johnstoni (20033)
Blancan4.9 - 1.8USA (Florida) H. sp. (113275)
Blancan4.9 - 1.8Mexico (Baja California Sur) Geochelone sp. (19673)
Blancan4.9 - 1.8USA (Kansas) Testudo rexroadensis (19936 19940)
Blancan4.9 - 1.8USA (Arizona) Geochelone sp. (19618)
Late/Upper Pliocene3.6 - 2.588USA (Kansas) Testudo riggsi (99819)
Late/Upper Pliocene3.6 - 2.588USA (Texas) Testudo turgida (191451)
Piacenzian - Gelasian3.6 - 1.806USA (California) H. sp. (166974)
Late/Upper Pliocene - Early/Lower Pleistocene3.6 - 0.781USA (Nebraska) Geochelone oelrichi (191910)
Early/Lower Pleistocene - Middle Pleistocene2.588 - 0.126El Salvador H. crassiscutata (70118)
Pleistocene2.588 - 0.0117USA (Florida) H. sp. (206227) Testudo incisa (195410) Testudo sellardsi (180339)
Pleistocene2.588 - 0.0117USA (Texas) Testudo crassiscutata (85137 196056)
Irvingtonian1.8 - 0.3USA (Florida) Geochelone crassiscutata, Geochelone sp. (20400) Geochelone mlynarskii (20289) H. crassiscutata (20402) H. mlynarsii, H. crassiscutata (58089) H. sp. (26550)
Irvingtonian1.8 - 0.3USA (California) H. sp. (28222)
Irvingtonian1.8 - 0.3USA (Texas) Geochelone sp. (20342)
Irvingtonian1.8 - 0.3USA (South Carolina) Hesperotestudo indet. (79919)
Rancholabrean0.3 - 0.012USA (Florida) Geochelone crassiscutata, Geochelone incisa (20491) H. crassiscutata (20440)
Late/Upper Pleistocene (glacial)0.126 - 0.0117USA (Mississippi) Geochelone crassiscutata (164396)
Late/Upper Pleistocene (interglacial)0.126 - 0.0117USA (Georgia) H. crassicutata, H. incisa (20604) H. incisa (20613)
Late/Upper Pleistocene (high glacial)0.126 - 0.0117USA (Texas) Geochelone crassiscutata (93103)
Late/Upper Pleistocene (interglacial)0.126 - 0.0117USA (Florida) Geochelone crassiscutata (79667) Testudo sellardsi (20616)
Late/Upper Pleistocene (glacial)0.126 - 0.0117USA (Florida) Geochelone crassiscutata (154921)
Late/Upper Pleistocene0.126 - 0.0117USA (Florida) Geochelone crassiscutata, Geochelone incisa (154805) H. crassiscutata (154804) Testudo sellardsi (34269)
Late/Upper Pleistocene0.126 - 0.0117USA (Texas) Testudo wilsoni (79632)
Late/Upper Pleistocene (late glacial)0.126 - 0.0117USA (Mississippi) Geochelone crassiscutata (164397)
Late/Upper Pleistocene0.126 - 0.0117Bermuda H. bermudae (190749)
Late/Upper Pleistocene (high glacial)0.126 - 0.0117USA (New Mexico) Geochelone wilsoni (76059)
Late/Upper Pleistocene (late glacial)0.126 - 0.0117USA (Florida) H. incisa (154722)
Late/Upper Pleistocene (glacial)0.126 - 0.0117USA (Texas) Geochelone wilsoni (76101)