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Hemiscylliidae

Chondrichthyes - Orectolobiformes - Hemiscylliidae

Synonymy list
YearName and author
1862Hemiscylliidae Gill
1963Hemiscylliidae Jordan p. 597
1991Hemiscyllidae Case p. 2
2001Hemiscylliidae Peng et al. p. 7
2003Hemiscylliidae Delsate p. 8
2003Hemiscylliidae Kriwet p. 585
2005Hemiscylliidae Tabuce et al. p. 385
2006Hemiscylliidae Underwood p. 234
2007Hemiscylliidae Cahuzac et al. p. 15
2007Hemiscylliidae Kriwet et al.
2014Hemiscylliidae Carnevale et al. p. 41
2016Hemiscylliidae Nelson et al. p. 61

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
classChondrichthyes
RankNameAuthor
subclassElasmobranchiiBonaparte 1838
infraclassEuselachii()
—NeoselachiiCompagno 1977
superorderGaleomorphiiCompagno 1973
orderOrectolobiformesApplegate 1972
familyHemiscylliidaeGill 1862
familyHemiscylliidaeGill 1862

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Hemiscylliidae Gill 1862
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G. †Adnetoscyllium Guinot et al. 2013
+
†Adnetoscyllium angloparisensis Guinot et al. 2013
G. †Almascyllium Cappetta 1980
G. Chiloscyllium Müller and Henle 1837
+
†Chiloscyllium frequens Guinot et al. 2013
†Chiloscyllium greeni Cappetta 1973
†Chiloscyllium meraense Noubhani and Cappetta 1997
†Chiloscyllium mereretiae Werner 1989
†Chiloscyllium salvani Noubhani and Cappetta 1997
†Chiloscyllium vulloi Guinot et al. 2013
G. †Cirrhoscyllium Smith 1913
G. †Folipistrix Kriwet 2003
+
†Folipistrix digitulus Kriwet 2003
G. Hemiscyllium Smith 1837
+
†Hemiscyllium hermani Müller 1989
†Hemiscyllium ocellatus Bonaterre 1788
G. †Palaeobrachaelurus Thies 1983
+
†Palaeobrachaelurus mitchelli Underwood 2004
†Palaeobrachaelurus mussetti Underwood and Ward 2004
G. †Pseudospinax Müller and Diedrich 1991
+
†Pseudospinax heterodon Underwood and Mitchell 1999
†Pseudospinax muftius Thies 1983
†Pseudospinax pusillus Müller and Diedrich 1991
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: marinesubp
Locomotion: actively mobileo
Life habit: nektobenthico
Diet: carnivoreo
Created: 2009-07-21 07:07:32
Modified: 2009-07-21 09:07:32
Source: o = order, subp = subphylum
References: Hendy et al. 2009, Wagner 2023, Carroll 1988

Age range

Maximum range based only on fossils: base of the Thouarsense to the top of the Middle Eocene or 180.10000 to 37.71000 Ma
Minimum age of oldest fossil (stem group age): 175.6 Ma

Collections (135 total)


Time interval Ma Country or state Original ID and collection number
Toarcian182.7 - 174.1Germany Chiloscyllium sp., Palaeobrachaelurus sp. (148936)
Bifrons182.0 - 175.6Luxembourg Palaeobrachaelurus sp. (141382)
Levesquei182.0 - 175.6Luxembourg Chiloscyllium sp. (148935)
Variabilis182.0 - 175.6Luxembourg Palaeobrachaelurus sp. (141384)
Thouarsense180.1 - 175.6Luxembourg Palaeobrachaelurus sp. (141386 141389)
Late/Upper Aalenian175.6 - 171.6Germany Palaeobrachaelurus sp. 1, Palaeobrachaelurus sp. 2 (139231)
Aalenian174.1 - 170.3Germany Palaeobrachaelurus aperizostus (137168)
Early/Lower Bajocian171.6 - 168.4France (Grand Est) Palaeobrachaelurus sp., Palaeobrachaelurus mussetti (179498)
Bathonian168.3 - 166.1United Kingdom (England) Palaeobrachaelurus mussetti (91448 91450 91453)
Middle Bathonian167.7 - 164.7Poland Palaeobrachaelurus sp. (141269 141299)
Callovian166.1 - 163.5United Kingdom Protospinax muftius, Palaeobrachaelurus bedfordensis, Palaeobrachaelurus alisonae (137185)
Late/Upper Callovian164.7 - 161.2Luxembourg Palaeobrachaelurus sp., Pseudospinax sp. (91447)
Late/Upper Callovian164.7 - 161.2Poland Palaeobrachaelurus sp., Pseudospinax sp. (117993)
Middle Callovian164.7 - 161.2Poland Pseudospinax sp. (97035)
Late/Upper Kimmeridgian - Early/Lower Tithonian155.7 - 145.0United Kingdom (England) Hemiscylliidae indet., Pseudospinax sp. (140224)
Late/Upper Barremian130.0 - 125.45United Kingdom (England) Palaeobrachaelurus mitchelli (213572)
Early/Lower Aptian121.4 - 122.46United Kingdom (England) Hemiscylliidae indet. (213576)
Middle Albian109.0 - 105.3United Kingdom (England) Chiloscyllium greeni (213325)
Late/Upper Albian105.3 - 99.6United Kingdom (England) Chiloscyllium greeni (213351) Hemiscyllium sp. (213371) Hemiscyllium sp., Chiloscyllium greeni (213331) Hemiscyllium sp., Pseudospinax heterodon (213345 213372) Pseudospinax heterodon (213333 213341 213350)
Late/Upper Albian105.3 - 99.6Ukraine Chiloscyllium sp. (214237)
Cenomanian100.5 - 93.9Germany (Westphalia) Chiloscyllium greenei, Hemiscyllium sp., Pseudospinax pusillus (210063)
Early/Lower Cenomanian99.6 - 93.5France (Poitou-Charentes) Chiloscyllium sp. (77229 77230 77232) Chiloscyllium sp., Almascyllium sp. (73862)
Early/Lower Cenomanian99.6 - 93.5United Kingdom (England) Chiloscyllium sp. 1, Pseudospinax heterodon, Adnetoscyllium sp. (213567) Pseudospinax heterodon (213355 213361 213362)
Late/Upper Cenomanian99.6 - 93.5Egypt Chiloscyllium mereretiae (214508)
Middle Cenomanian99.6 - 93.5United Kingdom (England) Chiloscyllium sp. 2 (213569)
Middle Cenomanian99.6 - 93.5USA (Nebraska) Chiloscyllium greeni (213308)
Late/Upper Turonian93.5 - 89.3USA (Texas) Chiloscyllium greeni (213299)
Late/Upper Turonian93.5 - 89.3USA (South Dakota) Brachaelurus greeni (213291)
Middle Turonian93.5 - 89.3France (Ardennes) Adnetoscyllium sp., Chiloscyllium vulloi (213564)
Middle Turonian93.5 - 89.3USA (Kansas) Chiloscyllium greeni (35365)
Middle Turonian93.5 - 89.3USA (Nebraska) Chiloscyllium greeni (213389)
Middle Turonian93.5 - 89.3USA (New Mexico) Chiloscyllium greeni (179267)
Coniacian89.8 - 86.3United Kingdom (Northern Ireland) Hemiscylliidae indet. 3, Chiloscyllium vulloi (213383)
Early/Lower Santonian - Middle Santonian85.8 - 83.5USA (Georgia) Chiloscyllium sp. (61769)
Early/Lower Santonian - Middle Santonian85.8 - 83.5USA (New Mexico) Chiloscyllium sp. (52037)
Late/Upper Santonian85.8 - 83.5Lebanon Almascyllium cheikhelissi (92241)
Late/Upper Santonian85.8 - 83.5United Kingdom (England) Adnetoscyllium angloparisensi, Chiloscyllium frequens (213377) Chiloscyllium frequens (213381)
Late/Upper Santonian85.8 - 83.5France (Aisne) Hemiscylliidae indet. 1, Adnetoscyllium angloparisensis, Chiloscyllium frequens (213563)
Middle Santonian85.8 - 83.5United Kingdom (England) Adnetoscyllium angloparisensis (213376) Chiloscyllium frequens (213380)
Early/Lower Campanian83.5 - 70.6France (Ardennes) Adnetoscyllium angloparisensis, Chiloscyllium frequens (157587)
Early/Lower Campanian83.5 - 70.6USA (Utah) Chiloscyllium missouriense (75839)
Early/Lower Campanian83.5 - 70.6France (Aisne) Adnetoscyllium angloparisensis, Chiloscyllium frequens (213559) Chiloscyllium frequens (213560)
Early/Lower Campanian83.5 - 70.6Sweden (Skåne) Chiloscyllium sp., Hemiscyllium hermani (74334)
Early/Lower Campanian83.5 - 70.6Sweden Chiloscyllium sp. (102883 102885)
Early/Lower Campanian83.5 - 70.6France (Somme) Chiloscyllium frequens (213565 213566)
Early/Lower Campanian83.5 - 70.6Sweden (Götaland) Chiloscyllium gaemersi, Chiloscyllium sp., Hemiscyllium hermani (102886)
Early/Lower Campanian83.5 - 70.6Sweden (Skane) Chiloscyllium gaemersi, Chiloscyllium sp., Hemiscylliidae indet., Hemiscyllium hermani (106411)
Early/Lower Campanian83.5 - 70.6USA (Texas) Chiloscyllium greeni (213303)
Early/Lower Campanian83.5 - 70.6United Kingdom (England) Adnetoscyllium angloparisensi, Chiloscyllium frequens (213378) Adnetoscyllium angloparisensis, Chiloscyllium frequens (213382 213561) Chiloscyllium frequens (213379)
Early/Lower Campanian83.5 - 70.6Germany (Niedersachsen) Hemiscyllium sp. (31992)
Late/Upper Campanian83.5 - 70.6USA (Wyoming) Chiloscyllium missouriense (70455)
Late/Upper Campanian83.5 - 70.6France (Marne) Hemiscylliidae indet. 2, Chiloscyllium frequens (157588)
Late/Upper Campanian83.5 - 70.6Canada (British Columbia) Hemiscyllium hermani (221571)
Late/Upper Campanian83.5 - 70.6USA (Montana) Chiloscyllium missouriensis (69150)
Late/Upper Campanian83.5 - 70.6Canada (Alberta) Chiloscyllium missouriense (45893)
Late/Upper Campanian83.5 - 70.6USA (Utah) Chiloscyllium missouriense (75849)
Late/Upper Campanian83.5 - 70.6Israel Chiloscyllium sp. (214862)
Late/Upper Campanian83.5 - 70.6Germany (Nordrhein-Westfalen) Chiloscyllium sp. (101701)
Middle Campanian83.5 - 70.6USA (Mississippi) Hemiscyllium sp. (164558)
Middle Campanian83.5 - 70.6USA (Utah) Squatirhina sp., Squatirhina americana (61844)
Middle Campanian83.5 - 70.6Canada (Alberta) Chiloscyllium missouriense (45560 45561 45566 45820)
Middle Campanian - Late/Upper Campanian83.5 - 70.6Mexico (Chihuahua) Chiloscyllium sp. (35353)
Middle Campanian - Late/Upper Campanian83.5 - 70.6USA (South Carolina) Chiloscyllium sp. (79363)
Late/Upper Campanian - Early/Lower Maastrichtian83.5 - 66.0USA (Colorado) Chiloscyllium sp. (167302)
Late/Upper Campanian - Early/Lower Maastrichtian83.5 - 66.0USA (New Jersey) Chiloscyllium greeni (221052)
Late/Upper Campanian - Early/Lower Maastrichtian83.5 - 66.0Syria Chiloscyllium sp. (117855 117857)
Middle Campanian - Early/Lower Maastrichtian83.5 - 66.0Peru (Amazonas) Chiloscyllium sp. (52839)
Maastrichtian72.1 - 66.0USA (Maryland) Chiloscyllium greeni (83331)
Maastrichtian72.1 - 66.0Jordan Hemiscyllium sp. (65221)
Early/Lower Maastrichtian70.6 - 66.0Spain (Cataluña) Hemiscyllium sp. (76212)
Early/Lower Maastrichtian70.6 - 66.0Jordan Hemiscyllium sp. (65223)
Early/Lower Maastrichtian70.6 - 66.0Morocco Chiloscyllium sp. 1, Chiloscyllium sp. 2 (116903) Hemiscyllium sp. (214826)
Lancian70.6 - 66.0USA (Montana) Chiloscyllium sp. (144502)
Late/Upper Maastrichtian70.6 - 66.0USA (New Jersey) Chiloscyllium sp., Hemiscyllium sp. (61807)
Late/Upper Maastrichtian70.6 - 66.0Spain (Burgos) Chiloscyllium sp. (176782)
Late/Upper Maastrichtian - Danian70.6 - 61.6Morocco Hemiscyllium sp. (214373)
Danian66.0 - 61.6Austria Hemiscyllium sp. (233361)
Selandian - Thanetian61.6 - 56.0Morocco Chiloscyllium salvani (214848)
Thanetian59.2 - 56.0Algeria Hemiscylliidae indet. (232262)
Ypresian56.0 - 47.8Morocco Chiloscyllium meraense (116904 214849 214850 214851)
Ypresian56.0 - 47.8USA (South Carolina) Hemiscyllium sp. (203258)
Ypresian56.0 - 47.8Senegal Chiloscyllium sp. (209379)
Ypresian - Lutetian56.0 - 41.3France (Landes) Hemiscyllium tailledisensis (161754)
Early/Lower Eocene - Middle Eocene55.8 - 37.71United Kingdom (England) Hemiscyllium bruxelliense (3256 3257)
Lutetian47.8 - 41.3Ukraine Hemiscyllium bruxelliensis (211009)
Lutetian47.8 - 41.3Morocco Hemiscyllium sp., Chiloscyllium meraense (91339)
Lutetian47.8 - 41.3Senegal Chiloscyllium meraense (133429)
Lutetian47.8 - 41.3United Kingdom (England) Hemiscyllium bruxelliense (41989 41992 41996) Hemiscyllium bruxelliensis (41888)
Lutetian47.8 - 41.3Togo Chiloscyllium sp. (32991)
Lutetian - Bartonian47.8 - 38.0France (Landes) Hemiscyllium tailledisensis (161753)
Lutetian - Bartonian47.8 - 38.0Egypt Hemiscyllium sp. (137540)
Middle Eocene47.8 - 37.71Ukraine (Crimea) Hemiscyllium bruxelliensis (140018)
Bartonian41.3 - 38.0Tunisia (Sidi Bouzid Township) Hemiscyllium sp. (173640)
Bartonian41.3 - 38.0Ukraine (Kirovograd) Hemiscyllium bruxelliensis (132381)
Priabonian38.0 - 33.9Egypt Chiloscyllium sp. (137568 137569 137579)
Priabonian38.0 - 33.9Western Sahara Chiloscyllium sp. (96686)