Basic info Taxonomic history Classification Included Taxa
Morphology Ecology and taphonomy External Literature Search Age range and collections

Hemicyoninae

Mammalia - Carnivora - Ursidae

Synonymy list
YearName and author
1926Hemicyoninae Frick p. 12
1991Hemicyoninae Qiu et al.
1998Hemicyoninae Hunt, Jr. p. 180
2019Hemicyoninae de Bonis et al.

Is something missing? Join the Paleobiology Database and enter the data

RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
subclassSynapsida
—Therapsida()
infraorderCynodontia()
—Mammaliamorpha
—Mammaliaformes
classMammalia
—Theriamorpha(Rowe 1993)
RankNameAuthor
—Theriiformes()
—Trechnotheria
—Cladotheria
—Zatheria
subclassTribosphenida()
subclassTheria
—Eutheria()
—Placentalia
—Boreoeutheria
—Laurasiatheria
—Scrotifera
—Ferae()
—CarnivoramorphaWyss and Flynn 1993
—CarnivoraformesFlynn et al.
orderCarnivora
suborderCaniformiaKretzoi 1943
infraorderCanoidea(Simpson 1931)
superfamilyArctoideaFlower 1869
—Ursida
superfamilyUrsoideaFischer de Waldheim 1817
familyUrsidaeGray 1825
subfamilyHemicyoninae
subfamilyHemicyoninae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. †Hemicyoninae Frick 1926
show all | hide all
Tr. †Cephalogalini de Bonis 2013
hide
G. †Adelpharctos De Bonis 1971
+
†Adelpharctos ginsburgi de Bonis 2011
†Adelpharctos mirus De Bonis 1971
G. †Cephalogale Jourdan 1862
+
†Cephalogale brevirostris Croizet 1845
†Cephalogale bugtiensis Forster-Cooper 1923
†Cephalogale geoffroyi Jourdan 1862
†Cephalogale meschethense Gabunia 1964
†Cephalogale ursinus Bonis 1973
G. †Cyonarctos de Bonis 2013
+
†Cyonarctos dessei de Bonis 2013
+
Invalid names: Cephalogale cadurcensis Filhol 1877 [synonym]
G. †Filholictis de Bonis 2013
+
†Filholictis filholi Filhol 1877
G. †Phoberogale Ginsburg and Morales 1995
+
†Phoberogale bonali Helbing 1928
†Phoberogale depereti Viret 1929
†Phoberogale gracile Pomel 1847
†Phoberogale minor Jourdan 1862
†Phoberogale shareri Wang et al. 2009
G. †Dinocyon Jourdan 1861
hide
†Dinocyon aurelianensis Frick 1926
†Dinocyon sansaniensis Frick 1926
†Dinocyon thenardi Jourdan 1861
G. †Hemicyon Lartet 1851
hide
†Hemicyon barbouri Colbert 1941
†Hemicyon goriachensis Toula 1884
†Hemicyon grivensis Frick 1926
†Hemicyon mayorali Astibia et al. 2000
†Hemicyon minor Dépéret 1887
†Hemicyon sansaniensis Lartet 1851
†Hemicyon statzlingii Frick 1926
G. †Phoberocyon Ginsburg 1955
hide
†Phoberocyon huerzeleri Ginsburg 1955
†Phoberocyon johnhenryi White 1947
†Phoberocyon youngi Xiang 1986
G. †Plithocyon Ginsburg 1955
hide
†Plithocyon armagnacensis Ginsburg 1955
+
Invalid names: Hemicyon californicus Frick 1926 [synonym]
†Plithocyon teilhardi Colbert 1939
†Plithocyon ursinus Cope 1875
G. †Zaragocyon Ginsburg and Morales 1995
hide
†Zaragocyon daamsi Ginsburg and Morales 1995
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: terrestrialsubc
Locomotion: actively mobilec
Life habit: ground dwellingf
Diet: herbivoref
Diet 2: carnivoref
Reproduction: viviparoussubc
Created: 2005-06-08 10:11:10
Modified: 2005-08-22 19:31:30
Source: f = family, subc = subclass, c = class, subp = subphylum
References: Lillegraven 1979, Nowak 1991, Hendy et al. 2009, Ji et al. 2002, Carroll 1988

Age range: base of the late Late Arikareean to the top of the Serravallian or 29.50000 to 11.62000 Ma

Collections (65 total)


Time interval Ma Country or state Original ID and collection number
Early/Lower Oligocene33.9 - 28.4China (Nei Mongol) Cephalogale sp. (175387)
MP 2133.9 - 28.4Czech Republic Cephalogale sp. (91467)
MP 2233.9 - 28.4France Cephalogale minor (183452)
MP 2233.9 - 28.4France (Quercy) Cephalogale sp. (179140)
MP 2333.9 - 28.4France (Quercy) Adelpharctos mirus (36709)
late Late Arikareean29.5 - 18.5USA (California) Phoberogale shareri (160461)
Late/Upper Oligocene28.4 - 23.03Germany Cephalogale gracile (183664)
Late/Upper Oligocene28.4 - 23.03Georgia Cephalogale meschethense (182744)
MP 2828.4 - 23.03France (Quercy) Adelpharctos ginsburgi, Cephalogale sp. (36712) Cyonarctos dessei (36715)
Chattian28.1 - 23.03France Phoberocyon hispanicus (187576)
Chattian28.1 - 23.03Switzerland (Canton Solothurn) Cephalogale sp. 1, Cephalogale sp. 2, Adelpharctos mirus (182805)
Harrisonian24.8 - 18.5USA (Wyoming) Cephalogale sp. (17839)
Harrisonian24.8 - 18.5USA (Nebraska) Cephalogale sp. (17741 28043)
Harrisonian24.8 - 18.5USA (New Mexico) Cephalogale sp. (17844)
Early/Lower Miocene - Middle Miocene23.03 - 11.608Spain (Castile-La Mancha) Plithocyon armagnecensis, Dinocyon thenardi (136269)
Miocene23.03 - 5.333USA (Montana) Hemicyon ursinus (19038)
Miocene23.03 - 5.333USA (Nebraska) Plithocyon sp. (18276)
Miocene23.03 - 5.333France Hemicyon vincenti (200296)
Miocene23.03 - 5.333Austria Dinocyon sp., Dinocyon thenardi (63085)
MN 222.4 - 20.0Spain (Zaragoza) Cephalogale sp., Zaragocyon daamsi, Phoberogale ginesticus (35750)
Burdigalian20.44 - 15.97France (Loiret) Phoberocyon aurelianensis (181642 181675)
Burdigalian20.44 - 15.97France Phoberocyon dehmi (148399) Plithocyon bruneti (148398)
Burdigalian20.44 - 15.97France (Maine-et-Loire) Hemicyon hareni, Phoberocyon dehmi, Plithocyon bruneti (48069)
Burdigalian20.44 - 15.97Spain (Zaragoza) Hemicyon sansaniensis (40716)
Burdigalian20.44 - 15.97Spain (Valencia) Hemicyon sansaniensis (40031)
Burdigalian - Tortonian20.44 - 7.246France Hemicyon gargan, Phoberocyon dehmi, Plithocyon bruneti, Phoberocyon aurelianensis (187579) Hemicyon sansaniensis (187580)
Early/Lower Hemingfordian20.43 - 15.97USA (Nebraska) Cephalogale sp. (17881 17887 17888 17921 17942)
Hemingfordian20.43 - 15.97USA (Florida) Aelurodon johnhenryi (18597)
MN 320.0 - 16.9Germany (Bavaria) Hemicyon spp. (68040)
Orleanian20.0 - 15.97France Hemicyon gargan, Hemicyon stehlini, Phoberocyon dehmi, Plithocyon bruneti (142146)
MN 416.9 - 15.97France (Centre) Hemicyon stehlini (69116)
MN 416.9 - 15.97France (Midi-Pyrénées) Hemicyon stehlini (71873)
MN 416.9 - 15.97France (Alsac) Hemicyon stehlini (69106)
MN 416.9 - 15.97France (Gers) Hemicyon stehlini (69099)
Barstovian16.3 - 12.5USA (New Mexico) Hemicyon ursinus (19535)
Barstovian16.3 - 12.5USA (Nebraska) Hemicyon sp. (18443)
Barstovian16.3 - 12.5USA (California) Plithocyon barstowensis (19364) Plithocyon sp. (19557 19581)
Early/Lower Barstovian16.3 - 12.5USA (New Mexico) Plithocyon sp. (19547)
Langhian15.97 - 13.82France (Maine-et-Loire) Hemicyon sansaniensis (148396)
Langhian15.97 - 13.82Spain (Catalonia) Hemicyon stehlini (180192)
Langhian15.97 - 13.82China (Hubei) Hemicyon youngi (42068)
MN 515.97 - 13.65Turkey (Bursa) Plithocyon sp., Hemicyon sansaniensis (68547)
MN 5 - MN 615.97 - 12.75Bosnia and Herzegovina Hemicyon goeriachensis (198251)
Langhian - Serravallian15.97 - 11.62France (Midi-Pyrenees) Plithocyon armagnacensis, Hemicyon sansaniensis (32191)
Langhian - Tortonian15.97 - 7.246France (Midi-Pyrenees Region) Hemicyon sansaniensis (31775) Plithocyon armagnacensis (31776)
Langhian - Tortonian15.97 - 7.246France (Maine-et-Loire) Hemicyon sp., Hemicyon gargan, Hemicyon stehlini, Hemicyon sansaniensis (142149)
Middle Miocene - Late/Upper Miocene15.97 - 5.333China (Nei Mongol) Hemicyon teilhardi (28781)
MN 613.65 - 12.75China (Gansu) Hemicyon teilhardi (71105)
MN 613.65 - 12.75Slovakia (Bratislava) Hemicyon sansaniensis (69711)
MN 6 - MN 7 + 813.65 - 11.1Slovakia Hemicyon goeriachensis (69712)
MN 7 + 812.75 - 11.1Spain Hemicyon sp. (70806)
Clarendonian12.5 - 9.4USA (Nebraska) Hemicyon barbouri (18229)
Late/Upper Clarendonian12.5 - 9.4USA (Florida) Plithocyon sp. (18554)