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Protospermophilus quatalensis

Mammalia - Rodentia - Sciuridae

Taxonomy
Citellus quatalensis was named by Gazin (1930). Its type specimen is CIT 30, a partial skull ( A skull without the cranium and the posterior half of the orbits, broken upper incisors and left P3 through M2 present; a left mandibular ramus with lower inci), and it is a 3D body fossil. Its type locality is Quatal Canyon South Side 3, which is in a Clarendonian terrestrial horizon in the MR Caliente Formation of California.

It was recombined as Protospermophilus quatalensis by Bryant (1945), James (1963) and Black (1963).

Synonymy list
YearName and author
1930Citellus quatalensis Gazin
1945Protospermophilus quatalensis Bryant p. 350 figs. (P1. 7, figs. b, c.
1963Protospermophilus quatalensis Black
1963Protospermophilus quatalensis James

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
subclassSynapsida
—Therapsida()
infraorderCynodontia()
—Mammaliamorpha
RankNameAuthor
—Mammaliaformes
classMammalia
—Theriamorpha(Rowe 1993)
—Theriiformes()
—Trechnotheria
—Cladotheria
—Zatheria
subclassTribosphenida()
subclassTheria
—Eutheria()
—Placentalia
—Boreoeutheria
—EuarchontogliresMurphy et al. 2001
—GliriformesWyss and Meng 1996
—Glires()
—Simplicidentata()
orderRodentiaBowdich 1821
superfamilySciuroidea
familySciuridae
genusProtospermophilusGazin 1930
speciesquatalensis()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

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Diagnosis
ReferenceDiagnosis
M. D. Bryant 1945Skull massive; width of rostrum greater than depth; ridges on pre- maxillae lateral to incisors prominent; palate wide in proportion to length of tooth row but actually narrower than in P. malheurensis and P. angusticeps; fossae anterolateral to incisive foramina deep; massereric tubercles moderately prominent. Lower jaw shallow and stout; dorsal surface between incisor and p4 slightly depressed; masseteric crests prominent; masseteric fossa terminates anteriorly below posterior margin of p4. Mandibular tooth row long in proportion to depth of jaw; p3 a minute cone; p4, m1 and m2 nearly quadrate in occlusal outline, low crowned, cingula separated from protocone by furrows; mesostyles and mesostylids present on molars.
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: terrestrialsubc
Locomotion: actively mobilec
Life habit: scansorialf
Diet: granivoref
Diet 2: frugivoref
Reproduction: viviparoussubc
Created: 2005-06-08 10:11:09
Modified: 2005-08-26 13:04:24
Source: f = family, subc = subclass, c = class, subp = subphylum
References: Carroll 1988, Ji et al. 2002, Hendy et al. 2009, Nowak 1991, Lillegraven 1979

Age range: base of the Barstovian to the top of the Clarendonian or 16.30000 to 9.40000 Ma

Collections (17 total)


Time interval Ma Country or state Original ID and collection number
Barstovian16.3 - 12.5USA (Texas) Protospermophilus quatalensis (18646 18664)
Early/Lower Barstovian16.3 - 12.5USA (Texas) Protospermophilus quatalensis (18665)
Clarendonian12.5 - 9.4USA (California) Protospermophilus quatalensis (19181 19182 19361 19453 type locality: 19489 19496 19497 19507 19512 19515 19539 19541 19594)
Late/Upper Clarendonian12.5 - 9.4USA (Nebraska) Protospermophilus quatalensis (18327)