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Paratomarctus temerarius

Mammalia - Carnivora - Canidae

Taxonomy
Canis temerarius was named by Leidy (1858). It is a 3D body fossil. Its type locality is Niobrara River (coll. Hayden 1857), which is in a Barstovian/Clarendonian terrestrial horizon in the Loup Fork Formation of Nebraska.

It was recombined as Tephrocyon temerarius by Matthew and Cook (1909), Peterson (1910), Merriam (1913), Matthew (1918) and Merriam (1919); it was recombined as Tomarctus temerarius by Matthew (1924), Green (1948), Galbreath (1953), Downs (1956), Sutton (1977), Voorhies (1990) and Munthe (1998); it was recombined as Paratomarctus temerarius by Wang et al. (1999) and Wang et al. (2008).

Synonyms
Synonymy list
YearName and author
1858Canis temerarius Leidy p. 21
1893Canis anceps Scott p. 660
1902Canis anceps Hay p. 775
1902Canis temerarius Hay p. 776
1909Tephrocyon temerarius Matthew and Cook
1910Tephrocyon temerarius Peterson
1913Tephrocyon temerarius Merriam
1918Tephrocyon temerarius Matthew
1919Tephrocyon temerarius Merriam
1924Tomarctus temerarius Matthew
1948Tomarctus temerarius Green
1953Tomarctus temerarius Galbreath
1956Tomarctus temerarius Downs
1977Tomarctus temerarius Sutton
1990Tomarctus temerarius Voorhies
1998Tomarctus temerarius Munthe
1999Paratomarctus temerarius Wang et al.
2008Paratomarctus temerarius Wang et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
subclassSynapsida
—Therapsida()
infraorderCynodontia()
—Mammaliamorpha
—Mammaliaformes
classMammalia
RankNameAuthor
—Theriamorpha(Rowe 1993)
—Theriiformes()
—Trechnotheria
—Cladotheria
—Zatheria
subclassTribosphenida()
subclassTheria
—Eutheria()
—Placentalia
—Boreoeutheria
—Laurasiatheria
—Scrotifera
—Ferae()
—CarnivoramorphaWyss and Flynn 1993
—CarnivoraformesFlynn et al.
orderCarnivora
familyCanidae
subfamilyBorophaginae
tribeBorophaginiSimpson 1945
subtribeBorophagina
genusParatomarctus
speciestemerarius()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

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Invalid names: Canis anceps Scott 1893 [synonym]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: terrestrialsubc
Locomotion: actively mobilec
Life habit: ground dwellingf
Diet: carnivoref
Diet 2: omnivoref
Reproduction: viviparoussubc
Created: 2005-06-08 10:11:09
Modified: 2005-08-22 19:28:06
Source: f = family, subc = subclass, c = class, subp = subphylum
References: Hendy et al. 2009, Ji et al. 2002, Carroll 1988, Lillegraven 1979, Nowak 1991

Age range: base of the Barstovian to the top of the Clarendonian or 16.30000 to 9.40000 Ma

Collections (47 total)


Time interval Ma Country or state Original ID and collection number
Hemingfordian20.43 - 15.97USA (Oregon) Paratomarctus temerarius (212623)
Barstovian16.3 - 12.5USA (Oregon) Tomarctus brevirostris (19056)
Barstovian16.3 - 12.5USA (Nebraska) Paratomarctus temerarius (18109 18123 18124 18131 18144 18155 18172 18265 18272 18281) Tomarctus temerarius (18164 18192 18261 18275 18284 18442)
Barstovian16.3 - 12.5USA (Texas) Carnivora indet. (18627) Paratomarctus temerarius (18642)
Barstovian16.3 - 12.5USA (Montana) Canis anceps (18894)
Barstovian16.3 - 12.5USA (New Mexico) Paratomarctus temerarius (19274 19287 19435 19520 19535) Tomarctus temerarius (19519)
Barstovian16.3 - 12.5USA (California) Paratomarctus temerarius (19362 19364 19366 19422 19432 19457 19463 19522 19542 19548 19556 19568 214888) Tephrocyon temerarius (19157)
Barstovian16.3 - 12.5USA (Nevada) Cynodesmus kelloggi (19088)
Early/Lower Barstovian16.3 - 12.5USA (Texas) Paratomarctus temerarius (18665)
Early/Lower Barstovian16.3 - 12.5USA (Montana) Tomarctus temerarius (18803)
Barstovian - Clarendonian16.3 - 9.4USA (Nebraska) Canis temerarius (190543)
Langhian15.97 - 13.82USA (California) Paratomarctus temerarius (59059)
Clarendonian - Hemphillian12.5 - 4.9USA (Wyoming) Tomarctus compressus (18424)