PBDB Taxon

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Ceratopsidae (ceratopsid)

Reptilia - Ceratopsidae

Taxonomy

Ceratopsidae was named by Marsh (1888). Its type is Ceratops. It was considered monophyletic by Sereno (2000).

It was corrected as Ceratopidae by Olshevsky and Ford (1994); it was considered a nomen dubium by Lull (1906).

It was assigned to Ornithopoda by Mansel-Pleydell (1902); to Orthopoda by Depéret (1896), Nopcsa (1901), Lambe (1905), Huene (1909) and Lambe (1911); to Postpubici by Jaekel (1911); to Phytophagi by Jaekel (1914); to Predentata by von Zittel (1911) and Gilmore (1919); to Dinosauria by Marsh (1888) and Joleaud (1922); to Thyreophora by Nopcsa (1923), Nopcsa (1923), Nopcsa (1923), Huene (1927) and Huene (1929); to Ceratopsoidae by Hay (1930); to Thyreophoroidea by Huene (1934); to Thyreophoroidea by Nopcsa (1928), Nopcsa (1929) and Huene (1934); to Ceratopsia by Parks (1935); to Ponderopoda by Huene (1948) and Huene (1959); to Ornithischia by Lapparent (1947) and Madsen and Miller (1979); to Neoceratopia by Olshevsky and Ford (1994); to Ceratopsia by Marsh (1890), Marsh (1895), Marsh (1896), Hatcher et al. (1907), Matthew (1915), Gregory and Mook (1925), Russell (1930), Roxo (1937), Kuhn (1946), Huene (1950), Parsch (1963), Tatarinov (1964), Kuhn (1964), Estes (1964), Kuhn (1966), Romer (1966), Charig (1967), Swinton (1970), Thulborn (1971), Ostrom (1980), McIntosh (1981), McGinnis (1982), Sues and Galton (1982), Russell (1984), Maryanska and Osmólska (1985), Ostrom and Wellnhofer (1986), Bakker (1986), Carroll (1988) and Nessov (1995); to Ceratopsomorpha by Wolfe and Kirkland (1998); to Coronosauria by Zheng et al. (2015); to Ceratopsoidea by Hay (1902), Zhao (1983), Norman (1984), Cooper (1985), Sereno (1986), Sereno (1997), Sereno (1998), Sereno (1999), Sereno (2000), Lambert et al. (2001), Makovicky (2001), You and Dodson (2003), Chinnery and Horner (2007), Farke et al. (2014) and Morschhauser et al. (2019); to Ceratopsia by Tanoue et al. (2009), Brown and Henderson (2015), Rivera-Sylva et al. (2017) and Fowler and Freedman Fowler (2020); and to Neoceratopsia by Dodson and Currie (1990), Dodson (1990), Dodson (1997), Chinnery and Weishampel (1998), Penkalski and Dodson (1999), Peng et al. (2001), Chinnery (2004), Ryan and Evans (2005), Ryan (2007), Ryan et al. (2017), Dalman et al. (2018) and Wilson et al. (2020).

Subtaxa

Agathaumas, Bison alticornis, Centrosaurinae, Ceratopsinae, Ceratopsipes, Chasmosaurinae, Monocloniinae, Pachyrhinosaurinae, Polyonax, Triceratops galeus, Triceratops ingens, Triceratops maximus, Ugrosaurus

Synonyms

Polyonax was named by Cope (1874). Its type is Polyonax mortuarius.

It was synonymized subjectively with Ceratops by Zittel (1890); it was synonymized subjectively with Monoclonius by Nopcsa (1901); it was synonymized subjectively with Agathaumas by Hay (1902) and Romer (1956); it was synonymized subjectively with Trachodon by Hatcher (1902); it was synonymized subjectively with Triceratops by Tatarinov (1964), Kuhn (1964) and Romer (1966); it was considered a nomen dubium by Hatcher et al. (1907); it was considered a nomen dubium by Dodson and Currie (1990), Olshevsky and Ford (1994), Dodson (1997), Breithaupt (1999) and Dodson et al. (2004).

It was assigned to Dinosauria by Cope (1874); to Agathaumidae by Cope (1889), Cope (1889) and Cope (1891); to Reptilia by Huene (1909); and to Ceratopsidae by Marsh (1895), Marsh (1896) and Hay (1930).
Agathaumas was named by Cope (1872). Its type is Agathaumas sylvestris. It is the type genus of Agathaumidae.

It was synonymized subjectively with Triceratops by Lambe (1899), Tatarinov (1964), Romer (1966) and Breithaupt (1997); it was considered a nomen dubium by Ostrom and Wellnhofer (1986), Dodson and Currie (1990), Olshevsky and Ford (1994), Dodson (1997) and Dodson et al. (2004).

It was assigned to Dinosauria by Cope (1872); to Madrosauridae by Sauvage (1882); to Hadrosauridae by Dollo (1882), Marsh (1882), Cope (1883), Dollo (1884), Marsh (1884) and Marsh (1885); to Ceratopsia by Zittel (1890); to Scelidosauria by Seeley (1892); to Agathaumidae by Cope (1889), Cope (1891), Cope (1892), Cope (1898) and Lull (1906); to Ceratopsia by Tornier (1913); and to Ceratopsidae by Marsh (1895), Marsh (1896), Nopcsa (1901), Hay (1902), Hatcher et al. (1907), Huene (1909), von Zittel (1911), Huene (1927), Hay (1930), Russell (1930), Romer (1956), Kuhn (1964) and Breithaupt (1985).
Agathaumas sylvestris was named by Cope (1872). Its type specimen is AMNH 4000, a partial skeleton, and it is a 3D body fossil. Its type locality is Black Buttes Station, which is in a Maastrichtian estuary/bay sandstone/coal in the Lance Formation of Wyoming. It is the type species of Agathaumas.

It was considered a nomen dubium by Ostrom and Wellnhofer (1986), Dodson and Currie (1990), Breithaupt (1997) and Dodson et al. (2004).
Bison alticornis was named by Marsh (1887). Its type specimen is USNM 4739 and is a 3D body fossil. Its type locality is Windsor Addition, Highland, which is in a Maastrichtian terrestrial sandstone in the Denver Formation of Colorado.

It was recombined as Polyonax alticornis by Cope (1889); it was recombined as Ceratops alticornis by Marsh (1889), Marsh (1897), Walcott (1900) and Hay (1902); it was recombined as Triceratops alticornis by Cannon (1906), Hatcher et al. (1907), Huene (1927), Hay (1930), Johnson (1931), Schlaikjer (1935), Kuhn (1964) and Ostrom (1965); it was considered a nomen dubium by Lull (1933), Ostrom and Wellnhofer (1986), Dodson and Currie (1990), Ostrom and Wellnhofer (1990) and Dodson et al. (2004).
Polyonax mortuarius was named by Cope (1874). Its type specimen is USNM 244535/AMNH 3950, a partial skeleton, and it is a 3D body fossil. Its type locality is Bijou Creek (USG&GS), which is in a Maastrichtian terrestrial horizon in the Laramie Formation of Colorado. It is the type species of Polyonax.

It was recombined as Agathaumas mortuarius by Hay (1902); it was recombined as Triceratops mortuarius by Kuhn (1964); it was considered a nomen dubium by Hatcher et al. (1907); it was considered a nomen dubium by Ostrom and Wellnhofer (1986), Dodson and Currie (1990), Breithaupt (1999) and Dodson et al. (2004).
Agathaumidae was named by Cope (1889) ["I shall call it the Agathaumidae (or hellenicé Agathaumantidae), from the longest known genus, Agathaumas." Cope 1889:715.]. Its type is Agathaumas.

It was synonymized subjectively with Ceratopsidae by Lambe (1902), Hay (1902) and Hatcher et al. (1907).

It was assigned to Dinosauria by Cope (1889); to Iguanodontia by Baur (1891); and to Orthopoda by Cope (1889), Cope (1891) and Cope (1898).
Triceratops maximus was named by Brown (1933). Its type specimen is AMNH 5040, a set of vertebrae, and it is not a trace fossil. Its type locality is spring on Rock Creek (AMNH), which is in a Maastrichtian terrestrial sandstone in the Hell Creek Formation of Montana.

It was synonymized subjectively with Triceratops prorsus by Sloan (1976); it was considered a nomen dubium by Lull (1933); it was considered a nomen dubium by Ostrom and Wellnhofer (1986), Dodson and Currie (1990), Ostrom and Wellnhofer (1990) and Dodson et al. (2004).
Triceratops galeus was named by Marsh (1889). Its type specimen is USNM 2410 and is a 3D body fossil. Its type locality is Brighton area, Henderson (1), which is in a Maastrichtian terrestrial horizon in the Arapahoe Formation of Colorado.

It was considered a nomen dubium by Lull (1933); it was considered a nomen dubium by Ostrom and Wellnhofer (1986), Dodson and Currie (1990), Ostrom and Wellnhofer (1990) and Dodson et al. (2004).
Ugrosaurus was named by CoBabe and Fastovsky (1987). It is not extant. Its type is Ugrosaurus olsoni.

It was considered a nomen dubium by Dodson and Currie (1990), Lehman (1990) and Dodson et al. (2004).

It was assigned to Ceratopsidae by CoBabe and Fastovsky (1987).
Ugrosaurus olsoni was named by CoBabe and Fastovsky (1987). Its type specimen is UCMP 128561, a skull, and it is not a trace fossil. Its type locality is Dave's Nose, UCMP V-83224, which is in a Maastrichtian fluvial-lacustrine siltstone in the Hell Creek Formation of Montana. It is the type species of Ugrosaurus.

It was considered a nomen dubium by Dodson and Currie (1990) and Dodson et al. (2004).
Triceratops ingens was named by Lull (1915). Its type specimen is YPM 1828 and is a 3D body fossil. Its type locality is Lance Creek, south of Dogie Creek, which is in a Maastrichtian terrestrial horizon in the Lance Formation of Wyoming.

It was considered a nomen nudum by Ostrom and Wellnhofer (1986); it was considered a nomen dubium by Dodson and Currie (1990); it was considered a nomen nudum by Ostrom and Wellnhofer (1990); it was considered a nomen dubium by Dodson et al. (2004).

Synonymy list

Year	Name and author
1872	Agathaumas Cope p. 2
1872	Agathaumas sylvestris Cope p. 2
1872	Agathaumas Cope pp. 481-483
1872	Agathaumas sylvestris Cope p. 483
1873	Agathaumas sylvestris Cope p. 1
1874	Polyonax Cope p. 10ff
1874	Polyonax mortuarius Cope p. 10ff
1874	Agathaumas sylvestris Cope p. 18
1874	Polyonax mortuarius Cope p. 25
1875	Agathaumas sylvestris Cope p. 54
1875	Polyonax mortuarius Cope p. 64
1875	Agathaumas sylvestris Cope p. 435
1875	Polyonax mortuarius Cope p. 448
1882	Agathaumas Dollo
1882	Agathaumas Marsh
1882	Agathaumas Sauvage p. 35
1883	Agathaumas Cope p. 99
1884	Agathaumas Dollo p. 298
1884	Agathaumas Marsh p. 69
1885	Agathaumas Marsh p. 764
1887	Bison alticornis Marsh
1888	Bison alticornis Cannon p. 143
1888	Ceratopsidae Marsh p. 478
1889	Agathaumas sylvestris Cope
1889	Agathaumidae Cope p. 16
1889	Polyonax alticornis Cope p. 290
1889	Agathaumas Cope p. 715
1889	Agathaumidae Cope p. 715
1889	Polyonax Cope p. 715–716
1889	Polyonax mortuarius Cope p. 715–716
1889	Polyonax Cope p. 906
1889	Ceratops alticornis Marsh pp. 174-175
1889	Triceratops galeus Marsh pp. 174-175
1890	Ceratopsidae Marsh p. 425
1890	Agathaumas Zittel p. 751
1891	Agathaumidae Baur p. 450
1891	Agathaumas Cope p. 43
1891	Agathaumas sylvestris Cope p. 43
1891	Agathaumidae Cope p. 43
1891	Polyonax Cope p. 43
1892	Agathaumas Cope p. 758
1892	Agathaumas sylvestre Cope p. 758
1892	Agathaumas Seeley
1895	Agathaumas Marsh
1895	Ceratopsidae Marsh
1895	Polyonax Marsh
1896	Ceratopsidae Depéret p. 184
1896	Agathaumas sylvestre Hatcher p. 113
1896	Agathaumas Marsh p. 243
1896	Ceratopsidae Marsh p. 243
1896	Polyonax Marsh p. 243
1897	Ceratops alticornis Marsh p. 512
1897	Triceratops galeus Marsh p. 527
1898	Agathaumas Cope p. 70
1898	Agathaumas sylvestris Cope p. 70
1898	Agathaumidae Cope p. 70
1900	Ceratops alticornis Walcott p. 23
1901	Ceratopsidae Nopcsa p. 209
1901	Agathaumas Nopcsa p. 216
1901	Agathaumas sylvestris Nopcsa p. 216
1902	Agathaumas Hay p. 498
1902	Agathaumas mortuarius Hay p. 498
1902	Agathaumas sylvestris Hay p. 498
1902	Ceratopsidae Hay p. 498
1902	Triceratops galeus Hay p. 498
1902	Ceratops alticornis Hay p. 500
1902	Polyonax mortuarius Lambe p. 68
1902	Ceratopidae Mansel-Pleydell p. lxxv
1905	Ceratopsidae Lambe p. 49
1906	Triceratops alticornis Cannon p. 198
1906	Agathaumas Lull p. 144
1907	Ceratopsidae Hatcher et al. p. 10
1907	Agathaumas Hatcher et al. pp. 11, 104
1907	Agathaumas sylvestris Hatcher et al. pp. 105, 168
1907	Triceratops alticornis Hatcher et al. p. 170
1909	Ceratopsidae Huene p. 17
1909	Agathaumas Huene p. 18
1909	Polyonax Huene p. 18
1909	Agathaumas sylvestris Knowlton p. 270
1911	Ceratopsidae Jaekel p. 161
1911	Ceratopsidae Lambe p. 9
1911	Ceratopsidae von Zittel p. 295
1911	Agathaumas von Zittel p. 296
1913	Agathaumas Tornier p. 375
1914	Ceratopsidae Jaekel p. 197
1915	Triceratops ingens Lull
1915	Ceratopsidae Matthew p. 32
1919	Ceratopsidae Gilmore p. 64
1922	Ceratopsidae Joleaud p. 59
1923	Ceratopsidae Nopcsa p. 106
1925	Ceratopsidae Gregory and Mook p. 4
1927	Ceratopsidae Huene p. 257
1927	Agathaumas Huene p. 261
1927	Agathaumas sylvestris Huene p. 261
1927	Triceratops alticornis Huene p. 261
1927	Triceratops galeus Huene p. 261
1928	Ceratopsidae Nopcsa p. 185
1929	Ceratopsidae Huene p. 155
1929	Ceratopsidae Nopcsa p. 71
1930	Ceratopsidae Hay p. 220
1930	Triceratops alticornis Hay p. 228
1930	Triceratops galeus Hay p. 229
1930	Triceratops ingens Hay p. 229
1930	Agathaumas Hay p. 230
1930	Agathaumas sylvestris Hay p. 231
1930	Polyonax Hay p. 231
1930	Polyonax mortuarius Hay p. 231
1930	Agathaumas Russell p. 140
1930	Agathaumas sylvestris Russell p. 140
1930	Ceratopsidae Russell p. 140
1931	Triceratops alticornis Johnson p. 368
1931	Triceratops galeus Johnson p. 368
1933	Triceratops maximus Brown
1934	Ceratopsidae Huene p. 15
1935	Ceratopsidae Parks p. 185
1935	Triceratops alticornis Schlaikjer p. 57
1937	Ceratopsidae Roxo p. 65
1946	Ceratopsidae Kuhn p. 66
1947	Ceratopsidae Lapparent pp. 17-18
1948	Ceratopsidae Huene p. 95
1950	Ceratopsidae Huene p. 351
1956	Agathaumas Romer p. 638
1959	Ceratopsidae Huene p. 122
1963	Ceratopsidae Parsch p. 4
1964	Ceratopsidae Estes p. 145
1964	Agathaumas Kuhn p. 54
1964	Agathaumas silvestris Kuhn p. 54
1964	Ceratopsidae Kuhn p. 54
1964	Triceratops mortuarius Kuhn p. 59
1964	Triceratops alticornis Kuhn p. 60
1964	Triceratops galeus Kuhn p. 60
1964	Triceratops ingens Kuhn p. 60
1964	Triceratops maximus Kuhn p. 60
1964	Triceratops maximus Russell p. 17
1964	Ceratopsidae Tatarinov p. 581
1965	Triceratops alticornis Ostrom p. 39
1965	Triceratops galeus Ostrom p. 39
1965	Triceratops maximus Ostrom p. 39
1966	Ceratopsidae Kuhn p. 106
1966	Ceratopsidae Romer p. 371
1967	Ceratopsidae Charig p. 717
1970	Ceratopsidae Swinton p. 254
1971	Ceratopsidae Thulborn pp. 77-78 fig. 4
1979	Ceratopsidae Madsen and Miller p. 5
1980	Ceratopsidae Ostrom p. 23
1981	Ceratopsidae McIntosh p. 41
1982	Ceratopsidae McGinnis p. 96
1982	Ceratopsidae Sues and Galton p. 189
1983	Ceratopsidae Zhao p. 300
1984	Ceratopsidae Norman p. 160 fig. 2
1984	Ceratopsidae Russell p. 29
1985	Agathaumas Breithaupt p. 170
1985	Agathaumas sylvestris Breithaupt p. 170
1985	Ceratopsidae Cooper pp. 292-293
1985	Ceratopsidae Maryanska and Osmólska pp. 143-144 fig. 1
1986	Ceratopsidae Bakker p. 461
1986	Ceratopsidae Ostrom and Wellnhofer p. 116
1986	Ceratopsidae Sereno
1987	Ugrosaurus CoBabe and Fastovsky p. 149 figs. 1-4
1987	Ugrosaurus olsoni CoBabe and Fastovsky p. 149 figs. 1-4
1988	Ceratopsidae Carroll
1990	Ceratopsidae Dodson p. 578
1990	Ceratopsidae Dodson and Currie p. 611
1994	Ceratopidae Olshevsky and Ford
1995	Ceratopsidae Nessov p. 54
1997	Ceratopsidae Dodson pp. 473, 475–476
1997	Ceratopsidae Sereno p. 443
1998	Ceratopsidae Chinnery and Weishampel pp. 578-579 fig. 10
1998	Ceratopsidae Sereno p. 62
1998	Ceratopsidae Wolfe and Kirkland p. 307 fig. 5
1999	Ceratopsidae Penkalski and Dodson
1999	Ceratopsidae Sereno p. 2138 fig. 2
2000	Ceratopsidae Sereno p. 482
2001	Ceratopsidae Lambert et al. p. 18
2001	Ceratopsidae Makovicky p. 254 fig. 18.3
2001	Ceratopsidae Peng et al. p. 35
2003	Ceratopsidae You and Dodson
2004	Ceratopsidae Chinnery pp. 584-585 fig. 9
2005	Ceratopsidae Ryan and Evans p. 323
2007	Ceratopsidae Chinnery and Horner p. 638 fig. 6
2007	Ceratopsidae Ryan p. 391 fig. 12
2009	Ceratopsidae Tanoue et al. p. 426
2014	Ceratopsidae Farke et al.
2015	Ceratopsidae Brown and Henderson
2015	Ceratopsidae Zheng et al. p. 6 fig. 3
2017	Ceratopsidae Rivera-Sylva et al.
2017	Ceratopsidae Ryan et al.
2018	Ceratopsidae Dalman et al.
2019	Ceratopsidae Morschhauser et al. p. 129 fig. 10
2020	Ceratopsidae Fowler and Freedman Fowler
2020	Ceratopsidae Wilson et al.

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
subclass	Eureptilia	()
—	Romeriida

Rank	Name	Author
—	Diapsida	()
—	Archosauromorpha	(Huene 1946)
—	Crocopoda
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
informal	Avemetatarsalia
—	Ornithodira
—	Dinosauromorpha
—	Dinosauriformes
—	Dinosauria	()
—	Ornithischia	()
—	Genasauria
—	Cerapoda
—	Marginocephalia
—	Ceratopsia	()
infraorder	Neoceratopsia
—	Coronosauria
superfamily	Ceratopsoidea
family	Ceratopsidae
family	Ceratopsidae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Ceratopsidae Marsh 1888 [ceratopsid]

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Subfm. †Centrosaurinae Lambe 1915

G. †Albertaceratops Ryan 2007

†Albertaceratops nesmoi Ryan 2007

Unr. †Centrosaurini Ryan et al. 2017

G. †Centrosaurus Lambe 1905

†Centrosaurus apertus Lambe 1905

Invalid names: Centrosaurus longirostris Sternberg 1940 [synonym], Monoclonius cutleri Brown 1917 [synonym], Monoclonius dawsoni Lambe 1902 [synonym], Monoclonius flexus Brown 1914 [synonym]

G. †Coronosaurus Ryan et al. 2012

†Coronosaurus brinkmani Ryan and Russell 2005

G. †Rubeosaurus McDonald and Horner 2010

†Rubeosaurus ovatus Gilmore 1930

G. †Spinops Farke et al. 2011

†Spinops sternbergorum Farke et al. 2011

G. †Styracosaurus Lambe 1913

†Styracosaurus albertensis Lambe 1913

Invalid names: Monoclonius nasicornus Brown 1917 [synonym], Styracosaurus parksi Brown and Schlaikjer 1937 [synonym]

G. †Diabloceratops Kirkland and Deblieux 2010

†Diabloceratops eatoni Kirkland and Deblieux 2010

G. †Machairoceratops Lund et al. 2016

†Machairoceratops cronusi Lund et al. 2016

G. †Menefeeceratops Dalman et al. 2021

†Menefeeceratops sealeyi Dalman et al. 2021

G. †Monoclonius Cope 1876

†Monoclonius crassus Cope 1876

Invalid names: Monoclonius lowei Sternberg 1940 [synonym]

Invalid names: Monoclonius fissus Cope 1889 [nomen dubium], Monoclonius recurvicornis Cope 1889 [nomen dubium], Monoclonius sphenocerus Cope 1889 [nomen dubium]

Unr. †Nasutoceratopsini Ryan et al. 2017

G. †Avaceratops Dodson 1986

†Avaceratops lammersi Dodson 1986

G. †Crittendenceratops Dalman et al. 2018

†Crittendenceratops krzyzanowskii Dalman et al. 2018

G. †Furcatoceratops Ishikawa et al. 2023

†Furcatoceratops elucidans Ishikawa et al. 2023

G. †Nasutoceratops Sampson et al. 2013

†Nasutoceratops titusi Sampson et al. 2013

G. †Yehuecauhceratops Rivera-Sylva et al. 2017

†Yehuecauhceratops mudei Rivera-Sylva et al. 2017

Unr. †Pachyrhinosaurini Fiorillo and Tykoski 2012

G. †Achelousaurus Sampson 1994

†Achelousaurus horneri Sampson 1994

G. †Einiosaurus Sampson 1994

†Einiosaurus procurvicornis Sampson 1994

Invalid names: Styracosaurus makeli Czerkas and Czerkas 1990 [replaced]

G. †Pachyrhinosaurus Sternberg 1950

†Pachyrhinosaurus canadensis Sternberg 1950

†Pachyrhinosaurus lakustai Currie et al. 2008

†Pachyrhinosaurus perotorum Fiorillo and Tykoski 2012

Invalid names: Pachyrostra Fiorillo and Tykoski 2012 [empty]

G. †Sinoceratops Xu et al. 2010

†Sinoceratops zhuchengensis Xu et al. 2010

G. †Stellasaurus Wilson et al. 2020

†Stellasaurus ancellae Wilson et al. 2020

G. †Wendiceratops Evans and Ryan 2015

†Wendiceratops pinhornensis Evans and Ryan 2015

G. †Xenoceratops Ryan et al. 2012

†Xenoceratops foremostensis Ryan et al. 2012

Invalid names: Brachyceratops Gilmore 1914 [nomen dubium]

G. †Ceratopsipes Lockley and Hunt 1995

†Ceratopsipes goldenensis Lockley and Hunt 1995

Subfm. †Chasmosaurinae Lambe 1915

G. †Agujaceratops Lucas et al. 2006 [Aguja horned face]

†Agujaceratops mariscalensis Lehman 1989

G. †Anchiceratops Brown 1914

†Anchiceratops ornatus Brown 1914

Invalid names: Anchiceratops longirostris Sternberg 1926 [synonym]

G. †Arrhinoceratops Parks 1925

†Arrhinoceratops brachyops Parks 1925

G. †Bisticeratops Dalman et al. 2022

†Bisticeratops froeseorum Dalman et al. 2022

G. †Bravoceratops Wick and Lehman 2013

†Bravoceratops polyphemus Wick and Lehman 2013

G. †Chasmosaurus Lambe 1914

†Chasmosaurus belli Lambe 1902

†Chasmosaurus russelli Sternberg 1940

Invalid names: Mojoceratops perifania Longrich 2010 [synonym]

Invalid names: Chasmosaurus brevirostris Lull 1933 [nomen dubium], Chasmosaurus kaiseni Brown 1933 [nomen dubium], Eoceratops Lambe 1915 [synonym], Mojoceratops Longrich 2010 [synonym], Monoclonius canadensis Lambe 1902 [nomen dubium], Protorosaurus Lambe 1914 [synonym]

G. †Coahuilaceratops Loewen et al. 2010

†Coahuilaceratops magnacuerna Loewen et al. 2010

G. †Judiceratops Longrich 2013

†Judiceratops tigris Longrich 2013

G. †Kosmoceratops Sampson et al. 2010

†Kosmoceratops richardsoni Sampson et al. 2010

G. †Medusaceratops Ryan et al. 2010

†Medusaceratops lokii Ryan et al. 2010

G. †Navajoceratops Fowler and Freedman Fowler 2020

†Navajoceratops sullivani Fowler and Freedman Fowler 2020

G. †Pentaceratops Osborn 1923

†Pentaceratops sternbergii Osborn 1923

Invalid names: Pentaceratops fenestratus Wiman 1930 [synonym]

G. †Sierraceratops Dalman et al. 2022

†Sierraceratops turneri Dalman et al. 2022

G. †Spiclypeus Mallon et al. 2016

†Spiclypeus shipporum Mallon et al. 2016

G. †Terminocavus Fowler and Freedman Fowler 2020

†Terminocavus sealeyi Fowler and Freedman Fowler 2020

Unr. †Triceratopsini Longrich 2011

G. †Eotriceratops Wu et al. 2007

†Eotriceratops xerinsularis Wu et al. 2007

G. †Regaliceratops Brown and Henderson 2015

†Regaliceratops peterhewsi Brown and Henderson 2015

G. †Titanoceratops Longrich 2011

†Titanoceratops ouranos Longrich 2011

G. †Torosaurus Marsh 1891

†Torosaurus latus Marsh 1891

Invalid names: Torosaurus gladius Marsh 1891 [objective synonym]

†Torosaurus utahensis Gilmore 1946

G. †Triceratops Marsh 1889

†Triceratops horridus Marsh 1889

Invalid names: Nedoceratops hatcheri Lull 1905 [synonym], Triceratops albertensis Sternberg 1949 [synonym], Triceratops brevicornus Hatcher and Lull 1905 [synonym], Triceratops calicornis Marsh 1898 [synonym], Triceratops elatus Marsh 1891 [synonym], Triceratops eurycephalus Schlaikjer 1935 [synonym], Triceratops flabellatus Marsh 1889 [synonym], Triceratops obtusus Marsh 1898 [synonym], Triceratops serratus Marsh 1890 [synonym]

†Triceratops prorsus Marsh 1890

Invalid names: Nedoceratops Ukrainsky 2007 [synonym], Ojoceratops Sullivan and Lucas 2010 [synonym], Sterrholophus Marsh 1891 [synonym], Tatankaceratops Ott and Larson 2010 [synonym], Triceratops sulcatus Marsh 1890 [nomen dubium]

G. †Utahceratops Sampson et al. 2010

†Utahceratops gettyi Sampson et al. 2010

G. †Vagaceratops Sampson 2010

†Vagaceratops irvinensis Holmes et al. 2001

Invalid names: Mercuriceratops gemini Ryan et al. 2014 [synonym], Pentaceratops aquilonius Longrich 2014 [synonym]

Invalid names: Mercuriceratops Ryan et al. 2014 [synonym]

Invalid names: Ceratops Marsh 1888 [nomen dubium]

Subfm. †Pachyrhinosaurinae Huene 1950

G. †Eucentrosaurus Chure and McIntosh 1989

Invalid names: Agathaumas Cope 1872 [nomen dubium], Agathaumidae Cope 1889 [synonym], Bison alticornis Marsh 1887 [nomen dubium], Ceratopsinae Abel 1919 [empty], Monocloniinae Nopcsa 1923 [empty], Polyonax Cope 1874 [nomen dubium], Triceratops galeus Marsh 1889 [nomen dubium], Triceratops ingens Lull 1915 [nomen dubium], Triceratops maximus Brown 1933 [nomen dubium], Ugrosaurus CoBabe and Fastovsky 1987 [nomen dubium]

Diagnosis

Reference	Diagnosis
B. F. Nopcsa 1901	Intermaxillare fogatlan. Szarvak es legyezöalaku parietale; praeorbitalis nyilás nincsen. Os rostrale. Csigolyák platycoelek. Elsö lábak a hátsókkal majdnem egyformák. Pubis durványos. Femur harmadik trochanter nélkül. Ersö pánczél; paták. (Az összes eddig ismert fajok a krétából valók.) [Intermaxillary toothless. Horns and fan-shaped parietal; there is no preorbital opening. Rostral bone. Vertebrae platycoelous. Forelegs almost identical to hind legs. Pubis is rough. Femur without a third trochanter. Strong armor; hooves. (All species known so far are from Chalk.)]
B. F. Nopcsa 1928	skull with nuchal fringe and generally with horns; no armour on body.
F. v. Huene 1948	Two interorbital horns and another one on nose; double skull-roof with empty space between frontal and parietal; prefrontal bordering on nasal.
P. Dodson 1997	Skulls are large, and prominent horns are found over either the nose or the eyes, but usually not both (except Pachyrhinosaurus). The frill formed by the parietal and squamosal bones typically is about as long as or longer than the rest of the skull. The frill may be additionally ornamented with scallops, spikes, or hooks. Parietal fenestrae may be large (Chasmosaurus and Pentaceratops) or absent (Avaceratops and Triceratops). The opening for the nostril is large. The sharp beak is toothless, as in most protoceratopsids. Ceratopsids had dental batteries consisting of numerous interlocking teeth in which the roots of each tooth are split by the tooth beneath it in the replacement series. Also unique to ceratopsids is the vertical orientation of the plane of contact between the upper and lower tooth batteries, resulting in a peculiar slicing action. The occipital condyle by which the back of the skull contacts the vertebral column is spherical and in some cases as large as a small grapefruit (up to 115 mm in diameter). There are 10 fused sacral vertebrae in ceratopsids. The femur is always longer than the tibia, andthe ungual phalanges are blunt, rounded, and hoof-like.

Measurements

No measurements are available

Composition:

hydroxyapatiteo

Entire body:

yeso

Adult length:

10 to < 100o

Adult width:

1.0 to < 10o

Adult height:

1.0 to < 10o

Maximum body mass:

10000 kgg

Minimum body mass:

1000 kgg

Architecture:

compact or denseo

Ontogeny:

accretion, modification of partso

Grouping:

gregariouso

Environment:

terrestrialsubo

Locomotion:

actively mobileo

Life habit:

ground dwellingsubo

Diet:

herbivoresubo

Reproduction:

oviparouso

Dispersal:

direct/internalo

Dispersal 2:

mobileo

Created:

2005-09-15 11:08:09

Modified:

2005-09-15 13:08:09

Source: g = genus, subo = suborder, o = order

References: Marsh 1875, Norman and Weishampel 1991, Peczkis 1995

Age range: base of the Late/Upper Turonian to the top of the Late/Upper Maastrichtian or 93.50000 to 66.00000 Ma

Collections (679 total)

Time interval	Ma	Country or state	Original ID and collection number
Late/Upper Cretaceous	100.5 - 66.0	Mexico (Chihuahua)	Centrosaurinae indet. (227180)
Late/Upper Turonian	93.5 - 89.3	Uzbekistan (Navoi)	Ceratopsidae indet. (53138)
Middle Turonian - Late/Upper Turonian	93.5 - 89.3	Uzbekistan (Navoi)	Ceratopsidae indet. (58168 75943) Monocloniinae indet. (58164)
Santonian	86.3 - 83.6	Kyrgyzstan (Jalal-Abad)	Ceratopsidae indet. (59597)
Santonian - Early/Lower Campanian	86.3 - 70.6	Kazakhstan (Qyzylorda)	Ceratopsidae indet. (59702)
Late/Upper Santonian	85.8 - 83.5	Canada (Alberta)	Brachyceratops sp. (48834)
Campanian	83.6 - 72.1	Mexico (Chihuahua)	Ceratopsidae indet. (170942) Chasmosaurinae indet. (226324)
Campanian	83.6 - 72.1	USA (Wyoming)	Ceratopsia indet. (45861) Ceratopsidae indet. (35367)
Campanian	83.6 - 72.1	USA (Montana)	Achelousaurus horneri (22652) Brachyceratops montanensis (50083) Brachyceratops sp., Styracosaurus sp. (22694) Centrosaurinae indet., Einiosaurus procurvicornis (59373) Monoclonius sp. (167423) Styracosaurus ovatus (45682) Styracosaurus sp., Centrosaurinae indet. (22693)
Campanian	83.6 - 72.1	China (Shandong)	Sinoceratops zhuchengensis (96815)
Campanian	83.6 - 72.1	Canada (Alberta)	Centrosaurus brinkmani (60364) Centrosaurus recurvicornis (64398) Chasmosaurus russelli (64401) Monoclonius cutleri (13340) Monoclonius sp. (52483 52485 70055)
Campanian - Maastrichtian	83.6 - 66.0	Kazakhstan (Aqmola)	Ceratopsidae indet. (53140)
Campanian - Maastrichtian	83.6 - 66.0	Kazakhstan (Almaty)	Ceratopsidae indet. (53143)
Campanian - Maastrichtian	83.6 - 66.0	USA (Colorado)	Centrosaurus sp. (189580)
Early/Lower Campanian	83.5 - 70.6	Mexico (Coahuila)	Ceratopsidae indet. (113768)
Early/Lower Campanian	83.5 - 70.6	USA (Utah)	Centrosaurinae indet. (84300 84301 230892)
Early/Lower Campanian	83.5 - 70.6	USA (Texas)	Ceratopsidae indet. (191055 200395 200399)
Early/Lower Campanian - Middle Campanian	83.5 - 70.6	USA (New Mexico)	Pachyrhinosaurinae indet. (13000 24924)
Early/Lower Campanian - Middle Campanian	83.5 - 70.6	USA (Utah)	Centrosaurinae indet. (230915)
Judithian	83.5 - 70.6	USA (Texas)	Ceratopsidae indet. (14488 14489) Chasmosaurus mariscalensis (48941 52462 52463 52464)
Judithian	83.5 - 70.6	USA (Utah)	Ceratopsidae indet. (75835)
Judithian	83.5 - 70.6	USA (Wyoming)	Ceratopsidae indet. (14458)
Late/Upper Campanian	83.5 - 70.6	Canada (Saskatchewan)	Ceratopsidae indet. (67885)
Late/Upper Campanian	83.5 - 70.6	USA (Arizona)	Crittendenceratops krzyzanowskii, Centrosaurinae indet. (68256)
Late/Upper Campanian	83.5 - 70.6	Mexico (Sonora)	Ceratopsidae indet. (46163)
Late/Upper Campanian	83.5 - 70.6	USA (Texas)	Agujaceratops sp. (94373) Pentaceratops sp., Brachyceratops sp. (191054)
Late/Upper Campanian	83.5 - 70.6	USA (Utah)	Centrosaurinae indet. (97803 192256) Ceratopsia indet. (97805 97810) Chasmosaurinae indet. (69799 97800 97801 97802 97806 97808 97809 97811 131868) Kosmoceratops richardsoni (97804)
Late/Upper Campanian	83.5 - 70.6	USA (Wyoming)	Ceratopsia indet. (45862)
Late/Upper Campanian	83.5 - 70.6	Mexico (Coahuila)	Ceratopsia indet. (55245) Ceratopsidae indet. (154244 170939 171003 227164 227175) Ceratopsidae indet., Centrosaurinae indet. (24261) Ceratopsidae indet., Ceratopsidae indet. (24258) Coahuilaceratops magnacuerna (97757) Dinosauria indet. (79611) Monoclonius sp. (51545)
Late/Upper Campanian	83.5 - 70.6	USA (New Mexico)	Ceratops sp. (62549 196540) Ceratopsia indet. (35518) Ceratopsidae indet. (47195 53982 62530 196541 196543 222078 222084 230007 230010) Ceratopsidae indet., Pentaceratops sp. (64094 64096) Chasmosaurinae indet. (138832 210631) Chasmosaurus sp. (53981) Monoclonius sp. (62528) Navajoceratops sullivani (138830) Pentaceratops fenestratus (39231) Pentaceratops sp. (46991) Pentaceratops sternbergi (52058) Pentaceratops sternbergii (34826 34827 34828 39229 196511 199764 222481) Torosaurus sp., Chasmosaurinae indet. (227186) Torosaurus utahensis (222091) Triceratops sp. (62533)
Late/Upper Campanian	83.5 - 70.6	USA (Montana)	Achelousaurus horneri, Stellasaurus ancellae (52435) Furcatoceratops elucidans (231025) Mercuriceratops gemini (156538) Spiclypeus shipporum (55274)
Late/Upper Campanian	83.5 - 70.6	Mexico (Chihuahua)	Centrosaurinae indet. (35707) Ceratopsidae indet. (210630 226998) Chasmosaurus mariscalensis (227230)
Late/Upper Campanian	83.5 - 70.6	Canada (Alberta)	Anchiceratops sp. (51317) Arrhinoceratops sp. (152403) Centrosaurus apertus (11894 58758 58945 59016 59018 59046 64427 65827 65895 65901 65903 65908 65925 65926 65928 65931 65932) Centrosaurus apertus, Chasmosaurus sp. (22659) Centrosaurus sp. (22657 58910 58952 59109 59110 60363 64336 65904 65906 67893 163047) Centrosaurus sp., Ceratops sp. (59025) Ceratops sp. (58950 59039) Ceratopsia indet. (59027 59228) Ceratopsidae indet. (22640 22695 22696 22697 22698 36264 45238 45260 45261 45264 45671 45674 45822 45892 45893 45897 45898 45899 46953 47784 47793 47807 47812 47814 47816 55317 65924 65927 67869 67870 67871 67872 67873 67874 67875 67876 67877 67881 67882 67883 67894) Ceratopsidae indet., Centrosaurinae indet., Centrosaurus apertus (221937) Chasmosaurus belli (45730 45731 58909 58954 59047 191258) Chasmosaurus brevirostris (58912) Chasmosaurus irvinensis (86515) Chasmosaurus kaiseni (35379) Chasmosaurus russelli (64400) Chasmosaurus sp. (22656 24891 24893 59006 59011 59014 59136 191255) Eoceratops canadensis (59026) Mercuriceratops gemini (156536) Monoclonius belli (60873) Monoclonius dawsoni, Monoclonius dawsoni (74657) Monoclonius flexus (58962 191270) Monoclonius nasicornus (13339) Monoclonius sp. (35511 55352 75476) Pachyrhinosaurinae indet. (97762) Pachyrhinosaurus sp. (22699 87125 220129) Styracosaurus albertensis (52487 65929 77259 77261 77265 77266 143007) Styracosaurus sp. (22655 59222) Triceratops horridus (162141)
Middle Campanian	83.5 - 70.6	USA (Utah)	Centrosaurinae indet. (178847)
Middle Campanian	83.5 - 70.6	Japan (Kagoshima)	Ceratopsidae indet. (143568)
Middle Campanian	83.5 - 70.6	Canada (Alberta)	Centrosaurinae indet. (69151) Ceratopsia indet. (59105) Ceratopsidae indet. (45240 45560 45561 45564 45565 45566 45567 45670 45820 47794 67880 67884 67892 181024) Nasutoceratopsini indet. (206061) Spinops sternbergorum (123351) Wendiceratops pinhornensis (171408) Xenoceratops foremostensis (135811)
Middle Campanian	83.5 - 70.6	USA (Montana)	Centrosaurus sp., Ceratopsidae indet. (14437) Judiceratops tigris (26779) Medusaceratops lokii, Styracosaurus albertensis (69152) Monoclonius crassus (46121) Monoclonius recurvicornis (228490)
Middle Campanian - Late/Upper Campanian	83.5 - 70.6	Mexico (Chihuahua)	Ceratopsidae indet., Chasmosaurus mariscalensis (35353)
Middle Campanian - Late/Upper Campanian	83.5 - 70.6	Canada (Alberta)	Anchiceratops sp. (51318) Centrosaurus sp. (53892) Ceratopsidae indet. (45819 45821 45894) Chasmosaurus russelli (64402) Monoclonius canadensis (60874) Monoclonius sp. (59118 64399 188846)
Middle Campanian - Late/Upper Campanian	83.5 - 70.6	USA (Texas)	Ceratopsidae indet. (68002 68006 150883 212185) Chasmosaurus mariscalensis (22709 25073)
Middle Campanian - Late/Upper Campanian	83.5 - 70.6	Mexico (Coahuila)	Ceratopsidae indet. (170937)
Middle Campanian - Late/Upper Campanian	83.5 - 70.6	Canada (Saskatchewan)	Ceratopsidae indet. (14494 229975) Chasmosaurinae indet. (75030)
Middle Campanian - Late/Upper Campanian	83.5 - 70.6	USA (Montana)	Avaceratops sp. (206062) Centrosaurus sp. (50909) Ceratops montanus (46123) Ceratopsidae indet. (14446 50906 142328 167801 167802) Chasmosaurus sp. (50907) Monoclonius sphenocerus (46122)
Late/Upper Campanian - Early/Lower Maastrichtian	83.5 - 66.0	Mexico (Coahuila)	Ceratopsidae indet. (227179) Chasmosaurinae indet., Ceratopsia indet. (226295) Chasmosaurus sp. (91053)
Late/Upper Campanian - Early/Lower Maastrichtian	83.5 - 66.0	USA (Colorado)	Chasmosaurinae indet. (76715)
Late/Upper Campanian - Early/Lower Maastrichtian	83.5 - 66.0	Canada (Alberta)	Anchiceratops longirostris (28548) Anchiceratops ornatus (191402) Anchiceratops sp., Pachyrhinosaurus canadensis (52072) Arrhinoceratops brachyops (67612) Ceratopsia indet. (64352 64629) Ceratopsidae indet. (162135 163050) Pachyrhinosaurus canadensis (52071 52249) Pachyrhinosaurus sp., Pachyrhinosaurus canadensis (34744)
Late/Upper Campanian - Late/Upper Maastrichtian	83.5 - 66.0	Mexico (Sonora)	Ceratopsidae indet. (170931)
Late/Upper Campanian - Maastrichtian	83.5 - 66.0	USA (New Mexico)	Ceratopsidae indet. (55291 55292 178746 178747) Torosaurus sp. (178745) Triceratops sp. (55286)
Maastrichtian	72.1 - 66.0	USA (Wyoming)	Ceratopsia indet. (165881) Triceratops sp. (225318 225319)
Maastrichtian	72.1 - 66.0	USA (North Dakota)	Ceratopsidae indet. (45100 45101 45104 45105 45106 45107 45113 45115 45116 45121 45125 45127 45129 45131 45132 45135 45139 45140 45142 45147 45297 45308 45311 45312 45313 45314 45320 45383 45384 45385) Torosaurus latus (45103)
Maastrichtian	72.1 - 66.0	USA (Texas)	Bravoceratops polyphemus (145482) Torosaurus sp. (52056) Torosaurus utahensis (227185)
Maastrichtian	72.1 - 66.0	USA (Montana)	Ceratopsia indet. (35510) Ceratopsidae indet. (85498 85499) Triceratops elatus (49016) Triceratops prorsus (173841) Triceratops sp. (38544 55353 55355 64428 64429 100852 109660 109661 142376 189572 228491) Ugrosaurus olsoni (53228)
Maastrichtian	72.1 - 66.0	Canada (Northwest Territories)	Ceratopsidae indet. (52893)
Maastrichtian	72.1 - 66.0	USA (Colorado)	Ceratopsidae indet. (49526 49537 49553 228914) Triceratops galeus (49525)
Maastrichtian	72.1 - 66.0	USA (South Dakota)	Torosaurus latus (53128) Triceratops horridus (225317 225924) Triceratops sp. (74175)
Early/Lower Maastrichtian	70.6 - 66.0	Canada (Alberta)	Anchiceratops longirostris (97797) Anchiceratops ornatus (64630 191401) Arrhinoceratops brachyops (156799) Ceratopsidae indet. (55363 69621 135739 135741) Ceratopsidae indet., Anchiceratops ornatus (76062) Pachyrhinosaurus canadensis (220119) Pachyrhinosaurus sp. (96455 96457 161248) Regaliceratops peterhewsi (191308)
Early/Lower Maastrichtian	70.6 - 66.0	Mexico (Coahuila)	Ceratopsidae indet. (219745)
Early/Lower Maastrichtian	70.6 - 66.0	USA (Alaska)	Ceratopsidae indet. (62631) Pachyrhinosaurus sp. (47787 51889)
Lancian	70.6 - 66.0	Canada (Saskatchewan)	Ceratopsidae indet. (14655) Triceratops sp. (14558 128461)
Lancian	70.6 - 66.0	USA (Montana)	Ceratopsidae indet. (14508 14513 14533 14536 14546 14559 14560 14562 14568 14574 14582 14613 14614 14624 14639 14653 14654) Ceratopsidae indet., Triceratops sp. (14516 14615 14616) Triceratops sp. (14550 14649 81041 81042 144502 179991)
Lancian	70.6 - 66.0	USA (South Dakota)	Ceratopsidae indet. (14571) Triceratops sp. (14537 192251)
Lancian	70.6 - 66.0	USA (North Dakota)	Ceratopsidae indet. (14610 14620 14621 137601 137704 137706 137707 137711 137712 137713 137715 137724) Ceratopsidae indet., Triceratops sp. (137710 137728) Triceratops sp. (224667)
Lancian	70.6 - 66.0	USA (Colorado)	Triceratops sp. (14640)
Lancian	70.6 - 66.0	USA (Wyoming)	Ceratopsia indet. (165877) Ceratopsidae indet. (45844) Chasmosaurinae indet. (14540 14541 14542 14543 14544 70341 70343 75280 75281 75284 75287 75289 75290 75292 75294 75297 75298 75429 75431 75432 75439 75441 75442 75443 75447) Triceratops sp. (14505 14519 14527 14538 14557 14579 14601 14609 165878)
Late/Upper Maastrichtian	70.6 - 66.0	USA (Wyoming)	Agathaumas sylvestris (46117) Ceratops horridus (46119) Ceratopsia indet. (52089) Ceratopsidae indet. (22638 46125 175972) Diceratops hatcheri (46133) Torosaurus gladius (46135) Torosaurus latus (46134) Triceratops brevicornus (46132 52683) Triceratops calicornis (50931) Triceratops elatus (46130 52682 52691 52692) Triceratops eurycephalus (52107) Triceratops flabellatus (46136 165876) Triceratops ingens (50930) Triceratops obtusus (46131) Triceratops prorsus (46127 46129 52690 52694 189513) Triceratops serratus (46126) Triceratops serratus, Ceratopsia indet. (52686) Triceratops sp. (14585 26760 45878 50086 50087 52088 52681 52684 52687 52688 52693 52695 54177 54179 57161 57162 57166 57168 142325 164991 164992 164993 191271 226054 226055) Triceratops sp., Triceratops sulcatus (52685) Triceratops sulcatus (46138 52689)
Late/Upper Maastrichtian	70.6 - 66.0	USA (South Dakota)	Ceratopsidae indet. (45390) Tatankaceratops sacrisonorum (97798) Torosaurus sp., Triceratops sp. (224484) Triceratops horridus (179969 224486) Triceratops sp. (50082 124934 150890 150892 150893 179970 216603 224488 224489 224490 224492)
Late/Upper Maastrichtian	70.6 - 66.0	USA (Texas)	Ceratopsidae indet. (52091) Pentaceratops sp. (85188) Torosaurus utahensis (69211)
Late/Upper Maastrichtian	70.6 - 66.0	Canada (Alberta)	Eotriceratops xerinsularis (75475) Leptoceratops sp., Iguanodontidae indet. (14569) Torosaurus sp. (47130) Triceratops albertensis (47127 47128)
Late/Upper Maastrichtian	70.6 - 66.0	USA (North Dakota)	Ceratopsidae indet. (45099 45102 45109 45110 45111 45112 45114 45117 45118 45120 45122 45124 45128 45130 45133 45134 45136 45137 45143 45144 45145 45299 45309 45381 45382 137720) Triceratops horridus (191374) Triceratops prorsus (24851) Triceratops sp. (45123 57919)
Late/Upper Maastrichtian	70.6 - 66.0	USA (Montana)	Ceratopsidae indet., Triceratops sp. (142326) Triceratops horridus (217183) Triceratops prorsus (14674 173842) Triceratops sp. (13142 55354 67129 72143 88990 99441 109659 157187 162855 179968 183109 183110 189003 221004 221005 221007 227182 227283)
Late/Upper Maastrichtian	70.6 - 66.0	USA (New Mexico)	Ceratopsidae indet. (59068 62529 62531 62532 62534 62546 138834 138836 171028 171031) Ceratopsidae indet., Ojoceratops fowleri (47819) Chasmosaurinae indet. (68042) Ojoceratops fowleri (97795 97796 138833 171027)
Late/Upper Maastrichtian	70.6 - 66.0	Canada (Saskatchewan)	Ceratopsidae indet. (152402) Chasmosaurinae indet. (52837) Triceratops prorsus (152400 152401 228825 228826 228827) Triceratops sp. (14555 52836 54105 152399 225845) Triceratops sp., Triceratops prorsus (45675)
Late/Upper Maastrichtian	70.6 - 66.0	USA (Utah)	Arrhinoceratops utahensis (48842) Ceratopsia indet. (45719) Ceratopsidae indet. (52926) Triceratops sp. (48549)
Late/Upper Maastrichtian	70.6 - 66.0	USA (Mississippi)	Ceratopsidae indet. (212621)
Late/Upper Maastrichtian	70.6 - 66.0	USA (Colorado)	Bison alticornis (46139) Ceratopsia indet. (49527) Ceratopsidae indet. (46210 49523 49528 49535 49539 49540 49541 49543 49547 49550 49551 49552 49555 49556 184700 219683) Dinosauria indet. (49533 49542) Polyonax mortuarius (49524) Triceratops sp. (49529 49531 49534 49536 49544 73927 182584)