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Bothremys
Taxonomy
Bothremys was named by Leidy (1865). Its type is Bothremys cookii. It is the type genus of Bothremydidae.
It was assigned to Chelonia by Leidy (1865); to Pleurodira by Hay (1908); to Bothremydidae by Hay (1930); to Pelomedusidae by Hay (1902), Thurmond and Jones (1981), Carroll (1988) and Zug (2001); and to Bothremydina by Gaffney et al. (2006), Gaffney et al. (2009) and Noto et al. (2022).
It was assigned to Chelonia by Leidy (1865); to Pleurodira by Hay (1908); to Bothremydidae by Hay (1930); to Pelomedusidae by Hay (1902), Thurmond and Jones (1981), Carroll (1988) and Zug (2001); and to Bothremydina by Gaffney et al. (2006), Gaffney et al. (2009) and Noto et al. (2022).
Species
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1865 | Bothremys Leidy p. 109 |
| 1902 | Bothremys Hay p. 438 |
| 1908 | Bothremys Hay p. 102 |
| 1930 | Bothremys Hay p. 74 |
| 1981 | Bothremys Thurmond and Jones p. 120 |
| 1988 | Bothremys Carroll |
| 2001 | Karkaemys Gaffney et al. p. 160 figs. 4-7 |
| 2001 | Bothremys Zug p. 207 |
| 2005 | Karkaemys Zalmout et al. |
| 2006 | Bothremys Gaffney et al. p. 79 |
| 2009 | Bothremys Gaffney et al. p. 1 |
| 2022 | Bothremys Noto et al. p. 3 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
G. †Bothremys Leidy 1865
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†Bothremys arabicus Zalmout et al. 2005
†Bothremys cooki Leidy 1865
†Bothremys kellyi Gaffney et al. 2006
†Bothremys maghrebiana Gaffney et al. 2006
Invalid names: Karkaemys Zalmout et al. 2005 [synonym]
Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| O. P. Hay 1908 | Vomer well developt, not separating the palatines. Choanae behind the centers of the orbits. The nasal passages underfloored by the surrounding bones. Crushing surfaces of jaws broad and occupying portions of the maxilla and of the palatines; the excavation contracting to a pit in each. Lower jaw with the dentaries co-ossified. Shell unknown. | |
| E. S. Gaffney et al. 2006 | A bothremydid pleurodire of the tribe Bothremydini with the following unique features: dorsal process of maxilla broad, separating apertura narium externa and orbital rim more than in any other Bothremydini; differs from Chedighaii in having well-developed maxilla-jugal pits on triturating surface; suborbital depth of maxilla deeper than in other Bothremydini except Chedighaii hutchisoni; orbits face strongly upward, in contrast to all other Bothremydini except Chedighaii; basisphenoid-quadrate contact very narrow, in contrast to all other Bothremydini except Chedighaii; lower jaw with deep pit on triturating surface. |
Measurements
No measurements are available
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| Source: subo = suborder, c = class, subp = subphylum | |||||
| References: Carroll 1988, Uetz 2005, Hendy et al. 2009 | |||||
Age range: base of the Santonian to the top of the Clarkforkian or 86.30000 to 54.90000 Ma
Collections (13 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Santonian | Jordan | Karkaemys arabicus (83312) | |
| Early/Lower Campanian | USA (New Jersey) | B. cooki (88610) | |
| Early/Lower Campanian - Middle Campanian | USA (Alabama) | Chedighaii barberi (129836) | |
| Early/Lower Campanian - Middle Campanian | USA (North Carolina) | B. cooki (129825) B. sp. (128412) B. sp., B. cooki (57918) | |
| Middle Campanian | USA (South Carolina) | B. sp. (180790) | |
| Maastrichtian | USA (New Jersey) | B. cookii (81999) | |
| Late/Upper Maastrichtian - Early/Lower Paleocene | USA (New Jersey) | B. sp. (26819) | |
| Danian | Morocco | B. maghrebiana (83311) | |
| Danian | Morocco (Khouribga) | B. maghrebiana (129831) | |
| Selandian | Morocco | B. kellyi (68176) | |
| Clarkforkian | USA (South Carolina) | B. sp. (15074) |