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Ustatochoerinae
Taxonomy
Ustatochoerinae was named by Stevens and Stevens (1996). Its type is Ustatochoerus.
It was assigned to Merycoidodontidae by Stevens and Stevens (1996), Stevens and Stevens (2007) and Stevens et al. (2023).
It was assigned to Merycoidodontidae by Stevens and Stevens (1996), Stevens and Stevens (2007) and Stevens et al. (2023).
Subtaxa
Mediochoerus, Ustatochoerus (type genus)
Synonymy list
Year | Name and author |
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1996 | Ustatochoerinae Stevens and Stevens p. 510 |
2007 | Ustatochoerinae Stevens and Stevens p. 164 |
2023 | Ustatochoerinae Stevens et al. p. 117 figs. Figure 95 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Subfm. †Ustatochoerinae Stevens and Stevens 1996
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G. †Mediochoerus Schultz and Falkenbach 1941
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†Mediochoerus blicki Schultz and Falkenbach 1941
†Mediochoerus johnsoni Schultz and Falkenbach 1941
†Mediochoerus mohavensis Schultz and Falkenbach 1941
G. †Ustatochoerus Schultz and Falkenbach 1941
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†Ustatochoerus californicus Merriam 1919
†Ustatochoerus leptoscelos Stevens et al. 1969
†Ustatochoerus major Leidy 1858
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Invalid names: Merychyus (Metoreodon) profectus Matthew and Cook 1909 [synonym], Ustatochoerus major texanus Schultz and Falkenbach 1941 [invalid subgroup], Ustatochoerus profectus espanolensis Schultz and Falkenbach 1941 [invalid subgroup], Ustatochoerus profectus nevadaensis Schultz and Falkenbach 1968 [invalid subgroup], Ustatochoerus profectus studeri Schultz and Falkenbach 1941 [invalid subgroup], Ustatochoerus skinneri Schultz and Falkenbach 1941 [synonym], Ustatochoerus skinneri santacruzensis Schultz and Falkenbach 1941 [invalid subgroup]
†Ustatochoerus medius Leidy 1858
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†Ustatochoerus medius mohavensis Schultz and Falkenbach 1941
†Ustatochoerus medius novomexicanus Frick 1929
Invalid names: Merychyus medius novomexicanus Frick 1929 [invalid subgroup], Merycochoerus coenopus Scott 1890 [synonym], Merycochoerus compressidens Douglass 1901 [synonym]
†Ustatochoerus tedfordi Stevens et al. 2023
Invalid names: Merychyus major major Leidy 1859 [invalid subgroup], Ustatochoerus californicus raki Schultz and Falkenbach 1941 [invalid subgroup]
Diagnosis
Reference | Diagnosis | |
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M. S. Stevens et al. 2023 | Oreodonts with relatively hypsodont cheek teeth, molars which narrow anteriorly, prominent labial cusps on P1 and P3, prominent labial cingula on the premolars except on the conical P1/p1, and a tendency towards early molarization of the premolars. The narial notch is retracted to at least the level of P4 (and sometimes M1), but there are no facial vacuities as in other taxa with retracted nasals. There is a fan-shaped occipital region, and a posterior extension of the palate toward the pterygoid region. |
Measurements
No measurements are available
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Source: f = family, o = order, subc = subclass, c = class, subp = subphylum | |||||
References: Carroll 1988, Hendy et al. 2009, Lander 1998, Lillegraven 1979, Nowak 1999 |
Age range: base of the Early/Lower Hemingfordian to the top of the Clarendonian or 20.43000 to 9.40000 Ma
Collections (120 total)
Time interval | Ma | Country or state | Original ID and collection number |
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Arikareean | USA (California) | Phenacocoelus leptoscelos (17800) | |
Arikareean | USA (Texas) | Phenacocoelus leptoscelos (178057 178059 178062) | |
Harrisonian | USA (Texas) | Hypsiops leptoscelos (17768) Ustatochoerus leptoscelos (17836) | |
Miocene | USA (Nebraska) | Ustatochoerus major (18189 18289 18400) Ustatochoerus major, Ustatochoerus medius (18049) Ustatochoerus medius (18264 18311 18319 18356 18469) Ustatochoerus profectus (18050 18136 18137) Ustatochoerus skinneri (18224) | |
Miocene | USA (New Mexico) | Ustatochoerus medius (19518) | |
Miocene | USA (South Dakota) | Ustatochoerus medius (18191) | |
Miocene | USA (Nevada) | Ustatochoerus profectus (18989) | |
Miocene | USA (Texas) | Ustatochoerus profectus (18103) | |
Miocene | USA (Kansas) | Ustatochoerus medius, Ustatochoerus profectus (18122) | |
Early/Lower Hemingfordian | USA (Nebraska) | Mediochoerus johnsoni (17946) Ustatochoerus tedfordi (17873) | |
Barstovian | USA (Nebraska) | Ustatochoerus medius (18031 18101 18124 18125 18141 18155 18164 18192 18216 18275 18281 18284 18442 18443) Ustatochoerus medius, Ustatochoerus profectus (18123) Ustatochoerus profectus (18119 18464) | |
Barstovian | USA (New Mexico) | Ustatochoerus californicus, Ustatochoerus medius (19535) Ustatochoerus medius (19287) Ustatochoerus novomexicanus, Ustatochoerus skinneri (19470) | |
Barstovian | USA (Colorado) | Ustatochoerus medius (18203 18220) | |
Barstovian | USA (California) | Ustatochoerus medius (19364) | |
Early/Lower Barstovian | USA (Nebraska) | Mediochoerus blicki (17972) | |
Barstovian - Clarendonian | USA (Colorado) | Ustatochoerus medius (18445) | |
Barstovian - Clarendonian | USA (Nebraska) | Merychyus major, Merychyus medius (190543) Ustatochoerus medius (18077 18270 18280 18290) Ustatochoerus medius, Ustatochoerus profectus (18070) | |
Clarendonian | USA (California) | Merycochoerus californicus (189022) Ustatochoerus californicus (19322 19323 19384 19417 19517) Ustatochoerus major (19185) Ustatochoerus profectus (19383 19391) | |
Clarendonian | USA (South Dakota) | Metoreodon profectus (18063) Ustatochoerus profectus (18200) Ustatochoerus skinneri (18268 18429) | |
Clarendonian | USA (Nebraska) | Ustatochoerus major (18028 18099 18135 18145 18156 18214 18218 18221 18227 18231 18300 18333 18425 18430 18458 18468 18496) Ustatochoerus medius (18260) Ustatochoerus profectus (18042 18060 18095 18204 18251 18301 18335 18362 18447) Ustatochoerus skinneri (18092 18315 18324 188779) | |
Clarendonian | USA (Oklahoma) | Ustatochoerus profectus (18045) | |
Clarendonian | USA (Texas) | Oreodontidae indet. (18645) Ustatochoerus major, Ustatochoerus profectus (18096) Ustatochoerus medius, Metoreodon sp. (18623) Ustatochoerus profectus (18104 18244 18355) | |
Late/Upper Clarendonian | USA (Nebraska) | Ustatochoerus major (18066) Ustatochoerus medius (18207) | |
Late/Upper Clarendonian | USA (New Mexico) | Merychyus major (157331) | |
Clarendonian - Hemphillian | USA (Nebraska) | Merychyus major (18295) Ustatochoerus major (18180) Ustatochoerus profectus (18069 18134) | |
Clarendonian - Hemphillian | USA (Wyoming) | Ustatochoerus medius (18424) | |
Tortonian | USA (California) | Ustatochoerus californicus (19466) Ustatochoerus major (52210) | |
Hemphillian | USA (Nebraska) | Ustatochoerus major (18033 18499) |