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Scombrimorpharia

Actinopteri

Taxonomy
Scombrimorpharia is an unranked clade. It is extant.

It was assigned to Percomorphaceae by Betancur et al. (2013).

Synonymy list
YearName and author
2013Scombrimorpharia Betancur et al.

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
superclassActinopterygii()
classActinopteri()
subclassNeopterygii(Regan 1923)
Teleosteomorpha
RankNameAuthor
Teleostei(Müller 1846)
Osteoglossocephalai
ClupeocephalaPatterson and Rosen 1977
Euteleosteomorpha
NeoteleosteiRosen 1973
Eurypterygia(Rosen 1973)
Ctenosquamata
Acanthomorphata(Rosen 1973)
Euacanthomorphacea
Percomorphaceae
Scombrimorpharia
Scombrimorpharia

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

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G. †Isurichthys Woodward 1901
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Isurichthys breviusculus Bannikov 2012
Isurichthys orientalis Woodward 1901
Invalid names: Isurus Agassiz 1842 [replaced]
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Subfm. Braminae
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G. †Paucaichthys Baciu and Bannikov 2003
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Paucaichthys neamtensis Baciu and Bannikov 2003
G. †Bramoides Casier 1966
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Bramoides brieni Casier 1966
G. †Digoria Danil'chenko 1980
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Digoria ambigua Danil'chenko 1980
G. †Goniocranion Casier 1966
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Goniocranion arambourgi Casier 1966
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G. †Absalomichthys Whitley 1933
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Absalomichthys velifer Jordan 1925
G. †Chalcidichthys David 1943
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G. †Centrolophidarum Gill 1861
G. Centrolophus Lacépède 1803
G. Hyperoglyphe Guenther 1859
G. Mupus Cocco 1833
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Mupus neumanni Schwarzhans 1974
Mupus sinuosus Stinton 1965
G. †Zorzinia Bannikov 2000
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Zorzinia postalensis Bannikov 2000
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G. †Bannikovichthys Carnevale 2007
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Bannikovichthys paelignus Carnevale 2007
G. Pseudoscopelus Luetken 1892
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Invalid names: Myersiscus Fowler 1933 [synonym]
Fm. Gempylidae Gill 1862
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G. †Abadzekhia Bannikov 1985
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Abadzekhia marinae Bannikov 1985
Abadzekhia tarletskovi Bannikov 2005
G. †Chelifichthys Carnevale 2006
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Chelifichthys goujeti Carnevale 2006
G. Diplospinus Maul 1948
G. †Gempylidarum Unknown
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Gempylidarum merus Schwarzhans 2003
G. †Nealotus Johnson 1865
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Nealotus tripes Johnson 1865
G. †Progempylus Casier 1966
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Progempylus edwardsi Casier 1966
G. †Thyrsites Lesson 1831
G. Thyrsitoides Fowler 1929
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G. Ariomma Jordan and Snyder 1904
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Ariomma geslini Carnevale and Bannikov 2006
G. Cubiceps Lowe 1843
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Cubiceps pauciradiatus Günther 1872
G. †Psenicubiceps Danil'chenko 1980
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Psenicubiceps alatus Danil'chenko 1980
G. †Rybapina Bannikov
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G. †Carangopsis Agassiz 1834
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Carangopsis analis Agassiz 1835
Carangopsis brevis Blainville 1818
Carangopsis dorsalis Agassiz 1834
Carangopsis latior Agassiz 1835
Carangopsis maximus Agassiz 1835
G. †Lophar Jordan and Gilbert 1919
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Lophar miocaenus Jordan and Gilbert 1919
G. Pomatomus Lacépède 1802 [bluefish]
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Pomatomus saltatrix Linnaeus 1766
G. †Pseudoseriola David 1943
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G. †Sullivanichthys Weems 1999
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Fm. Scombridae Rafinesque 1815
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G. Acanthocybium Gill 1862
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Acanthocybium proosti Storms 1897
Acanthocybium solandri Cuvier 1831
G. †Acanthonotos Sauvage 1870
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Acanthonotos armatus Sauvage 1870
G. †Ardiodus White 1931
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Ardiodus mariotti White 1931
G. Cariniceps Owen 1854
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G. †Caucombrus Bannikov 2020
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Caucombrus histiopterygius Bannikov 2020
G. †Cybium Cuvier 1829
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Cybium bartonense Woodward 1901
Cybium bleekeri Storms 1892
Cybium excelsum Woodward 1901
Cybium loczyi Weiler 1938
Cybium macropomum Agassiz 1839
G. †Eoscomber Claeson et al. 2015
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Eoscomber senegalicus Claeson et al. 2015
G. †Eoscombrus David 1946
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Eoscombrus chaneyensis David 1946
G. †Eucynodus Owen 1854
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G. Gasterochisma Richardson 1845
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Gasterochisma melampus Richardson 1845
G. †Gasteronemus Agassiz 1833
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Gasteronemus oblongus Agassiz 1833
Gasteronemus rhombeus Volta 1796
G. †Godsilia Monsch 2006
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Godsilia lanceolata Agassiz 1835
G. †Lepidocybium Gill
G. †Ocystias Jordan and Gilbert 1920
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Ocystias sagitta Jordan and Gilbert 1920
G. †Pelamys Cuvier and Valenciennes 1831
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Pelamys delheidi Leriche 1905
Pelamys robusta Van Beneden 1871
G. Pneumatophorus Jordan and Gilbert 1883
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Pneumatophorus euodus Nolf 1972
G. †Pseudauxides Monsch 2006
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Pseudauxides speciosus Agassiz 1835
Tr. †Sardini Starks 1910
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G. Cybiosarda Whitley 1935
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Cybiosarda elegans Whitley 1935
G. Gymnosarda Gill 1863
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Gymnosarda disodilica Ciobanu 1977
Gymnosarda prisca Monsch 2000
Gymnosarda unicolor Ruppell 1838
Gymnosarda zsivnyi Weiler 1938
G. Orcynopsis Gill 1862
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Orcynopsis unicolor Geoffroy St Hilaire 1817
G. Sarda Lacépède 1802
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Sarda australis Macleay 1881
Sarda chiliensis Cuvier 1832
Sarda delheidi Leriche 1905
Sarda memorabilis Danil'chenko 1980
Sarda orientalis Temminck and Schlegel 1844
Sarda rara Bannikov 1979
Sarda remota Danil'chenko 1980
Sarda roulei Arambourg 1927
Sarda sarda Bloch 1791
Sarda stockii David 1943
G. Scomber Linnaeus 1758
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Scomber caucasica Bogachev 1933
Scomber colias Gmelin 1789
Scomber cubanicus Danil'chenko 1960
Scomber gnarus Bannikov 1979
Scomber japonicus Houttuyn 1782
Scomber oligocenicus Pauca 1929
Scomber sanctaemonicae Jordan 1919
Scomber scombrus Linnaeus 1758
Scomber voitestii Pauca 1929
G. †Scomberodon Van Beneden 1871
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Scomberodon dumontii Van Beneden 1871
G. †Scombridarum Rifinesque 1815
Subfm. †Scombrinae Rafinesque-Schmaltz 1815
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Tr. †Eocoelopomini Monsch and Bannikov 2011
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G. †Eocoelopoma Woodward 1901
G. †Palaeothunnus Bannikov 1978
Tr. Scomberomorini Starks 1910
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G. †Neocybium Leriche 1906
G. †Palaeocybium Monsch 2004
G. Scomberomorus Lacépède 1802
Tr. †Scombrini Rafinesque-Schmaltz 1815
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G. †Auxides Jordan 1919
G. Rastrelliger Jordan and Starks 1908
G. †Scombrinus Woodward 1901
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Scombrinus nuchalis Woodward 1901
Tr. Thunnini Starks 1910
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G. Allothunnus Serventy 1948
G. Auxis Cuvier 1829
G. †Eothynnus Woodward 1901
G. Euthynnus Jordan and Gilbert 1882
G. Katsuwonus Kishinouye 1915
G. Thunnus South 1845
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Invalid names: Albacora Jordan 1888 [synonym], Orcynus Agassiz 1835 [replaced], Thynnus Cuvier 1816 [objective synonym]
G. †Woodwardella Casier 1966
G. †Scombrosarda Danilt'chenko 1962
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Scombrosarda limata Danil'chenko 1980
G. †Thunnoscomberoides Monsch 2006
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G. Toxopoma Owen 1854
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Toxopoma politum Owen 1854
G. †Tunita Jordan and Gilbert 1919
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Tunita octavia Jordan and Gilbert 1919
G. †Turio Jordan and Gilbert 1920
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Turio wilburi Jordan and Gilbert 1920
G. †Wetherellus Casier 1966
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Wetherellus brevior Casier 1966
Wetherellus cristatus Agassiz 1844
Wetherellus longior Casier 1966
G. †Xestias Jordan 1920
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Xestias iratus Jordan 1920
Invalid names: Acanthocybiinae Starks 1910 [empty], Sardinae Starks 1910 [empty]
Subor. †Scombroidei Bleeker 1859
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G. †Duplexdens Monsch 2004
Fm. †Euzaphlegidae Daniltshenko 1960
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Subfm. †Dipterichthyinae Arambourg 1967
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G. †Palimphyes Agassiz 1835
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Invalid names: Krambergeria Simonescu 1905 [synonym]
G. †Euzaphleges White and Moy-Thomas 1941
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Euzaphleges longurio Jordan and Gilbert 1920
G. †Lednevia Danil'chenko 1960
G. †Thyrsocles Jordan and Gilbert 1920
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Thyrsocles escharion Jordan and Gilbert 1920
Thyrsocles kriegeri Jordan and Gilbert 1919
Invalid names: Thyrsion Jordan and Gilbert 1920 [synonym]
G. †Trossulus Jordan 1921
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Trossulus exoletus Jordan 1921
G. †Veronaphleges Bannikov 2008
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Veronaphleges brunae Bannikov 2008
G. †Zaphlegulus David 1943
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G. †Micrornatus Monsch 2004
G. †Scombramphodon Woodward 1901
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Scombramphodon crassidens Agassiz 1844
Superfm. †Scombroidea Rafinesque-Schmaltz 1815
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Fm. †Cybiidae
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G. †Escharion Jordan 1925
G. †Starrias Jordan 1925
G. †Sphyraenodus Agassiz 1839
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Sphyraenodus crassidens Agassiz 1844
Sphyraenodus hastatus Bohm 1926
Sphyraenodus lerichei Casier 1944
Sphyraenodus priscus Agassiz 1839
Sphyraenodus rupeliensis Dollo and Storms 1888
Sphyraenodus speciosus Leidy 1856
Invalid names: Dictyodus Owen 1839 [nomen dubium]
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Fm. †Zaphlegidae David 1943
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G. †Zaphleges Jordan and Gilbert 1920
Superfm. Trichiuroidea Rafinesque-Schmaltz 1810
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G. †Argestichthys Prokofiev 2002
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Argestichthys vysotzkyi Prokofiev 2002
G. †Hemithyrsites Sauvage 1873
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Hemithyrsites alatus Sauvage 1870
Hemithyrsites armatus Sauvage 1873
Hemithyrsites maicopicus Danil'chenko 1960
Superfm. †Xiphioidea Rafinesque-Schmaltz 1810
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Fm. †Hemingwayidae Sytchevskaya and Prokofiev 2002
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G. †Hemingwaya Sytchevskaya and Prokofiev 2002
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G. †Aglyptorhynchus Casier 1966
G. †Enniskillenus Casier 1966
G. †Homorhynchus Van Beneden 1873
G. †Palaeorhynchus de Blainville 1818
G. †Pseudotetrapturus Danil'chenko 1960
Invalid names: Aglyptorhynchinae Fierstine and Weems 2009 [empty]
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G. Pseudohistiophorus Buen 1950
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G. Scombrolabrax Roule 1921
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G. †Pinichthys Bannikov 1985
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Pinichthys fractus Bannikov 1985
Pinichthys shirvanensis Bannikov 2021
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G. Tetragonurus Risso 1810
Fm. Thunnidae Starks 1910
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G. †Pinulothunnus Ciobanu 1977
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G. †Anenchelum de Blainville 1818
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Anenchelum angustum Danil'chenko 1980
Anenchelum dorsale Agassiz 1842
Anenchelum eocaenicus Danilt'chenko 1962
Anenchelum glarisianum de Blainville 1818
Anenchelum heteropleurum Agassiz 1842
Anenchelum isopleurum Agassiz 1842
Anenchelum latum Agassiz 1842
Anenchelum lednevi Menner 1949
Anenchelum paucivertebrale Bannikov and Parin 1995
G. Aphanopus Lowe 1839
G. Benthodesmus Goode and Bean 1882
G. Lepidopus Gouan 1770
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Lepidopus albyi Sauvage 1870
Lepidopus anguis Sauvage 1870
Lepidopus caudatus Euphrasen 1788
Lepidopus gouanianus Lacépède 1800
Lepidopus lateralis Danil'chenko 1980
Lepidopus lerichei Casier 1958
Lepidopus proargenteus Arambourg 1927
G. †Trichiuridarum Rafinesque 1810
Subfm. Trichiurinae Rafinesque-Schmaltz 1815
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G. †Eutrichiurides Casier 1944
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Eutrichiurides winkleri Casier 1946
G. Trichiurides Winkler 1874
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Trichiurides sagittidens Winkler 1874
G. Trichiurus Linnaeus 1758
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Trichiurus lepturus Linnaeus 1758
Trichiurus miocaenus Delfortrie 1876
Trichiurus plicidens Arambourg 1952
Invalid names: Amarsipidae [empty], Icosteidae [empty]
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G. †Aulorhamphus Zigno 1890
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Aulorhamphus bolcensis de Zigno 1887
Aulorhamphus capellinii de Zigno 1887
Aulorhamphus chiarasorbiniae Bannikov and Tyler 2011
G. †Pesciarhamphus Bannikov and Tyler 2011
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Pesciarhamphus carnevalei Bannikov and Tyler 2011
G. †Veronarhamphus Tyler 2004
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Veronarhamphus canossae Heckel 1856
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G. †Eoaulostomus Blot 1981
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Eoaulostomus bolcensis Blainville 1818
Eoaulostomus gracilis Blot 1980
G. †Jungersenichthys Blot 1981
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G. †Macroaulostomus Blot 1981
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G. †Synhypuralis Blot 1981
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G. †Tyleria Parin 1993
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Tyleria necopinata Parin 1993
Invalid names: Aulostomes [replaced]
Superfm. †Aulostomoidea
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G. †Aulostomoides Blot 1981
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Aulostomoides tyleri Blot 1980
Fm. †Eekaulostomidae Cantalice and Alvarado-Ortega 2016
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G. †Eekaulostomus Cantalice and Alvarado-Ortega 2016
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Eekaulostomus cuevasae Cantalice and Alvarado-Ortega 2016
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G. Callionymus Linnaeus 1758
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Callionymus eocaenus Casier 1946
Callionymus hakatarameaensis Schwarzhans 2019
Callionymus macrocephalus Gorjanovic-Kramberger 1882
Callionymus triquetrus Schwarzhans 2019
G. †Protonymus Sytchevskaya and Prokofiev 2007
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Protonymus miocenicus Pobedina 1954
Protonymus primus Weiler 1943
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Invalid names: Protonymus gontsharovae Sytchevskaya and Prokofiev 2007 [synonym]
G. †Gilmourella Carnevale and Bannikov 2019
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Gilmourella minuta Carnevale and Bannikov 2019
Fm. Centriscidae Rafinesque 1826
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G. †Aeoliscoides Blot 1981
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Aeoliscoides longirostris Blainville 1818
G. Aeoliscus Jordan and Starks 1902
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Aeoliscus heinrichi Heckel 1850
Aeoliscus strigatus Güther 1861
G. †Amphisile Cuvier 1816
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Amphisile teleajensis jonet 1949
G. Centriscus Linnaeus 1758
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Centriscus heinrichi Heckel 1850
Centriscus longispinus Rozhdestvenski 1949
G. †Paramphisile Blot 1981
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Paramphisile weleri Blot 1980
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G. †Fistularioides Blot 1981
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G. †Pseudosyngnathus Kner and Steindachner 1863
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G. †Macrorhamphosidarum Bleeker 1879
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Subor. Mulloidei
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G. Mullus Linnaeus 1758
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Mullus bifurcatus Strashimirov 1972
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Invalid names: Mullus fuscatus Rafinesque 1966 [synonym]
Mullus elongatus Steurbaut 1984
Mullus moldavicus Switchenska 1959
Invalid names: Creediidae [empty]
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G. †Paraeoliscus Blot 1981
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G. †Parasynarcualis Blot 1981
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Parasynarcualis longirostris Blainville 1818
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G. †Ramphosus Agassiz 1835
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Ramphosus aculeatus Blainville 1818
Ramphosus biserratus Bassani 1876
Ramphosus rastrum Volta 1796
Rhamphosus biserratus Bassani 1876
Rhamphosus rastrum Volta 1796
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Subfm. †Solenorhynchinae Bannikov and Carnevale 2017
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G. †Calamostoma Agassiz 1833
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Invalid names: Calamostoma breviculum Blainville 1818 [synonym]
G. †Solenorhynchus Heckel 1853
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Solenorhynchus elegans Heckel 1854
G. Solenostomus Gronovius 1763
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Superfm. †Centriscoidea
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Fm. †Gerpegezhidae Bannikov and Carnevale 2012
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G. †Gerpegezhus Bannikov and Carnevale 2012
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G. Fistularia Linnaeus 1758
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Fistularia contermina Danil'chenko 1960
Fistularia koenigii Agassiz 1839
Fistularia licatae Sauvage 1880
Fistularia tenuirostris Agassiz 1835
Infraor. Macrorhamphosa Pietsch 1978
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Invalid names: Macrorhamphosoidea Pietsch 1978 [empty]
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G. Acanthognathus Duncker 1912
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Acanthognathus incolumis Danil'chenko 1960
G. Bryx Herald 1940
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Subg. Bryx (Bryx) Herald 1940
Subg. Bryx (Simocampus) Fritzsche 1980
Bryx veleronis Herald 1940
G. Doryrhamphus Kaup 1853
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Doryrhamphus paulus Fritzsche 1980
Doryrhamphus squalidus Danil'chenko 1960
Invalid names: Pristidoryrhamphus Fowler 1944 [synonym]
G. Dunckerocampus Whitley 1933
Subfm. †Eogastrophinae Jerzmanska 1968
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G. †Hipposyngnathus Danil'chenko 1960
G. Hippocampus Rafinesque 1810 [sea horse]
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Hippocampus ingens Girard 1859
Hippocampus sarmaticus Zalohar et al. 2009
Hippocampus slovenicus Zalohar et al. 2009
Invalid names: Farlapiscus Whitley 1931 [synonym], Hippohystrix Whitley 1940 [synonym]
G. Leptonotus Kaup 1853
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Leptonotus blainvilleanus Eydoux and Gervais 1837
G. †Prosolenostomus Blot 1981
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Subfm. †Pshekhagnathinae Bannikov et al. 2017
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G. †Pshekhagnathus Bannikov et al. 2017
Subfm. †Syngnathinae
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G. †Maroubrichthys Parrin 1992
G. †Nepigastrosyngnathus Pharisat 1993
G. Syngnathus Linnaeus 1758 [pipefish]
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Syngnathus albyi Sauvage 1870
Syngnathus altus Danil'chenko 1960
Syngnathus avus Jordan and Gilbert 1919
Syngnathus breviculus Blainville 1818
Syngnathus carinatus Gilbert 1892
Syngnathus emeritus Fritzsche 1980
Syngnathus euchrous Fritzsche 1980
Syngnathus incertus Danil'chenko 1960
Syngnathus incompletus Cosmovici 1887
Syngnathus insulae Fritzsche 1980
Syngnathus macrobrachium Fritzsche 1980
Syngnathus opisthopterus Agassiz 1833
Invalid names: Dermatostethus Gill 1863 [synonym], Siphonostoma Rafinesque 1810 [synonym], Siphostoma Rafinesque 1810 [synonym], Syrictes Jordan and Evermann 1927 [synonym]
Invalid names: Hipposyngnathinae Fritzsche 1980 [empty], Syngnathus bolcensis de Zigno 1887 [invalid subgroup], Syngnathus heckeli de Zigno 1874 [invalid subgroup]
Invalid names: Macroramphosa Pietsch 1978 [empty], Pegasoidea [empty], Syngnatha Pietsch 1978 [empty], Syngnathoidea Pietsch 1978 [empty]
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G. †Urosphen Agassiz 1835
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Urosphen dubius Blainville 1818
Urosphen fistularis Agassiz 1835
Invalid names: Pegasidae [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: marinesubp
Locomotion: actively mobileo
Life habit: nektonico
Diet: omnivoreo
Created: 2009-07-20 20:44:22
Modified: 2009-07-20 22:44:22
Source: o = order, subp = subphylum
References: Hendy et al. 2009, Carroll 1988

Age range

Maximum range based only on fossils: base of the Late/Upper Santonian to the top of the Holocene or 85.80000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 83.5 Ma

Collections (301 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Late/Upper Santonian85.8 - 83.5Lebanon Solenognathus lineolatus (221226)
Late/Upper Maastrichtian70.6 - 66.0India (Manharashtra) Sphyraenodus sp. (63896)
Selandian61.6 - 59.2Denmark Mupus sinuosus (159792 159793)
Selandian - Thanetian61.6 - 56.0Ukraine Scombridarum sp. (159509)
Thanetian59.2 - 56.0USA (Maryland) Cybium sp. (84045)
Thanetian59.2 - 56.0USA (New Jersey) Sphyraenodus speciosus (3113)
Ypresian56.0 - 47.8United Kingdom (England) Aglyptorhynchus venablesi (8050) Eocoelopoma sp. (76601) Scombrinus nuchalis (185905) Sphyraenodus pirscus, Trichiurides sagittidens, Aglyptorhynchus venablesi, Eothynnus salmoneus, Scombrinus nuchalis, Gymnosarda prisca, Eutrichiurides winkleri, Progempylus edwardsi, Cybium proosti, Enniskillenus radiatus, Scombramphodon crassidens, Wetherellus cristatus, Woodwardella patellifrons, Thunnini indet., Bramoides brieni, Goniocranion arambourgi, Wetherellus brevior, Wetherellus longior, Scombramphodon sheppeyensis, Scombrinus macropomus, Aglyptorhynchus sulcatus (13299)
Ypresian56.0 - 47.8United Kingdom Eutrichiurides winkleri (160682 185917)
Ypresian56.0 - 47.8India (Gujarat) Eutrichiurides sp. (71438)
Ypresian56.0 - 47.8USA (Maryland ) Cybium sp., Sarda delheidi (124192)
Ypresian56.0 - 47.8Belgium (Brabant) Pelamys sp. (188444)
Ypresian56.0 - 47.8Italy Calamostoma breviculum, Carangopsis dorsalis, Auxides propterygius, Rhamphosus rastrum, Rhamphosus biserratus, Urosphen dubius, Eoaulostomus bolcensis, Eoaulostomus gracilis, Synhypuralis jungerseni, Synhypuralis banisteri, Jungensenichthys elongatus, Macraulostomus veronensis, Tyleria necopinata, Parasynarcualis longirostris, Fistularioides veronensis, Fistularioides phyllolepis, Pseudosyngnathus opisthopterus, Aulostomoides tyleri, Aulorhamphus bolcensis, Aulorhamphus capellinii, Aulorhamphus chiarasorbiniae, Veronarhamphus canossae, Pesciarhamphus carnevalei, Paraeoliscus robinetae, Paramphisile weleri, Syngnathus heckeli, Syngnathus bolcensis, Prosolenostomus lessinii, Solenorhynchus elegans, Carangopsis brevis, Veronaphleges brunae, Thunnoscomberoides bolcensis, Godsilia lanceolata, Palaeorhynchus zorzini, Zorzinia postalensis, Ramphosus rastrum, Ramphosus biserratus, Calamostoma lesiniforme, Gilmourella minuta, Aeoliscoides longirostris (94830) Orcynus lanceolatus, Orcynus latior, Thynnus bolcensis (55239)
Ypresian56.0 - 47.8Italy (Vicenze) Zorzinia postalensis (127545)
Ypresian56.0 - 47.8Russian Federation (Kabardino-Balkarian) Gerpegezhus paviai (218152)
Ypresian56.0 - 47.8Senegal Auxides huberti, Eoscomber senegalicus (190952)
Ypresian56.0 - 47.8Turkmenistan Palimphyes palaeocenicus, Argestichthys vysotzkyi, Auxides turkmenicus, Palaeothunnus parvidentatus, Scomberomorus avitus, Hemingwaya sarissa, Eocoelopoma portentosum (186444)
Ypresian56.0 - 47.8Belgium Cybium proosti (219003 219008) Cybium proosti, Cybium stormsi, Sphyraenodus lerichei, Cybium bleekeri (219005) Scomber sp., Eutrichiurides winkleri, Cybium proosti, Cybium stormsi, Sphyraenodus lerichei (219006) Scomber sp., Trichiurides sagittidens, Eutrichiurides winkleri, Cybium proosti, Glyptorhynchus sulcatus, Cybium stormsi, Sphyraenodus lerichei, Cybium bleekeri, Pelamys delheidi, Callionymus eocaenus (219001)
Ypresian56.0 - 47.8Italy (Verona) Euzaphlegidae n. gen. n. sp, Anenchelum sp., Thyrsitoides sp. (190957)
Ypresian - Lutetian56.0 - 41.3USA (Alabama) Gymnosarda sp., Scomberomorus sp., Trichiurides sagittidens, Scomberomorus bleekeri, Scombridae indet., Palaeocybium proosti, Trichiurus oshosunensis, Eutrichiurides plicidens, Scomberomorus stormsi (81179) Scomberomorus bleekeri, Scomberomorus stormsi (209524) Trichiurus sp., Trichiurides sp. (83330)
Eocene56.0 - 33.9USA (Alabama) Trichiurus sp. (83333)