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Hardella thurjii

Reptilia - Testudines - Geoemydidae

Taxonomy
Emys thurgi was named by Gray (1831). It is extant.

It was recombined as Hardella thurgi by Lydekker (1889) and Swinton (1934); it was recombined as Hardella thurjii by van Dijk et al. (2014) and Garbin et al. (2020).

Synonyms
  • Clemmys watsoni was named by Lydekker (1886). Its type specimen is number not provided, a partial shell, and it is a 3D body fossil. Its type locality is Piram Island, which is in a Miocene coastal conglomerate in the Piram Conglomerate Formation of India.

    It was synonymized subjectively with Hardella thurjii by Garbin et al. (2020).
  • Batagur falconeri was named by Lydekker (1885). Its type specimen is BMNH 39835, a partial shell, and it is a 3D body fossil. Its type locality is Siwalik Hills, which is in a Pliocene fluvial horizon in the Pinjor Formation of Pakistan.

    It was synonymized subjectively with Hardella thurjii by Garbin et al. (2020).
Synonymy list
YearName and author
1831Emys thurgi Gray p. 22
1885Batagur falconeri Lydekker p. 187
1886Clemmys watsoni Lydekker p. 541 figs. Pl. XV
1889Hardella thurgi Lydekker pp. 120-123
1934Hardella thurgi Swinton p. 70
2014Hardella thurjii van Dijk et al.
2020Hardella thurjii Garbin et al. p. 25

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
RankNameAuthor
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
Testudinata(Oppel 1811)
orderTestudinesBatsch 1788
suborderCryptodira
Pantestudinoidea
superfamilyTestudinoidea
familyGeoemydidaeTheobald 1868
Palatochelydia
genusHardellaGray 1870
speciesthurjii(Gray 1831)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Hardella thurjii Gray 1831
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Invalid names: Batagur falconeri Lydekker 1885 [synonym], Clemmys watsoni Lydekker 1886 [synonym]
Diagnosis
ReferenceDiagnosis
R. Lydekker 1886 (Clemmys watsoni)The species may be defined from the characters of the shell as follows: Shell moderately vaulted, globose, broad; posterior margin entire; nuchal plate narrow, and broadest posteriorly; first vertebral narrowest anteriorly, and showing a tendency to a bellshape ; second and third vertebrals hexagonal, fourth short, narrowest posteriorly, and giving off a process jutting into the posterior border of the third; an interrupted vertebral, but no costal keel; costal areolse well-marked; suture separating the postgulars much shorter than the intergular suture.
R. Garbin et al. 2020Hardella thurjii can be differentiated from other geoemydids by large carapace size (up to 60 cm in females), presence of hexagonal, nearly square second to fourth vertebrals, a first vertebral scute that is wider posteriorly, presence of an inflection at the margin of the gulohumeral sulcus, and an entoplastron that is not intersected by the humeropectoral sulcus.
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: freshwatersubo
Locomotion: actively mobilec
Life habit: aquaticsubo
Diet: omnivoresubo
Reproduction: oviparoussubo
Created: 2005-08-26 14:49:00
Modified: 2005-08-26 16:49:00
Source: subo = suborder, c = class, subp = subphylum
References: Carroll 1988, Hendy et al. 2009, Uetz 2005

Age range: base of the Late/Upper Miocene to the top of the Late/Upper Pliocene or 11.60800 to 2.58800 Ma

Collections (4 total)


Time interval Ma Country or state Original ID and collection number
Late/Upper Miocene11.608 - 5.333India (Gujarat) Clemmys watsoni (87489)
Pliocene5.333 - 2.588Pakistan Hardella thurjii, Batagur falconeri (92070)
Late/Upper Pliocene3.6 - 2.588Pakistan Hardella thurjii (224302)
Late/Upper Pliocene3.6 - 2.588India Hardella thurjii (224301)