PBDB Taxon

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Eugeneodontiformes

Chondrichthyes - Eugeneodontiformes

Taxonomy

Eugeneodontida was named by Zangerl (1981).

It was corrected as Eugeneodontiformes by Cappetta (1987), Goto and Okura (2004), Nelson (2006), Ginter et al. (2010), Long (2011) and Nelson et al. (2016).

It was assigned to Chondrichthyes by Sepkoski (2002); to Elasmobranchii by Zangerl (1981), Cappetta (1987) and Goto and Okura (2004); to Paraselachimorpha by Nelson (2006); to Euchondrocephali by Ginter et al. (2010); and to Holocephali by Long (2011) and Nelson et al. (2016).

Subtaxa

Campyloprion, Campyloprion annectans, Caseodontoidea, Edestoidea, Edistidae, Helicampodontidae, Helicoprionidae (syn. Agassizodontidae), Pseudodontichthys

Synonyms

Campyloprion annectans was named by Eastman (1902). Its type specimen is MCZ 2039, a tooth, and it is a 3D body fossil. Its type locality is Campyloprion type locality, which is in a Pennsylvanian marine horizon in Unknown.

It was considered a nomen dubium by Lebedev et al. (2023).
Campyloprion was named by Eastman (1902) [Sepkoski's age data: C Step Sepkoski's reference number: 451].

It was considered a nomen dubium by Lebedev et al. (2023).

It was assigned to Edestidae by Jordan (1963); to Eugeneodontida by Sepkoski (2002); to Agassizodontidae by Zangerl (1981) and Ginter et al. (2010); and to Helicoprionidae by Itano and Lucas (2018).

Synonymy list

Year	Name and author
1902	Campyloprion Eastman p. 151
1902	Campyloprion annectans Eastman p. 151 figs. pl. VIII, fig. 2, text-fig. 3
1963	Campyloprion Jordan p. 596
1981	Eugeneodontida Zangerl p. 74
1981	Campyloprion Zangerl p. 86
1987	Eugeneodontiformes Cappetta p. 28
2002	Campyloprion Sepkoski
2002	Eugeneodontida Sepkoski p. 162
2004	Eugeneodontiformes Goto and Okura p. 218
2006	Eugeneodontiformes Nelson p. 44
2010	Eugeneodontiformes Ginter et al. p. 117
2010	Campyloprion Ginter et al. p. 125
2011	Eugeneodontiformes Long p. 242
2016	Eugeneodontiformes Nelson et al. p. 49
2018	Campyloprion Itano and Lucas p. 408
2018	Campyloprion annectans Itano and Lucas p. 410 figs. 11A-C

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata

Rank	Name	Author
superclass	Gnathostomata
class	Chondrichthyes
subclass	Holocephali	(Bonaparte 1832)
order	Eugeneodontiformes	()
order	Eugeneodontiformes	()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Or. †Eugeneodontiformes Zangerl 1981

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Superfm. †Caseodontoidea Zangerl 1981

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G. †Campodus de Koninck 1844

†Campodus agassizianus Koninck 1844

†Campodus corrugatus Newberry and Worthen 1870

†Campodus scitulus Saint John and Worthen 1875

†Campodus virginianus Saint John and Worthen 1875

Fm. †Caseodontidae Zangerl 1981

G. †Caseodus Zangerl 1981

†Caseodus basalis Cope 1894

†Caseodus eatoni Zangerl 1981

†Caseodus varidentis Mutter and Neuman 2008

G. †Erikodus Nielsen 1952

†Erikodus groenlandicus Nielsen 1932

G. †Fadenia Nielsen 1932

†Fadenia crenulata Nielsen 1932

†Fadenia gigas Eaton 1962

†Fadenia uroclasmato Mutter and Neuman 2008

G. †Ornithoprion

†Ornithoprion hertwigi Zangerl 1966

G. †Pirodus Lebedev 2001

G. †Romerodus Zangerl 1981

†Romerodus orodontus Zangerl 1981

G. †Chiastodus Trautschold 1879

Fm. †Eugeneodontidae Zangerl 1981

G. †Bobbodus Zangerl 1981

†Bobbodus schaefferi Zangerl 1981

†Bobbodus xerxesi Hampe et al. 2013

G. †Eugeneodus Zangerl 1981

†Eugeneodus richardsoni Zangerl 1981

G. †Gilliodus Zangerl 1981

†Gilliodus orvellei Zangerl 1981

†Gilliodus peyeri Zangerl 1981

Invalid names: Orodus corrugatus Romer 1942 [nomen dubium]

G. †Tiaraju Richter 2007

Superfm. †Edestoidea Hay 1930

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Fm. †Edestidae Jaekel 1899

G. †Edestus Leidy 1855

†Edestus crenulatus Hay 1909

†Edestus giganteus Newberry 1888

†Edestus heinrichi Newberry and Worthen 1870

†Edestus karpinskii Missuna 1908

†Edestus minisculus Hay 1909

†Edestus minor Newberry 1866

†Edestus minusculus Hay 1909

†Edestus mirus Hay 1912

†Edestus newtoni Woodward 1916

†Edestus serratus Hay 1909

†Edestus triserratus Newton 1904

†Edestus vorax Leidy 1855

G. †Euctenius Traquair 1881

†Euctenius unilateralis Barkas 1874

G. †Lestrodus

G. †Prospiraxis Williams 1887

†Prospiraxis major Newberry 1885

†Prospiraxis randallii Newberry 1885

G. †Protopirata Trautschold 1888

G. †Spiraxis Newberry 1885

†Spiraxis bivalvis Ward 1885

G. †Syntomodus Obruchev 1964

†Syntomodus abbreviatus Obruchev 1964

G. †Paredestus Mutter and Neuman 2008

†Paredestus bricircum Mutter and Neuman 2008

Fm. †Helicampodontidae Itano and Lebedev 2023

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G. †Helicampodus Branson 1935

†Helicampodus egloni Obruchev 1965

†Helicampodus kokeni Branson 1935

†Helicampodus qomolangma Zhang 1976

G. †Hunanohelicoprion Liu 1994

†Hunanohelicoprion xiandongensis Liu 1994

G. †Parahelicampodus Nielsen 1952

†Parahelicampodus sparcki Nielsen 1952

G. †Sinohelicoprion Liu and Chang 1963

†Sinohelicoprion changhsingensis Liu and Chang 1963

†Sinohelicoprion macrodontus Lei 1983

Fm. †Helicoprionidae Karpinsky 1911

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G. †Helicoprion Karpinsky 1899

†Helicoprion bessonowi Karpinsky 1899

Invalid names: Helicoprion nevadensis Wheeler 1939 [synonym]

†Helicoprion davisii Woodward 1886

Invalid names: Helicoprion jingmenense Chen et al. 2007 [synonym], Helicoprion sierrensis Wheeler 1939 [synonym], Lissoprion ferrieri Hay 1907 [synonym]

†Helicoprion ergassaminon Bendix-Almgreen 1966

Invalid names: Lissoprion Hay 1907 [synonym]

G. †Karpinskiprion Lebedev and Itano 2023

†Karpinskiprion ivanovi Karpinsky 1924

Invalid names: Agassizodontidae Zangerl 1981 [synonym]

G. †Pseudodontichthys

Invalid names: Campyloprion Eastman 1902 [nomen dubium], Campyloprion annectans Eastman 1902 [nomen dubium], Edistidae Jaekel 1899 [empty]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

phosphaticsubp

Environment:

marinesubp

Locomotion:

actively mobileo

Life habit:

nektonico

Diet:

carnivoreo

Created:

2009-07-21 07:03:14

Modified:

2009-07-21 09:03:14

Source: o = order, subp = subphylum

References: Carroll 1988, Hendy et al. 2009, Wagner 2023

Age range: base of the Bashkirian to the top of the Griesbachian or 323.20000 to 250.50000 Ma

Collections (107 total)

Time interval	Ma	Country or state	Original ID and collection number
Bashkirian	323.2 - 315.2	USA (Oklahoma)	Edestus vorax (132557)
Bashkirian	323.2 - 315.2	United Kingdom	Edestus minor, Edestus triserratus (132706) Edestus newtoni (132821)
Pennsylvanian	323.2 - 298.9	USA (Unknown)	Campyloprion annectans (133378)
Pennsylvanian	323.2 - 298.9	USA (Iowa)	Campodus variabilis, Protopirata crenulata (74898)
Moscovian	315.2 - 307.0	USA (Missouri)	Fadenia gigas (133335)
Moscovian	315.2 - 307.0	USA (Indiana)	Edestus heinrichii (131614) Gilliodus orvillei, Ornithoprion hertwigi, Eugeneodus richardsoni, Caseodus eatoni (131229) Ornithoprion hertwigi (131224 131225 131226 131227 131228)
Moscovian	315.2 - 307.0	USA (Colorado)	Edestus minor (105146 132076)
Moscovian	315.2 - 307.0	Japan	Agassizodus sp. (90282)
Moscovian	315.2 - 307.0	United Kingdom	Ctenoptychius unilateralis (137306 137319)
Moscovian	315.2 - 307.0	USA (Illinois)	Edestus crenulatus, Edestus serratus (149418) Orodus basalis (132163)
Moscovian	315.2 - 307.0	USA (South Dakota)	Edestus sp., Caseodus eatoni (132341)
Moscovian	315.2 - 307.0	USA (Texas)	Edestus minor (131885 132077)
Moscovian	315.2 - 307.0	USA (indiana)	Gilliodus peyeri (132245)
Moscovian	315.2 - 307.0	USA (Iowa)	Edestus mirus (132827)
Moscovian	315.2 - 307.0	Russian Federation	Edestus karpinskii (132707) Edestus minor (149417)
Kasimovian	307.0 - 303.7	USA (New Mexico)	Campyloprion ivanovi (196772)
Kasimovian	307.0 - 303.7	USA (Iowa)	Bobbodus schaefferi (132118)
Kasimovian	307.0 - 303.7	USA (Nebraska)	Campodus variabilis (132971 133262 133265) Gilliodus peyeri (132280) Romerodus orodontus (132113 132117)
Kasimovian	307.0 - 303.7	USA (Kansas)	Caseodus eatoni, Edestus heinrichi (132333)
Late/Upper Pennsylvanian	307.0 - 298.9	United Kingdom (Bollington)	Edestus sp. (222586)
Gzhelian	303.7 - 298.9	Russian Federation	Campodus sp. (51474) Helicoprion ivanovi, Campyloprion annectans (173816)
Gzhelian	303.7 - 298.9	USA (Texas)	Campyloprion sp. (196773 196774)
Gzhelian	303.7 - 298.9	Russian Federation (Volgograd)	Karpinskiprion ivanovi (232358)
Gzhelian	303.7 - 298.9	USA (Nebraska)	Bobbodus schaefferi (132177)
Asselian	298.9 - 295.5	USA (Kansas)	Agassizodus sp., Agassizodus variabilis (132372) Bobbodus schaefferi (118270)
Asselian	298.9 - 295.5	Bolivia	Parahelicoprion mariosuarezi (117946)
Asselian - Sakmarian	298.9 - 290.1	USA (Nevada)	Helicoprion sp. (130653)
Cisuralian	298.9 - 273.01	USA (Nevada)	Edestus lecontei (196771)
Cisuralian	298.9 - 273.01	USA (California)	Helicoprion sierrensis (118113)
Permian	298.9 - 251.902	USA (Nevada)	Helicoprion nevadensis (118114)
Sakmarian	293.52 - 290.1	USA (Nevada)	Helicoprion sp. (130655)
Sakmarian	293.52 - 290.1	USA (New Mexico)	Agassizodus sp. (39154)
Artinskian	290.1 - 279.3	Russian Federation	Helicoprion bessonowi (149416)
Artinskian	290.1 - 279.3	USA (Texas)	Helicoprion ferrieri (105721) Orodus corrugatus (132655)
Artinskian	290.1 - 279.3	Kazakhstan	Helicoprion bessonowi (119029)
Artinskian - Kungurian	290.1 - 272.3	USA (Texas)	Orodus corrugatus (85301)
Kungurian	279.3 - 272.3	China (Hunan)	Hunanohelicoprion xiandongensis (50995)
Kungurian	279.3 - 272.3	Australia (Western Australia)	Helicoprion davisii (118112)
Kungurian	279.3 - 272.3	Mexico (Puebla)	Helicoprion sp. (90679)
Kungurian	279.3 - 272.3	Brazil (Maranhao)	Edestidae indet. (192146)
Kungurian	279.3 - 272.3	USA (Texas)	Helicoprion sp. (90276)
Kungurian	279.3 - 272.3	China (Hubei)	Helicoprion jingmenense (120307)
Guadalupian	273.01 - 259.51	China (Guizhou)	Hunanohelicoprion xiandongensis (118031)
Guadalupian	273.01 - 259.51	Canada (British Columbia)	Helicoprion sp. (149354)
Roadian	272.3 - 268.8	Canada (Alberta)	Helicoprion sp. (112844)
Roadian	272.3 - 268.8	USA (Montana)	Helicoprion sp. (11280)
Roadian	272.3 - 268.8	Canada (Nunavut)	Helicoprion sp. (90234 152126)
Roadian	272.3 - 268.8	USA (Texas)	Helicoprion sp. (152125)
Roadian	272.3 - 268.8	USA (Idaho)	Helicoprion davisii (152127 152129 152130) Helicoprion davisii, Helicoprion ergassaminon (152128) Helicoprion ferrieri (140078) Helicoprion sp. (11285 152131) Helicoprion sp., Helicoprion ergassaminon (140145) Lissoprion ferrieri (132826)
Roadian	272.3 - 268.8	Japan	Helicoprion bessonowi (133437)
Roadian	272.3 - 268.8	USA (Wyoming)	Campodus sp. (149346) Helicoprion sp. (11229) Lissoprion ferrieri (108931)
Roadian	272.3 - 268.8	USA (Utah)	Helicoprion sp. (11271)
Wordian	266.9 - 264.28	Oman	Eugeneodontiformes indet. (140671 140673)
Capitanian	264.28 - 259.51	Japan	Helicoprion sp. (149339)
Wuchiapingian	259.9 - 254.17	Iran	Eugeneodontidae indet., Bobbodus xerxesi (137398)
Wuchiapingian	259.9 - 254.17	Pakistan	Helicoprion sp. (149440)
Wuchiapingian	259.9 - 254.17	USA (Wyoming)	Campodus variabilis, Edestus sp. (90235)
Wuchiapingian	259.9 - 254.17	Greenland	Agassizodus groenlandicus, Fadenia crenulata (122142) Fadenia crenulata, Sarcoprion edax, Erikodus groenlandicus (136371 136372) Parahelicampodus sparcki (136373)
Wuchiapingian - Changhsingian	259.9 - 252.17	Japan	Helicampodus sp. (133502)
Changhsingian	254.17 - 252.17	Azerbaijan	Helicampodus egloni (97207)
Changhsingian	254.17 - 252.17	Pakistan	Helicampodus kokeni (27264)
Changhsingian	254.17 - 252.17	China (Hunan)	Sinohelicoprion macrodontus (123423)
Changhsingian	254.17 - 252.17	China (Zhejiang)	Sinohelicoprion changhsingensis (121648 149441 149442)
Griesbachian	251.902 - 250.5	India (Kashmir)	Helicampodus sp. (48914)
Griesbachian	251.902 - 250.5	China (Xizang)	Sinohelicoprion qomolangma (148625)
Smithian	251.3 - 247.2	Canada (British Columbia)	Eugeneodontida indet., Caseodontidae indet., Fadenia uroclasmato (105083) Eugeneodontida indet., Fadenia sp., Caseodontidae indet., Caseodus varidentis, Fadenia uroclasmato, Paredestus bricircum (119095) Fadenia sp. (122091)
Spathian	251.3 - 247.2	USA (Utah)	Caseodontidae indet. (63143)