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Bothremydini
Taxonomy
Bothremydini was named by Gaffney et al. (2006). It was considered monophyletic by Pérez-García (2017).
It was assigned to Bothremydodda by Gaffney et al. (2006), Pérez-García et al. (2013), Pérez-García (2017), Pérez-García and Ortega (2018), Pérez-Garcia (2020) and Pérez-García et al. (2021); to Bothremydidae by Pérez-García (2020) and López-Conde et al. (2021); to Bothremydinae by Gaffney et al. (2009) and Noto et al. (2022); and to Bothremydidae by Joyce et al. (2021), Cadena et al. (2024) and Antunes and Pérez-García (2024).
It was assigned to Bothremydodda by Gaffney et al. (2006), Pérez-García et al. (2013), Pérez-García (2017), Pérez-García and Ortega (2018), Pérez-Garcia (2020) and Pérez-García et al. (2021); to Bothremydidae by Pérez-García (2020) and López-Conde et al. (2021); to Bothremydinae by Gaffney et al. (2009) and Noto et al. (2022); and to Bothremydidae by Joyce et al. (2021), Cadena et al. (2024) and Antunes and Pérez-García (2024).
Subtaxa
Synonymy list
Year | Name and author |
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2006 | Bothremydini Gaffney et al. p. 60 |
2009 | Bothremydini Gaffney et al. p. 2 |
2013 | Bothremydini Pérez-García et al. |
2017 | Bothremydini Pérez-García p. 2 |
2018 | Bothremydini Pérez-García and Ortega |
2020 | Bothremydini Pérez-Garcia |
2020 | Bothremydini Pérez-García |
2021 | Bothremydini Joyce et al. |
2021 | Bothremydini López-Conde et al. |
2021 | Bothremydini Pérez-García et al. |
2022 | Bothremydini Noto et al. p. 3 |
2024 | Bothremydini Antunes and Pérez-García |
2024 | Bothremydini Cadena et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Tr. †Bothremydini Gaffney et al. 2006
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Subtr. †Bothremydina Gaffney et al. 2006
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G. †Algorachelus Pérez-García 2017
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†Algorachelus peregrina Pérez-García 2017
†Algorachelus tibert Joyce et al. 2016
G. †Bothremys Leidy 1865
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†Bothremys arabicus Zalmout et al. 2005
†Bothremys cooki Leidy 1865
†Bothremys kellyi Gaffney et al. 2006
†Bothremys maghrebiana Gaffney et al. 2006
Invalid names: Karkaemys Zalmout et al. 2005 [synonym]
G. †Zolhafah Lapparent de Broin and Werner 1998
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†Zolhafah bella Lapparent de Broin and Werner 1998
Subtr. †Foxemydina Gaffney et al. 2006
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G. †Foxemys Tong et al. 1998
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†Foxemys mechinorum Tong et al. 1998
†Foxemys trabanti Rabi et al. 2012
G. †Iberoccitanemys Pérez-García et al. 2012
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†Iberoccitanemys atlanticum Lapparent de Broin and Murelaga 1996
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Invalid names: Elochelys convenarum Laurent et al. 2002 [synonym]
G. †Polysternon Portis 1882
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Invalid names: Polysternon isonae Marmi et al. 2012 [nomen dubium]
Diagnosis
Reference | Diagnosis | |
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O. A. López-Conde et al. 2021 | It is recognized as a member of Bothremydidae based on rectangular neural I; contact of the axillary buttress with the main body of peripheral III; contact of the inguinal buttress with costal V; absence of a cervical scute; and costal I twice as long as costal II. |
Measurements
No measurements are available
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Source: subo = suborder, c = class, subp = subphylum | |||||
References: Uetz 2005, Carroll 1988, Hendy et al. 2009 |
Age range: base of the Middle Cenomanian to the top of the Late/Upper Paleocene or 99.60000 to 55.80000 Ma
Collections (67 total)
Time interval | Ma | Country or state | Original ID and collection number |
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Cenomanian | Portugal (Lisbon) | Algorachelus peregrina (234569) | |
Cenomanian | Madagascar (Atsimo-Andrefana) | Akoranemys madagasika (214154) | |
Cenomanian | Spain (Soria) | Algorachelus sp. (215933) | |
Early/Lower Cenomanian | Palestinian Territory | Podocnemis parva, Podocnemis judaea (11804) | |
Middle Cenomanian | Portugal (Leiria) | Algorachelus peregrinus (186171) | |
Middle Cenomanian - Late/Upper Cenomanian | Spain (Castilla-La Mancha) | Eupleurodira indet. (132494) | |
Middle Cenomanian - Late/Upper Cenomanian | USA (Utah) | Paiutemys tibert (181692) | |
Santonian - Coniacian | USA (Kansas) | Chedighaii barberi (129833) | |
Santonian | USA (Alabama) | Podocnemis alabamae (81928) | |
Santonian | Hungary (Veszprém) | Testudines indet. (48722) | |
Santonian | Jordan | Karkaemys arabicus (83312) | |
Campanian - Maastrichtian | Spain (Vasco-Cantabrica) | Polysternon sp. (208188 208189) | |
Early/Lower Campanian | France | Polysternon provinciale (83309) | |
Early/Lower Campanian | USA (New Jersey) | Bothremys cooki, Chedighaii barberi (88610) | |
Early/Lower Campanian | France (Languedoc-Roussillon) | Polysternon provinciale (47782) | |
Early/Lower Campanian | France (Provence-Alpes-Côte d'Azur) | Elochelys major (114968) Pleurosternon provinciale (114967) Polysternon sp. (220117) | |
Early/Lower Campanian - Middle Campanian | USA (North Carolina) | Bothremys cooki (129825) Bothremys sp., Bothremys cooki, Chedighaii barberi (57918) Bothremys sp., Chedighaii barberi (128412) | |
Early/Lower Campanian - Middle Campanian | USA (Alabama) | Chedighaii barberi (129836) | |
Late/Upper Campanian | Spain (Castilla y Leon) | Iberoccitanemys convenarum (48861 193947) Polysternon atlanticum (28406 37199) | |
Late/Upper Campanian | France (Provence-Alpes-Côte d'Azur) | Foxemys mechinorum (89990) Podocnemidae indet. (28454) Testudinata indet. (220134) | |
Late/Upper Campanian | Mexico (Coahuila) | Bothremydina indet. (35360) | |
Middle Campanian | France (Provence-Alpes-Côte d'Azur) | Polysternon sp. (48849) | |
Middle Campanian | Egypt (New Valley) | Khargachelys caironensis (229326) | |
Middle Campanian | USA (South Carolina) | Bothremys sp. (180790) | |
Middle Campanian - Late/Upper Campanian | France (Languedoc-Roussillon) | Polysternon sp. (74660) | |
Late/Upper Campanian - Early/Lower Maastrichtian | Spain (Valencia) | Polysternon sp. (53918) | |
Late/Upper Campanian - Early/Lower Maastrichtian | Mexico (Coahuila) | Palauchelys montellanoi (215838) | |
Late/Upper Campanian - Early/Lower Maastrichtian | Spain (Castilla-La Mancha) | Foxemydina indet., Bothremydidae indet. (91781) Iberoccitanemys convenarum (128273) | |
Late/Upper Campanian - Early/Lower Maastrichtian | France (Languedoc-Roussillon) | Bothremydidae indet. (48870) Foxemys mechinorum (59887) Pelomedusidae indet. (59686) Polysternon provinciale (88584) | |
Maastrichtian | France (Provence-Alpes-Côte d'Azur) | Polysternon provinciale (51692) | |
Maastrichtian | Egypt | Zolhafah bella (83310) | |
Maastrichtian | USA (New Jersey) | Bothremys cookii (81999) | |
Early/Lower Maastrichtian | Spain (Cataluña) | Polysternon isonae (132488) | |
Early/Lower Maastrichtian | France (Provence-Alpes-Côte d'Azur) | Polysternon sp. (47893) | |
Early/Lower Maastrichtian | France (Languedoc-Roussillon) | Testudines indet. (48850) | |
Late/Upper Maastrichtian | France (Haute-Garonne) | Foxemys sp. (148954) Foxemys sp., Elochelys sp. (67196) | |
Late/Upper Maastrichtian | Spain (Aragón) | Polysternon sp. (45437 45438 45439 45440) | |
Late/Upper Maastrichtian | Spain (Burgos) | Polysternon atlanticum (176782) | |
Late/Upper Maastrichtian - Early/Lower Paleocene | USA (New Jersey) | Bothremys sp. (26819) | |
Danian | Morocco (Khouribga) | Bothremys maghrebiana (129831) | |
Danian | Morocco | Araiochelys hirayamai, Bothremys maghrebiana (83311) | |
Middle Paleocene - Late/Upper Paleocene | Colombia (Guajira) | Puentemys mushaisaensis (106271 132012 132013 132014) | |
Selandian | Morocco | Bothremys kellyi (68176) | |
Thanetian | Colombia (Socha) | Puentemys mushaisaensis (234570) | |
Clarkforkian | USA (South Carolina) | Bothremys sp. (15074) | |
Ypresian | France (Aude) | Foxemydina indet., Tartaruscola teodorii (78236) |