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Gwyneddichnium majore

Osteichthyes - Gwyneddichniidae

Taxonomy
Gwyneddichnium majore was named by Bock (1952). It is considered to be a form taxon. Its type specimen is ANSP 15212-15213, a footprint, and it is a trace fossil. Its type locality is Gwynedd Tunnel (track layers), which is in a Norian floodplain claystone in the Lockatong Formation of Pennsylvania. It is the type species of Gwyneddichnium.

Sister species lacking formal opinion data
G. minor

Synonymy list
YearName and author
1952Gwyneddichnium majore Bock p. 417 figs. Pl. 49, 50:1
1958Gwyneddichnium majore Kuhn p. 20
2007Gwyneddichnium majore Hunt and Lucas p. 216
2021Gwyneddichnium majore Klein and Lucas p. 73

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
classOsteichthyes
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
RankNameAuthor
—Amniota
—Sauropsida
classReptilia
subclassEureptilia()
—Romeriida
—Diapsida()
—Archosauromorpha(Huene 1946)
—Crocopoda
—ArchosauriformesGauthier 1986
—Eucrocopoda
—Archosauria()
—Pseudosuchia(Zittel 1890)
familyGwyneddichniidae()
genusGwyneddichnium
speciesmajore

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Gwyneddichnium majore Bock 1952
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Diagnosis
ReferenceDiagnosis
H. Klein and S. G. Lucas 2021Footprints of a quadruped in which the manus and pes are pentadactyl and mesaxonic, the pes is substantially larger than the manus, digits are thin, long, straight to slightly curved, have nodular phalangeal pad impressions and terminate in claws. Differs from the most similar, lacertoid ichnogenus Rhynchosauroides in the digit proportions, with digits III > II ≥ IV > I in Gwyneddichnium, whereas in Rhynchosauroides the proportions are IV > III > II > I. Also, the digits in Rhynchosauroides are thicker and display a greater curvature, whereas the digits of Gwyneddichnium are nearly straight or only slightly curved. Digit V in Rhynchosauroides is strongly recurved laterally, whereas in Gwyneddichnium digit V is often nearly parallel to digit IV.
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: terrestrialsubc
Locomotion: actively mobilec
Diet: carnivoresubo
Created: 2009-01-03 20:38:45
Modified: 2009-01-03 22:38:45
Source: subo = suborder, subc = subclass, c = class, subp = subphylum
References: Benton 1983, Hendy et al. 2009, Kiessling 2004, Carroll 1988

Age range: Norian or 227.30000 to 205.70000 Ma

Collections (3 total)


Time interval Ma Country or state Original ID and collection number
Norian227.3 - 205.7USA (Pennsylvania) Gwyneddichnium majore (type locality: 68706)
Norian227.3 - 205.7USA (Virginia) Gwyneddichnium majore (105101)
Norian - Rhaetian227.3 - 201.4USA (New Jersey) Gwyneddichnium majore (92107)