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Cantharinae
Taxonomy
Cantharinae was named by Imhoff (1856). It is extant. Its type is Cantharis.
It was assigned to Cantharidae by Lawrence and Newton (1995), Kuśka and Kania (2010), Bouchard et al. (2011), Kazantsev (2013), Fanti and Ellenberger (2016), Poinar and Fanti (2016), Fanti (2017), Fanti (2017), Fanti (2017), Fanti and Damgaard (2018), Fanti and Ellenberger (2018), Fanti and Pankowski (2018), Peris and Fanti (2018), Fanti and Kupryjanowicz (2018), Fanti et al. (2018), Fanti and Poschmann (2019), Fanti and Damgaard (2019), Poinar and Fanti (2019), Ellenberger and Fanti (2019), Bukejs et al. (2019), Kupryjanowicz and Fanti (2019), Parisi and Fanti (2020), Fanti and Damgaard (2020), Fanti and Pankowski (2020), Kazantsev (2020), Kazantsev and Perkovsky (2020), Fanti (2021), Fanti (2021), Fanti and Müller (2022), Wentzel et al. (2022), Bramanti and Fanti (2022), Cai et al. (2022), Fanti and Vitali (2023), Fanti and Pankowski (2023), Qu et al. (2023), Pankowski and Fanti (2023), Fanti (2025), Fanti (2025), Kazantsev and Perkovsky (2025).
It was assigned to Cantharidae by Lawrence and Newton (1995), Kuśka and Kania (2010), Bouchard et al. (2011), Kazantsev (2013), Fanti and Ellenberger (2016), Poinar and Fanti (2016), Fanti (2017), Fanti (2017), Fanti (2017), Fanti and Damgaard (2018), Fanti and Ellenberger (2018), Fanti and Pankowski (2018), Peris and Fanti (2018), Fanti and Kupryjanowicz (2018), Fanti et al. (2018), Fanti and Poschmann (2019), Fanti and Damgaard (2019), Poinar and Fanti (2019), Ellenberger and Fanti (2019), Bukejs et al. (2019), Kupryjanowicz and Fanti (2019), Parisi and Fanti (2020), Fanti and Damgaard (2020), Fanti and Pankowski (2020), Kazantsev (2020), Kazantsev and Perkovsky (2020), Fanti (2021), Fanti (2021), Fanti and Müller (2022), Wentzel et al. (2022), Bramanti and Fanti (2022), Cai et al. (2022), Fanti and Vitali (2023), Fanti and Pankowski (2023), Qu et al. (2023), Pankowski and Fanti (2023), Fanti (2025), Fanti (2025), Kazantsev and Perkovsky (2025).
Subtaxa
Synonymy list
| Year | Name and author |
|---|---|
| 1856 | Cantharinae Imhoff |
| 1995 | Cantharinae Lawrence and Newton, Jr. |
| 2010 | Cantharinae Kuśka and Kania p. 50 |
| 2011 | Cantharinae Bouchard et al. p. 330 |
| 2013 | Cantharinae Kazantsev p. 290 |
| 2016 | Cantharinae Fanti and Ellenberger p. 166 |
| 2016 | Cantharinae Poinar and Fanti p. 2 |
| 2017 | Cantharinae Fanti p. 2 |
| 2018 | Cantharinae Fanti and Damgaard p. 2 |
| 2018 | Cantharinae Fanti and Ellenberger p. 202 |
| 2018 | Cantharinae Fanti and Kupryjanowicz p. 150 |
| 2018 | Cantharinae Fanti and Pankowski p. 514 |
| 2018 | Cantharinae Peris and Fanti p. 264 |
| 2019 | Cantharinae Bukejs et al. p. 2 |
| 2019 | Cantharinae Ellenberger and Fanti p. 595 |
| 2019 | Cantharinae Fanti and Damgaard p. 102 |
| 2019 | Cantharinae Fanti and Poschmann p. 364 |
| 2019 | Cantharinae Kupryjanowicz and Fanti p. 2 |
| 2019 | Cantharinae Poinar and Fanti p. 99 |
| 2020 | Cantharinae Fanti and Damgaard p. 126 |
| 2020 | Cantharinae Fanti and Pankowski p. 402 |
| 2020 | Cantharinae Kazantsev p. 260 |
| 2020 | Cantharinae Kazantsev and Perkovsky p. 396 |
| 2020 | Cantharinae Parisi and Fanti p. 547 |
| 2021 | Cantharinae Fanti p. 2 |
| 2022 | Cantharinae Bramanti and Fanti p. 202 |
| 2022 | Cantharinae Cai et al. |
| 2022 | Cantharinae Fanti and Müller p. 332 |
| 2022 | Cantharinae Wentzel et al. p. 314 |
| 2023 | Cantharinae Fanti and Pankowski p. 294 |
| 2023 | Cantharinae Fanti and Vitali p. 234 |
| 2023 | Cantharinae Pankowski and Fanti p. 301 |
| 2023 | Cantharinae Qu et al. p. 4 |
| 2025 | Cantharinae Fanti p. 2 |
| 2025 | Cantharinae Kazantsev and Perkovsky p. 55 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Subfm. Cantharinae Imhoff 1856
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G. †Brevipterus Yang et al. 2022
+
†Brevipterus acutiapicis Yang et al. 2022
†Brevipterus megacephalus Yang et al. 2022
†Brevipterus obtusiapicis Yang et al. 2022
Tr. †Cacomorphocerini Fanti and Kupryjanowicz 2018
+
G. †Cacomorphocerus Schaufuss 1891
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†Cacomorphocerus bentifabrici Fanti and Damgaard 2018
†Cacomorphocerus cerambyx Schaufuss 1891
†Cacomorphocerus coleae Fanti and Pankowski 2019
†Cacomorphocerus endecacerus Poinar and Fanti 2019
†Cacomorphocerus eocenicus Bukejs et al. 2019
†Cacomorphocerus incurvus Fanti and Vitali 2023
†Cacomorphocerus jantaricus Kuśka and Kania 2010
†Cacomorphocerus madseni Fanti and Damgaard 2018
†Cacomorphocerus marki Fanti and Pankowski 2020
†Cacomorphocerus meridionalis Kazantsev and Perkovsky 2020
†Cacomorphocerus obstinatus Parisi and Fanti 2020
†Cacomorphocerus simkei Fanti 2024
†Cacomorphocerus thomasiwentzeli Wentzel et al. 2022
†Cacomorphocerus wiszniewskii Fanti and Kupryjanowicz 2018
Invalid names: Hoffeinsensia Kuśka and Kania 2010 [synonym]
G. †Electronycha Kazantsev 2013
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†Electronycha arturmichalskii Fanti 2025
†Electronycha prussica Kazantsev 2013
Tr. Cantharini Imhoff 1856
+
G. †Burmomiles Fanti et al. 2018
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†Burmomiles blixenae Fanti and Damgaard 2019
†Burmomiles brevicollis Yang et al. 2024
†Burmomiles laticollis Yang et al. 2021
†Burmomiles parisii Fanti and Müller 2022
†Burmomiles volpei Fanti and Müller 2022
†Burmomiles willerslevorum Fanti et al. 2018
G. Cantharis Linnaeus 1758
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Subg. Cantharis (Cantharis) Linnaeus 1758
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†Cantharis (Cantharis) attavina Heer 1847
†Cantharis (Cantharis) borki Fanti and Damgaard 2019
†Cantharis (Cantharis) bradburyi Fanti and Walker 2019 [solider beetle]
†Cantharis (Cantharis) brodiei von Heyden and von Heyden 1866
†Cantharis (Cantharis) caduca von Heyden and von Heyden 1866
†Cantharis (Cantharis) carbonaria von Heyden and von Heyden 1866
†Cantharis (Cantharis) crisantha Fanti and Pankowski 2020
†Cantharis (Cantharis) doernerorum Fanti and Poschmann 2019
†Cantharis (Cantharis) dougi Kupryjanowicz and Fanti 2019
†Cantharis (Cantharis) exauctarata von Heyden and von Heyden 1866
Cantharis (Cantharis) figurata Mannerheim 1843
†Cantharis (Cantharis) fragilis Heer 1847
†Cantharis (Cantharis) hanswerneri Kazantsev 2018
†Cantharis (Cantharis) hoffeinsorum Kazantsev 2018
†Cantharis (Cantharis) kviumi Fanti and Damgaard 2020
†Cantharis (Cantharis) lidiae Fanti and Walker 2019 [solider beetle]
†Cantharis (Cantharis) macilenta Heer 1864
†Cantharis (Cantharis) michaeli Fanti and Pankowski 2023
Cantharis (Cantharis) paludosa Fallen 1807
†Cantharis (Cantharis) raeorum Fanti and Pankowski 2020
†Cantharis (Cantharis) rottensis Fanti and Walker 2019 [solider beetle]
Cantharis (Cantharis) rufa Linnaeus 1758
†Cantharis (Cantharis) sesermanae Manci et al. 2025
†Cantharis (Cantharis) sucinonigra Kuśka 1992
†Cantharis (Cantharis) zabolica Fanti and Poschmann 2019
Subg. Cantharis (Cyrtomoptila) Motschulsky 1860
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†Cantharis (Cyrtomoptila) mikkelsenorum Fanti and Damgaard 2018
†Cantharis (Cyrtomoptila) sucinokotejai Kuśka 1996
Invalid names: Cantharis (Absidiella) Wittmer 1972 [synonym]
Invalid names: Telephorus Schaeffer 1766 [synonym]
G. †Elektrokleinia Ellenberger and Fanti 2019
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†Elektrokleinia bilineatimaculata Yang et al. 2021
†Elektrokleinia oblongoculus Yang et al. 2021
†Elektrokleinia panna Hsiao et al. 2021
†Elektrokleinia picta Ellenberger and Fanti 2019
†Elektrokleinia steffenseni Fanti and Damgaard 2020
†Elektrokleinia thiloi Fanti and Müller 2022
†Elektrokleinia zahradniki Fanti and Müller 2022
Invalid names: Palaeocantharis Hsiao et al. 2021 [synonym]
G. †Hukawngichthyurus Fanti and Ellenberger 2018
+
†Hukawngichthyurus barsevskisi Fanti and Müller 2022
†Hukawngichthyurus ditaddeoi Fanti and Müller 2022
†Hukawngichthyurus kyawkhaingwini Fanti and Ellenberger 2018
†Hukawngichthyurus maha Hsiao et al. 2021
G. Lycocerus Gorham 1889
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†Lycocerus albovacca Kazantsev and Perkovsky 2024
†Lycocerus christelae Kazantsev 2018
†Lycocerus crassepunctatus Kazantsev 2021
†Lycocerus dentantennatus Kazantsev 2018
†Lycocerus elzbietae Kazantsev 2020
†Lycocerus guttula Zhang 1989
†Lycocerus jesperibuchi Fanti and Damgaard 2019
†Lycocerus jonasi Kazantsev 2020
†Lycocerus xenium Fanti 2020
G. Podistra Motschulsky 1839
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Subg. Podistra (Absidia) Mulsant 1862
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†Podistra (Absidia) jirii Fanti 2020
†Podistra (Absidia) kloevedali Fanti and Damgaard 2018
†Podistra (Absidia) mattheseni Fanti and Damgaard 2020
†Podistra (Absidia) quies Fanti and Walker 2019 [soldier beetle]
†Podistra madelineae Pankowski and Fanti 2023
G. †Poinarelektronmiles Fanti and Damgaard 2020
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†Poinarelektronmiles cuaroni Bramanti and Fanti 2022
†Poinarelektronmiles ellenbergeri Fanti and Damgaard 2020
G. Rhagonycha Eschscholtz 1830
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Subg. Rhagonycha (Rhagonycha) Eschscholtz 1830
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†Rhagonycha (Rhagonycha) carolynae Fanti and Walker 2019 [soldier beetle]
†Rhagonycha (Rhagonycha) germari Heer 1847
†Rhagonycha (Rhagonycha) hesperus Wickham 1914
†Rhagonycha (Rhagonycha) kryshtofovichi Yablokov-Khnzorian 1960
†Rhagonycha (Rhagonycha) maryae Fanti and Pankowski 2018
†Rhagonycha (Rhagonycha) micans Piton 1939
†Rhagonycha (Rhagonycha) nielsenae Fanti and Damgaard 2018
†Rhagonycha (Rhagonycha) sucinobaltica Poinar and Fanti 2016
†Rhagonycha (Rhagonycha) tertiaria Heer 1847
†Rhagonycha (Rhagonycha) ultramundana Fanti and Walker 2019 [solider beetle]
†Rhagonycha acarigera Kazantsev 2020
G. †Sanaungulus Fanti et al. 2018
+
†Sanaungulus angustipennis Yang et al. 2024
†Sanaungulus christensenae Fanti and Damgaard 2019
†Sanaungulus curtipennis Fanti et al. 2018
†Sanaungulus dentatus Yang et al. 2024
†Sanaungulus dominikiweissbachi Fanti and Müller 2022
†Sanaungulus dunlopi Fanti and Müller 2022
†Sanaungulus electrum Fanti and Müller 2022
†Sanaungulus elongaticollis Yang et al. 2024
†Sanaungulus emarginaticollis Yang et al. 2022
†Sanaungulus fabriciusi Fanti and Damgaard 2019
†Sanaungulus franziskaeweissbachae Fanti and Müller 2022
†Sanaungulus ghitaenoerbyae Fanti et al. 2018
†Sanaungulus havai Fanti and Müller 2022
†Sanaungulus imparitibius Yang et al. 2022
†Sanaungulus kachinensis Fanti and Müller 2022
†Sanaungulus kirstenaeweissbachae Fanti and Müller 2022
†Sanaungulus laticoxa Yang et al. 2022
†Sanaungulus leniae Fanti and Müller 2022
†Sanaungulus lethi Fanti and Damgaard 2020
†Sanaungulus longicornis Yang et al. 2024
†Sanaungulus marginalis Yang et al. 2024
†Sanaungulus morellii Fanti and Damgaard 2020
†Sanaungulus multiramus Yang et al. 2022
†Sanaungulus myanmaricus Fanti and Müller 2022
†Sanaungulus myitkyinaensis Fanti and Müller 2022
†Sanaungulus nalae Fanti and Müller 2022
†Sanaungulus nilsi Fanti and Müller 2022
†Sanaungulus perkovskyi Fanti and Müller 2022
†Sanaungulus peteriruedeli Fanti and Müller 2022
†Sanaungulus recticollis Yang et al. 2024
†Sanaungulus rosenzweigi Fanti and Damgaard 2020
†Sanaungulus ruficollis Yang et al. 2022
†Sanaungulus ruicheni Hsiao and Huang 2018
†Sanaungulus strungei Fanti and Damgaard 2019
†Sanaungulus temporiscapsula Fanti and Müller 2022
†Sanaungulus troelsikloevedali Fanti and Damgaard 2019
†Sanaungulus ypogaeum Fanti and Müller 2022
G. †Sucinorhagonycha Kuśka 1996
+
†Sucinorhagonycha carsteni Fanti 2023
†Sucinorhagonycha groehni Kazantsev 2020
†Sucinorhagonycha kulickae Kuśka 1996
†Sucinorhagonycha maryae Pankowski and Fanti 2023
†Sucinorhagonycha pugachensis Kazantsev and Perkovsky 2025
†Sucinorhagonycha samsockorum Fanti and Pankowski 2018
G. Themus Motschulsky 1858
+
Subg. Themus (Haplothemus) Wittmer 1973
+
†Themus (Haplothemus) bennyianderseni Fanti and Damgaard 2018
†Themus (Haplothemus) pristinus Kazantsev 2013
Subg. Themus (Themus) Motschulsky 1858
+
†Themus (Themus) capacis Zhang 1989
†Themus (Themus) thermophilus Zhang 1989
†Themus (Themus) trapezialis Zhang et al. 1994
Invalid names: Curticantharis Zhang 1989 [synonym]
G. †Vitalfranzius Fanti and Müller 2022
+
†Vitalfranzius burmiticus Fanti and Müller 2022
†Vitalfranzius cretaceus Fanti and Müller 2022
Invalid names: Sanaungulus undecimus Yang et al. 2024 [nomen dubium]
Tr. Podabrini Gistel 1856
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G. Malthacus Kirby 1837
G. Podabrus Leconte 1850
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Podabrus alpinus Paykull 1798
†Podabrus cupesoides Wickham 1917
†Podabrus florissantensis Wickham 1914
†Podabrus fragmentatus Wickham 1914
†Podabrus santaritensis Cockerell 1936
†Podabrus wheeleri Wickham 1909
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: c = class, p = phylum | |||||
| References: Bush and Bambach 2015, Kiessling 2004 | |||||
Age range
Maximum range based only on fossils: base of the Early/Lower Albian to the top of the Late/Upper Pleistocene or 113.20000 to 0.01170 Ma
Minimum age of oldest fossil (stem group age): 110.1 Ma
Minimum age of oldest fossil (stem group age): 110.1 Ma
Collections (63 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Early/Lower Albian | Spain | Molliberus albae (109438) | |
| Early/Lower Cenomanian | Myanmar | Burmomiles blixenae, Sanaungulus christensenae, Sanaungulus fabriciusi, Sanaungulus strungei, Sanaungulus troelsikloevedali, Elektrokleinia steffenseni, Poinarelektronmiles ellenbergeri, Sanaungulus lethi, Sanaungulus morellii, Sanaungulus rosenzweigi (178122) Burmomiles laticollis, Burmomiles bilineatimaculatus, Sanaungulus multiramus, Sanaungulus imparitibius, Sanaungulus ruficollis, Brevipterus obtusiapicis, Brevipterus acutiapicis, Brevipterus megacephalus, Burmomiles brevicollis, Burmomiles angustipennis, Burmomiles recticollis, Burmomiles dentatus, Sanaungulus undecimus, Sanaungulus marginalis, Sanaungulus longicornis, Sanaungulus elongaticollis (222554) Burmomiles oblongoculus, Sanaungulus emarginaticollis (178121) Burmomiles parisii, Sanaungulus electrum, Sanaungulus havai, Sanaungulus kirstenaeweissbachae, Sanaungulus leniae, Sanaungulus myanmaricus, Sanaungulus myitkyinaensis, Sanaungulus nalae, Sanaungulus peteriruedeli (228869) Burmomiles volpei, Elektrokleinia thiloi, Elektrokleinia zahradniki, Hukawngichthyurus barsevskisi, Hukawngichthyurus ditaddeoi, Sanaungulus dominikiweissbachi, Sanaungulus dunlopi, Sanaungulus franziskaeweissbachae, Sanaungulus kachinensis, Sanaungulus nilsi, Sanaungulus perkovskyi, Sanaungulus temporiscapsula, Sanaungulus ypogaeum, Vitalfranzius burmiticus, Vitalfranzius cretaceus (216411) Burmomiles willerslevorum, Sanaungulus ghitaenoerbyae (192152) Elektrokleinia picta (172900) Hukawngichthyurus maha (224861) Ornatomalthinus ruicheni (196206) Palaeocantharis panna, Sanaungulus laticoxa (165561) Poinarelektronmiles cuaroni (228604) Sanaungulus curtipennis, Hukawngichthyurus kyawkhaingwini (123053) | |
| Late/Upper Paleocene | Argentina (Jujuy) | Podabrus santaritensis (151868) | |
| Lutetian | Russian Federation | Cnathrion sakhalinense (107340) | |
| Late/Upper Eocene | USA (Colorado) | Telephorus humatus (122093) | |
| Priabonian | Russian Federation (Kaliningrad) | Cacomorphocerus cerambyx (123771) Cacomorphocerus eocenicus (180657) Cacomorphocerus obstinatus (202735) Cacomorphocerus sp., Cacomorphocerus bentifabrici, Cacomorphocerus madseni, Eridanula susannaebierae, Noergaardia dinae, Cantharis mikkelsenorum, Juratelacrima ballingi, Palmnickeneoceras ejersboi, Podistra kloevedali, Rhagonycha nielsenae, Themus bennyianderseni, Cantharis borki, Lycocerus jesperibuchi, Cantharis kviumi, Podistra mattheseni (172936) Cacomorphocerus thomasiwentzeli (228858) Cantharis crisantha, Cantharis raeorum, Arturmiles pankowskiarum (124205) Electronycha prussica (123764) Rhagonycha maryae, Sucinorhagonycha samsockorum, Cacomorphocerus coleae, Cacomorphocerus marki, Sucinorhagonycha maryae, Podistra madelineae (147652) Rhagonycha sucinobaltica, Cacomorphocerus endecacerus (125139) Sucinorhagonycha groehni, Sucinorhagonycha carsteni, Groehnionycha yantarnyca, Podistra carstengroehni, Cacomorphocerus simkei (110735) | |
| Priabonian | United Kingdom (England) | Themus polyaki (123960) | |
| Priabonian | Lithuania | Malchinus kryshtofovichi (113470) Podistra guthriei (154362) | |
| Priabonian | Russian Federation | Hoffeinsensia jantarica (153735) Hoffeinsensia jantarica, Sucinocantharis baltica, Electronycha prussica, Themus pristinus, Cantharis hanswerneri, Cantharis hoffeinsorum, Lycocerus christelae, Lycocerus dentantennatus (123419) Michalskantharis bursztynica, Lycocerus xenium, Podistra jirii, Cacomorphocerus incurvus, Electronycha arturmichalskii (185560) | |
| Priabonian | Ukraine | Cantharis michaeli (231022) Lycocerus albovacca (236501) Sucinorhagonycha pugachensis, Cacomorphocerus sp. (122997) | |
| Priabonian | Poland | Cacomorphocerus cerambyx (123438) Cacomorphocerus wiszniewskii, Cantharis dougi (44615) Lycocerus elzbietae, Lycocerus jonasi, Rhagonycha acarigera, Lycocerus crassepunctatus (110737) Sucinorhagonycha kulickae (123215) Sucinorhagonycha kulickae, Absidiella sucinokotejai, Cantharis sucinonigra (120813) | |
| Chadronian | USA (Colorado) | Podabrus fragmentatus, Podabrus wheeleri, Podabrus florissantensis, Telephorus hesperus (106384) Podabrus wheeleri (106379) Podabrus wheeleri, Podabrus cupesoides (124624) | |
| Chattian | Germany (North Rhine-Westphalia) | Cantharis bradburyi, Cantharis lidiae, Cantharis rottensis, Podistra quies, Rhagonycha carolynae, Rhagonycha ultramundana (122010) | |
| Late/Upper Oligocene | France (Puy-de-Dome) | Telephorus sp. (155204) | |
| MP 28 | Germany (Rheinland-Pfalz) | Cantharis doernerorum (211302) Cantharis zabolica (211301) | |
| MP 30 | Germany | Telephorus carbonarius, Telephorus caducus, Telephorus brodiei (123945) Telephorus exauctaratus (154946) | |
| MP 30 | Germany (Nordrhein-Westfalen) | Telephorus carbonarius (138696) | |
| Burdigalian | China (Shandong) | Curticantharis capacis, Cantharis guttula (128551) Curticantharis sp. (128552) Curticantharis thermophila (130412) Curticantharis trapezialis (138944) | |
| Sarmatian | Germany (Baden-Württemberg) | Telephorus germari, Telephorus fragilis (115305) Telephorus macilentus (117252) | |
| Sarmatian | Croatia | Telephorus attavinus (117250) | |
| Tortonian | Romania | Cantharis sesermanae (221024) | |
| Gelasian | France | Rhagonycha micans (113996) | |
| Gelasian | Greenland | Podabrus alpinus (160109) | |
| Late/Upper Pleistocene | United Kingdom (Worcestershire) | Cantharis figurata, Cantharis rufa, Cantharis paludosa (163070) | |
| Tarantian | USA (Pennsylvania) | Podabrus sp. (137205 137206) |