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Pachyrhachis problematicus
Taxonomy
Pachyrhachis problematicus was named by Haas (1979). Its type specimen is HUJ-PAL 3659, a skeleton (almost complete), and it is a 3D body fossil. Its type locality is 'Ein Yabrud quarries, which is in a Cenomanian shallow subtidal limestone in the Bet Meir or Amminadav Formation of Palestine. It is the type species of Pachyrhachis.
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1979 | Pachyrhachis problematicus Haas p. 62 |
| 1980 | Ophiomorphus colberti Haas |
| 1984 | Estesius colberti Wallach p. 329 |
| 1998 | Pachyrhachis problematicus Lee and Caldwell p. 1524 |
| 2013 | Pachyrhachis problematicus Palci et al. p. 1335 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
†Pachyrhachis problematicus Haas 1979
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Invalid names: Ophiomorphus colberti Haas 1980 [synonym]
Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| M. Y. Lee and M. Caldwell 1998 | A long-bodied marine squamate. The head is small, and the cervical region narrow. The shoulder girdle and forelimbs are absent, a small pelvis and hindlimbs are present. It exhibits the following autapomorphies not found in other squamates: (i) the frontal is extremely long and narrow; (ii) the quadrate is sheet-like, being greatly expanded anteroposteriorly; (iii) there is a large rectangular coronoid process; (iv) the coronoid bone has an extremely long anterior £ange which extends horizontally along the medial surface of the dentary; (v) the splenial-angular joint is located very far anteriorly, approximately mid-way between the symphysis and the coronoid process; and (vi) the middle trunk vertebrae and ribs are pachyostotic. | |
| A. Palci et al. 2013 | Snake with well-developed hind limbs consisting at least of femur, tibia, fibula, astragalus, and calcaneum; ilium, ischium, and pubis present; long and nar- row frontals; quadrate expanded anteroposteriorly and sheet-like; dorsal process of coronoid high, taller than maximum depth of compound bone; retains a jugal; long anterior flange of coronoid extends horizontally along medial surface of dentary; splenial- angular joint located far anteriorly, midway between symphysis and coronoid process; numerous (at least four) small mental foramina on dentary; pachyostotic middle trunk vertebrae and ribs; retains a sacral vertebra with sacral ribs articulated to the centrum; at least 13 tooth positions on dentary, at least seven on palatine, and at least six on pterygoid and maxilla. |
Measurements
No measurements are available
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| Source: g = genus, c = class, subp = subphylum | |||||
| References: Hendy et al. 2009, Haas 1979, Carroll 1988 | |||||
Age range: Early/Lower Cenomanian or 100.50000 to 93.90000 Ma
Collections: one only
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Early/Lower Cenomanian | Palestine | Pachyrhachis problematicus, Ophiomorphus colberti (type locality: 11804) |