PBDB Taxon

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Clavagnostus

Artiopoda - Agnostida - Clavagnostidae

Taxonomy

Clavagnostus was named by Howell (1937) [Sepkoski's age data: ]. It is not extant. It is the type genus of Clavagnostidae, Clavagnostinae.

It was assigned to Agnostida by Sepkoski (2002); to Clavagnostidae by Whittington et al. (1997), Jell and Adrain (2002); and to Clavagnostinae by Jago et al. (2004), Bentley et al. (2006).

Species

C. (Paraclavagnostus), C. burnsi, C. calensis (syn. C. (Leptagnostus) lanceolatus, C. canotensis, C. rawlingi), C. cuneatus, C. repandus, C. spinosus (syn. C. bisectus, C. sulcatus, C. longispinus), C. subconicus, C. taoyuanensis, C. trispinus

Synonyms

Acanthagnostus was named by Qian (1982). It is not extant.

It was synonymized subjectively with Clavagnostus by Peng and Robison (2000), Jell and Adrain (2002).
Tomorhachis was named by Resser (1938). It is not extant. Its type is Tomorhachis spinosa.

It was synonymized subjectively with Clavagnostus by Peng and Robison (2000).
Clavagnostus (Leptagnostus) was named by Lu and Lin (1989). It is not extant.

It was synonymized subjectively with Clavagnostus by Shergold and Laurie (1997), Jell and Adrain (2002).

It was assigned to Clavagnostus by Lu and Lin (1989).
Culipagnostus was named by Rusconi (1952).

It was synonymized subjectively with Clavagnostus by Jell and Adrain (2002).

Synonymy list

Year	Name and author
1937	Clavagnostus Howell
1938	Tomorhachis Resser
1952	Culipagnostus Rusconi
1982	Acanthagnostus Qian
1989	Clavagnostus (Leptagnostus) Lu and Lin
1997	Clavagnostus Whittington et al. p. 371
2002	Clavagnostus Jell and Adrain p. 358
2002	Clavagnostus Sepkoski
2004	Clavagnostus Jago et al. p. 29
2006	Clavagnostus Bentley et al. p. 172

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Gravenhorst 1843

Rank	Name	Author
class	Artiopoda	Hou and Bergstrom
order	Agnostida	Salter 1864
suborder	Agnostina	Salter 1864
superfamily	Condylopygoidea	Raymond 1913
family	Clavagnostidae	Howell 1937
subfamily	Clavagnostinae	Howell 1937
genus	Clavagnostus	Howell 1937

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Clavagnostus Howell 1937

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Subg. †Clavagnostus (Paraclavagnostus) Yang et al. 1993

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†Clavagnostus (Paraclavagnostus) variatus Yang et al. 1993

†Clavagnostus burnsi Jago and Daily 1974

†Clavagnostus calensis Rusconi 1950

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Invalid names: Clavagnostus (Leptagnostus) lanceolatus Lu and Lin 1989 [synonym], Clavagnostus canotensis Rusconi 1951 [synonym]

†Clavagnostus cuneatus Romanenko 1967

†Clavagnostus repandus Westergard 1930

†Clavagnostus spinosus Resser 1938

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Invalid names: Clavagnostus bisectus Opik 1967 [synonym], Clavagnostus longispinus Qian 1982 [synonym], Clavagnostus sulcatus Westergard 1946 [synonym]

†Clavagnostus subconicus Lu and Lin 1989

†Clavagnostus taoyuanensis Lin 1991

†Clavagnostus trispinus Zhou and Yang 1977

Invalid names: Acanthagnostus Qian 1982 [synonym], Culipagnostus Rusconi 1952 [synonym], Leptagnostus Lu and Lin 1989 [synonym], Tomorhachis Resser 1938 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Entire body:

yesg

Adult length:

1.0 to < 10g

Adult width:

1.0 to < 10g

Locomotion:

slow-movingo

Life habit:

epifaunalo

Diet:

detritivoreo

Vision:

blindf

Comments:

BODY SIZE: Data from Williams et al. (2002).g

Created:

2009-06-02 14:45:14

Modified:

2009-06-02 16:45:14

Source: g = genus, f = family, o = order

References: Whittington et al. 1997, Kiessling 2004, Hendy 2009

Age range: base of the Zhanaarykian to the top of the Mindyallan or 503.00000 to 497.00000 Ma

Collections (59 total)

Time interval	Ma	Country or state	Original ID and collection number
Miaolingian	506.5 - 497.0	China	C. sp. (228206)
Middle Cambrian	506.5 - 497.0	Argentina (Mendoza)	C. calensis, C. repandus (9811)
Middle Cambrian	506.5 - 497.0	India	C. calensis, C. trispinus (114635) C. trispinus (114663)
Mayan	504.5 - 499.5	Russian Federation (Gorniy Altay)	C. repandus (10174)
Menevian	504.5 - 499.5	USA (Utah)	C. repandus (10572)
Marjumian	504.5 - 497.0	USA (Utah)	C. spinosus (230681)
Wangcunian	504.5 - 497.0	China (Hunan)	C. trispinus (128337 128339)
Wangcunian - Guzhangian	504.5 - 497.0	China (Zhejiang)	C. lanceolatus (125541) C. sp. (125683)
Wangcunian - Guzhangian	504.5 - 497.0	China (Hunan)	C. taoyuanensis (128927)
Wangcunian - Guzhangian	504.5 - 497.0	China (Sichuan)	C. repandus (127066 127067)
Zhanaarykian	503.0 - 500.5	Kazakhstan	C. calensis, C. repandus (9958 118102)
Boomerangian	500.5 - 499.0	Australia (Tasmania)	C. burnsi (69973)
Guzhangian	500.5 - 497.0	China (Sichuan)	C. repandus, C. variatus (127068 127069 127070)
Guzhangian	500.5 - 497.0	China (Zhejiang)	C. sp., C. sp. (125673) C. sp., C. subconicus (125675)
Guzhangian	500.5 - 497.0	China (Shandong)	Acanthagnostus longispinus (124885) C. longispinus (126818)
Guzhangian	500.5 - 497.0	China (Hunan)	C. burnsi (128930) C. repandus (49169 49170 49174 49303 128356) C. sulcatus (128931) C. trispinus (49168 49175 49295 128365) C. trispinus, C. repandus (49172 49176)
Guzhangian	500.5 - 497.0	China (Yunnan)	C. spinosus (219647)
Ayusokkanian	499.5 - 497.0	Russian Federation (Altai (Region))	C. cuneatus (10064)
Ayusokkanian	499.5 - 497.0	Kazakhstan	C. calensis (9950) C. calensis, C. repandus (118158 118164 118165) C. repandus (9943 9944 9945 9947 118166 118168) C. spinosus (118191 118236 118237 118241) C. trispinus (118179 118250) C. trispinus, C. spinosus (118178 118184) C. trispinus, C. spinosus, C. repandus (118177)
Mindyallan	499.0 - 497.0	Antarctica	C. spinosus (242394)