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Orthophlebiidae
Taxonomy
Orthophlebidae was named by Handlirsch (1906). It is not extant.
It was reranked as the subfamily Orthophlebiinae by Sukatsheva (1985); it was corrected as Orthophlebiidae by Willmann (1978), Willmann (1987), Krzemiński et al. (2015), Soszyńska-Maj et al. (2019).
It was assigned to Panorpata by Handlirsch (1906); to Panorpidae by Sukatsheva (1985); to Eumecoptera by Tillyard (1933), Riek (1955), Whalley (1985); to Mecoptera by Martynov (1927), Martynov (1937), Riek (1950), Bode (1953), Willmann (1978), Carpenter (1992), Zhang (1996); and to Panorpoidea by Willmann (1987), Krzemiński et al. (2015), Soszyńska-Maj et al. (2019).
It was reranked as the subfamily Orthophlebiinae by Sukatsheva (1985); it was corrected as Orthophlebiidae by Willmann (1978), Willmann (1987), Krzemiński et al. (2015), Soszyńska-Maj et al. (2019).
It was assigned to Panorpata by Handlirsch (1906); to Panorpidae by Sukatsheva (1985); to Eumecoptera by Tillyard (1933), Riek (1955), Whalley (1985); to Mecoptera by Martynov (1927), Martynov (1937), Riek (1950), Bode (1953), Willmann (1978), Carpenter (1992), Zhang (1996); and to Panorpoidea by Willmann (1987), Krzemiński et al. (2015), Soszyńska-Maj et al. (2019).
Subtaxa
Entered
by E. Leckey (authorized by D. Smith) on 2008-12-02; modified by M. Clapham on 2021-03-24
Synonymy list
| Year | Name and author |
|---|---|
| 1906 | Orthophlebidae Handlirsch p. 479 |
| 1927 | Orthophlebidae Martynov |
| 1933 | Orthophlebidae Tillyard p. 23 |
| 1937 | Orthophlebidae Martynov |
| 1950 | Orthophlebidae Riek p. 254 |
| 1953 | Orthophlebidae Bode |
| 1955 | Orthophlebidae Riek p. 662 |
| 1978 | Orthophlebiidae Willmann p. 64 |
| 1985 | Orthophlebiinae Sukatsheva p. 97 |
| 1985 | Orthophlebidae Whalley |
| 1987 | Orthophlebiidae Willmann |
| 1992 | Orthophlebidae Carpenter |
| 1996 | Orthophlebidae Zhang p. 446 |
| 2015 | Orthophlebiidae Krzemiński et al. p. 502 |
| 2019 | Orthophlebiidae Soszyńska-Maj et al. p. 1082 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Fm. †Orthophlebiidae Handlirsch 1906
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G. †Burmorthophlebia Soszyńska-Maj et al. 2022
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†Burmorthophlebia macularis Zhang et al. 2023
†Burmorthophlebia multiprocessa Soszyńska-Maj et al. 2022
G. †Choristopanorpa Riek 1950
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†Choristopanorpa bifasciata Riek 1950
†Choristopanorpa drinnani Jell and Duncan 1986
†Choristopanorpa opinata Novokshonov 2001
†Choristopanorpa ridibunda Novokshonov 2001
†Choristopanorpa temperata Novokshonov 2001
Subfm. †Gigaphlebiinae Soszyńska-Maj and Krzemiński 2018
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G. †Gigaphlebia Soszyńska-Maj and Krzemiński 2018
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†Gigaphlebia grandis Martynov 1927
†Gigaphlebia palmaris Martynova 1948
†Gigaphlebia riccardii Petrulevičius and Ren 2012
G. †Longiphlebia Soszyńska-Maj and Krzemiński 2018
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†Longiphlebia incompleta Lian et al. 2021
†Longiphlebia stigmosa Qiao et al. 2012
G. †Jibeiorthophlebia Hong 1983
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†Jibeiorthophlebia internata Hong 1983
†Jibeiorthophlebia xiaofanzhangziensis Hong 1983
G. †Juraphlebia Soszyńska-Maj and Krzemiński 2019
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†Juraphlebia eugeniae Soszyńska-Maj and Krzemiński 2019
G. †Mesopanorpa Handlirsch 1906
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†Mesopanorpa angarensis Martynov 1927
†Mesopanorpa brooksorum Jarzembowski and Soszyńska-Maj 2018
†Mesopanorpa felix Martynov 1927
†Mesopanorpa formosa Bode 1953
†Mesopanorpa gambra Lin 1980
†Mesopanorpa hartungi Brauer et al. 1889
†Mesopanorpa incerta Martynov 1927
†Mesopanorpa kuschmurunensis Martynova 1956
†Mesopanorpa maculata Handlirsch 1939
†Mesopanorpa martynovae Sukacheva 1990
†Mesopanorpa obtusa Bode 1953
†Mesopanorpa praeangustipennis Martynova 1956
†Mesopanorpa tergenica Sukacheva 1990
†Mesopanorpa turgaica Martynova 1956
†Mesopanorpa umbrata Martynov 1937
†Mesopanorpa unica Sukatsheva 1985
†Mesopanorpa unicolor Martynov 1937
Subfm. †Orthophlebiinae Handlirsch 1906
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G. †Orthophlebia Westwood 1845
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Subg. †Orthophlebia (Dolichophlebia) Hong and Zhang 2004
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†Orthophlebia (Dolichophlebia) ladinica Hong 2009
†Orthophlebia (Dolichophlebia) diminuta Bode 1953
Subg. †Orthophlebia (Mesopanorpa) Handlirsch 1906
Subg. †Orthophlebia (Orthophlebia) Westwood 1845
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Invalid names: Orthophlebia (Brachyophlebia) Hong and Zhang 2004 [synonym]
Subg. †Orthophlebia (Parisophlebia) Hong and Zhang 2004
†Orthophlebia aequalis Martynov 1937
†Orthophlebia anglica Handlirsch 1939
†Orthophlebia angustata Martynov 1937
†Orthophlebia bella Handlirsch 1939
†Orthophlebia bifurcata Giebel 1856
†Orthophlebia bolboica Sukacheva 1990
†Orthophlebia brodiei Tillyard 1933
†Orthophlebia brunsvicensis Bode 1905
†Orthophlebia capillata Whalley 1985
†Orthophlebia chinensis Soszyńska-Maj et al. 2019
†Orthophlebia colorata Zhang 1996
†Orthophlebia compacta Bode 1953
†Orthophlebia confusa Willmann 1977
†Orthophlebia curta Martynova 1948
†Orthophlebia deformis Lin 1986
†Orthophlebia densa Zhang 1996
†Orthophlebia effusa Martynova 1948
†Orthophlebia elenae Willmann and Novokshonov 1998
†Orthophlebia elongata Handlirsch 1939
†Orthophlebia exculpta Zhang 1996
†Orthophlebia extensa Martynov 1937
†Orthophlebia fallerslebensis Bode 1953
†Orthophlebia fanshanensis Ren 1995
†Orthophlebia fracta Sukacheva 1990
†Orthophlebia furcata Handlirsch 1939
†Orthophlebia fuscipennis Handlirsch 1906
†Orthophlebia germanica Handlirsch 1906
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Invalid names: Orthophlebia geinitzi Handlirsch 1906 [synonym], Synorthophlebia dubia Handlirsch 1939 [synonym]
†Orthophlebia gracilis Handlirsch 1939
†Orthophlebia gubini Novokshonov and Sukatsheva 2003
†Orthophlebia heidemariae Willmann and Novokshonov 1998
†Orthophlebia intermedia Giebel 1856
†Orthophlebia jejuna Sukatsheva 1985
†Orthophlebia karabonica Sukacheva 1990
†Orthophlebia kuliki Martynova 1948
†Orthophlebia laesa Handlirsch 1939
†Orthophlebia latebrosa Sukatsheva 1985
†Orthophlebia latipennisimilis Bode 1953
†Orthophlebia lauta Sukatsheva 1985
†Orthophlebia liadis Handlirsch 1939
†Orthophlebia liaoningensis Ren 1997
†Orthophlebia liassica Mantell 1844
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Invalid names: Orthophlebia communis Westwood 1845 [synonym], Orthophlebia lata Giebel 1856 [synonym], Orthophlebia similis Giebel 1856 [synonym], Orthophlebia simillima Handlirsch 1939 [synonym]
†Orthophlebia limnophila Handlirsch 1906
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Invalid names: Orthophlebioides latipennis Handlirsch 1906 [synonym], Orthophlebioides reticulatus Handlirsch 1906 [synonym]
†Orthophlebia lini Lian and Huang 2024
†Orthophlebia lithographica Willmann and Novokshonov 1998
†Orthophlebia longicauda Willmann and Novokshonov 1998
†Orthophlebia luanpingensis Hong 1983
†Orthophlebia maculata Martynov 1927
†Orthophlebia mongolica Sukatsheva 1985
†Orthophlebia monstrosa Zhang 1996
†Orthophlebia nana Handlirsch 1939
†Orthophlebia nervulosa Qiao et al. 2012
†Orthophlebia obunca Sukatsheva 1985
†Orthophlebia parvula Handlirsch 1939
†Orthophlebia phryganoides Martynov 1925
†Orthophlebia picta Lian 2024
†Orthophlebia pictipennis Tillyard 1933
†Orthophlebia pulchra Martynova 1956
†Orthophlebia pygmaea Handlirsch 1939
†Orthophlebia quadrimacula Lin 1982
†Orthophlebia radialis Handlirsch 1939
†Orthophlebia retorrida Sukatsheva 1985
†Orthophlebia rossica Martynova 1948
†Orthophlebia rotundipennis Martynov 1937
†Orthophlebia shartegica Novokshonov and Sukatsheva 2003
†Orthophlebia shurabica Martynov 1937
†Orthophlebia speciosa Bode 1953
†Orthophlebia stigmatica Handlirsch 1939
†Orthophlebia varia Martynova 1948
†Orthophlebia venosa Martynov 1937
†Orthophlebia vernacula Martynova 1948
†Orthophlebia vicina Handlirsch 1939
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Invalid names: Synorthophlebia elegans Handlirsch 1939 [synonym]
†Orthophlebia xiangyu Lian 2025
†Orthophlebia yangjuanxiangensis Hong 1985
†Orthophlebia yaogouensis Hong 1983
†Orthophlebia yaojiashanensis Lin 1980
Invalid names: Callopanorpa Handlirsch 1906 [synonym], Orthophlebiites Handlirsch 1939 [synonym], Orthophlebioides Handlirsch 1906 [synonym], Synorthophlebia Handlirsch 1939 [synonym]
G. †Parachorista Lin 1976
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: c = class, p = phylum | |||||
| References: Kiessling 2004, Bush and Bambach 2015 | |||||
Age range: base of the Anisian to the top of the Early/Lower Cenomanian or 247.00000 to 93.90000 Ma
Collections (122 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Anisian | Australia (New South Wales) | Choristopanorpa bifasciata (106483) | |
| Ladinian | Kyrgyzstan (Osh) | Choristopanorpa opinata (106090 113406) Choristopanorpa opinata, Choristopanorpa ridibunda (110763) Choristopanorpa ridibunda (110765) Choristopanorpa temperata (105007) | |
| Longobardian | China (Shaanxi) | Orthophlebia ladinica (117138) Orthophlebia xiangyu (183209) | |
| Norian | Australia (Queensland) | Orthophlebidae indet. (111558) | |
| Rhaetian | United Kingdom (England) | Orthophlebia communis (124687) Orthophlebia communis, Orthophlebia liassica, Orthophlebioides anglicus (115222) Orthophlebia gracilis (139595) Orthophlebia liassica (113957 114643 114677 114680) Orthophlebia liassica, Chauliodes sp. 3 (113961) Orthophlebia liassica, Orthophlebia pictipennis, Orthophlebia confusa (115223) Panorpa liassica, Orthophlebia simillima (115220) | |
| Rhaetian - Early/Lower Hettangian | United Kingdom (England) | Orthophlebia communis, Chauliodes sp. (124688) | |
| Rhaetian - Hettangian | United Kingdom | Orthophlebia pygmaea, Orthophlebia bella (139594) | |
| Rhaetian - Hettangian | United Kingdom (Gloucestershire) | Orthophlebia nana, Orthophlebia laesa, Orthophlebioides stigmaticus, Orthophlebioides furcatus, Orthophlebioides parvulus (139593) | |
| Early/Lower Hettangian | United Kingdom (England) | Orthophlebia communis, Orthophlebia liassica (113956) Orthophlebia liassica (113960) | |
| Hettangian | United Kingdom (England) | Orthophlebia liassica (115224) | |
| Planorbis | United Kingdom (England) | Orthophlebia pictipennis (123982) | |
| Hettangian - Sinemurian | Kyrgyzstan (Tonskiy) | Mesopanorpa kuliki, Orthophlebia effusa, Orthophlebia varia (105086) Orthophlebia maculata, Mesopanorpa hartungi, Orthophlebia extensa, Mesopanorpa umbrata, Mesopanorpa unicolor, Orthophlebia curta, Orthophlebia churabica, Orthophlebia effusa, Orthophlebia varia, Orthophlebia vernacula, Mesopanorpa palmaris, Mesopanorpa ampla, Orthophlebia rossica (105087) | |
| Hettangian - Sinemurian | China (Xinjiang) | Orthophlebia latebrosa, Orthophlebia colorata, Mesopanorpa brodiei, Mesopanorpa kuliki, Mesopanorpa densa, Mesopanorpa monstrosa (103480) | |
| Hettangian - Sinemurian | United Kingdom (Warwickshire) | Orthophlebia communis, Chauliodes sp. (133185) | |
| Early/Lower Jurassic - Middle Jurassic | Kazakhstan | Mesopanorpa kuschmurunensis (172378) Mesopanorpa sp. (172379 172380) Mesopanorpa turgaica (172377) Orthophlebia pulchra (172375) Orthophlebia sp. (172376) | |
| Jurassic | Kazakhstan | Mesopanorpa praeangustipennis (172381) | |
| Obtusum | United Kingdom (England) | Orthophlebia capillata (114638) | |
| Pliensbachian | Kyrgyzstan | Orthophlebia aequalis (114701) | |
| Early/Lower Toarcian | Germany (Niedersachsen) | Orthophlebia brunsvicensis (107556) Orthophlebia compacta, Orthophlebia latipennisimilis, Orthophlebia diminuta, Orthophlebia formosa, Synorthophlebia speciosa, Synorthophlebia obtusa (105002) Orthophlebia fallerslebensis (108447) Orthophlebia formosa (107343) Orthophlebia latipennisimilis (107344) Parorthophlebia grasselensis, Orthophlebia latipennisimilis (107321) | |
| Early/Lower Toarcian | Germany | Orthophlebia geinitzi, Orthophlebia germanica, Synorthophlebia dubia, Orthophlebia elongata, Orthophlebioides fuscipennis, Synorthophlebia vicina, Synorthophlebia elegans, Synorthophlebia liadis, Orthophlebiites radialis, Synorthophlebia maculata, Orthophlebia intermedia, Orthophlebia intermedia (123987) | |
| Early/Lower Toarcian | United Kingdom (Gloucestershire) | Orthophlebia brodiei (153651) | |
| Early/Lower Toarcian | Germany (Mecklenburg-Vorpommern) | Orthophlebia brunsvicensis, Orthophlebia germanica, Orthophlebia elongata (114993) | |
| Falciferum | Germany | Orthophlebia elongata (192037) | |
| Falciferum | United Kingdom (England) | Orthophlebia brodiei (132073) Orthophlebia sp. (234545) | |
| Toarcian | Kyrgyzstan | Mesopanorpa unicolor (114705) Orthophlebia angustata (114704) Orthophlebia extensa, Orthophlebia shurabica, Orthophlebia venosa, Mesopanorpa umbrata (114702) Orthophlebia rotundipennis (114703) | |
| Toarcian | Russian Federation (Irkutsk) | Mesopanorpa angarensis, Mesopanorpa incerta (114694) Orthophlebia latebrosa (104946) Orthophlebia lauta, Orthophlebia retorrida (113471) Panorpa hartungi (114696) | |
| Toarcian | China (Guangxi) | Protorthophlebia deformis (151540) | |
| Toarcian | Russian Federation (Krasnoyarsk) | Orthophlebia jejuna, Orthophlebiinae indet. (110819) | |
| Toarcian | Germany | Orthophlebioides limnophilus (131767) | |
| Toarcian - Aalenian | China (Hebei) | Protorthophlebia yangjuanxiangensis (164199) | |
| Aalenian | China (Shaanxi) | Orthophlebia lini (222545 230680) | |
| Aalenian - Bajocian | China (Xinjiang) | Orthophlebia exculpta (117193) | |
| Middle Jurassic | China (Gansu) | Orthophlebia quadrimacula (154785) | |
| Bajocian - Bathonian | China (Anhui) | Orthophlebia rotundipennis (106695) | |
| Bajocian - Bathonian | Russian Federation (Krasnoyarsk) | Orthophlebia sp., Mesopanorpa sp., Orthophlebia obunca, Orthophlebiinae indet. (118010) | |
| Bathonian | Mongolia | Orthophlebia mongolica (119621) | |
| Callovian | China (Henan) | Orthophlebiidae indet., Orthophlebia quadrimacula, Jiyuanphlebia incompleta, Orthophlebia picta, Quadrivena jiyuanensis (202211) | |
| Callovian - Oxfordian | China (Nei Mongol) | Juraphlebia eugeniae (172659) Longiphlebia stigmosa (155751) Orthophlebia chinensis, Juraphlebia eugeniae (112663 196979) Orthophlebia extensa, Mesopanorpa densa, Mesopanorpa luanpingensis, Orthophlebia elenae, Juraphlebia eugeniae (123761 183155) Orthophlebia extensa, Mesopanorpa luanpingensis, Orthophlebia elenae (113360) Orthophlebia extensa, Mesopanorpa luanpingensis, Orthophlebia elenae, Juraphlebia eugeniae (211893) Orthophlebia extensa, Orthophlebia nervulosa, Orthophlebia stigmosa (113365) Orthophlebia riccardii, Orthophlebia nervulosa, Orthophlebia stigmosa (117610) | |
| Callovian - Oxfordian | China (Liaoning) | Jibeiorthophlebia internata, Jibeiorthophlebia xiafanzhangziensis (113475) | |
| Callovian - Oxfordian | China (Hebei) | Mesopanorpa luanpingensis (154786) Protorthophlebia yaogouensis (154788) | |
| Oxfordian | Russian Federation (Buryat) | Mesopanorpa unica (105017) | |
| Oxfordian | China (Hebei) | Longiphlebia incompleta (224167) | |
| Oxfordian - Kimmeridgian | Russian Federation (Buryat) | Mesopanorpa sp. (125482) | |
| Kimmeridgian | Kazakhstan | Mesopanorpa felix (114692) Orthophlebia elenae (105971 122330) Orthophlebia elenae, Orthophlebia longicauda (105426) Orthophlebia grandis (105970) Orthophlebia grandis, Orthophlebia phryganoides (115717) Orthophlebia heidemariae (106101) Orthophlebia maculata (114693) | |
| Early/Lower Tithonian | Germany (Bayern) | Orthophlebia lithographica (105797) | |
| Tithonian | Russian Federation | Mesopanorpa tergenica (138150) Orthophlebia bolboica (150500) Orthophlebidae indet. (123198) | |
| Tithonian | Russian Federation (Chita) | Mesopanorpa martynovae (119397) Orthophlebidae indet. (114307) | |
| Tithonian | Mongolia | Orthophlebia shartegica (118966 121394 152133) Orthophlebia sp., Orthophlebia gubini, Orthophlebia shartegica, Orthophlebiidae indet. (104748) | |
| Middle Berriasian | United Kingdom (England) | Orthophlebia sp. 2, Orthophlebia sp. 1 (104008) | |
| Valanginian - Hauterivian | China (Hebei) | Sinorthophlebia weichangensis (276554) | |
| Late/Upper Hauterivian | United Kingdom (England) | Mesopanorpa brooksorum (106417) | |
| Late/Upper Hauterivian | United Kingdom (Surrey) | Mesopanorpa brooksorum (74972) | |
| Barremian | China (Zhejiang) | Mesopanorpa gambra (155607) Mesopanorpa yaojiashanensis (155606) | |
| Late/Upper Barremian | Spain (Castilla-La Mancha) | Orthophlebiinae indet. (28006) | |
| Late/Upper Barremian | China (Liaoning) | Orthophlebia liaoningensis (106966 138436) | |
| Aptian | Russian Federation (Chita) | Orthophlebia fracta (138072) Orthophlebia karabonica (126001) | |
| Aptian | Russian Federation (Buryatia) | Orthophlebiidae indet. (113349 113350) | |
| Late/Upper Aptian | Australia (Victoria) | Choristopanorpa drinnani, Cretacochorista parva, Orthophlebiidae indet. (77192) | |
| Late/Upper Aptian | China (Beijing Shi) | Orthophlebia fanshanensis (106965) | |
| Early/Lower Cenomanian | Myanmar | Burmorthophlebia macularis (165561) Burmorthophlebia multiprocessa (187929) |