PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Keroplatidae (predatory fungus gnat)

Insecta - Diptera - Keroplatidae

Taxonomy

Keroplatidae was named by Rondani (1856). It is extant. Its type is Keroplatus.

It was reranked as the subfamily Ceroplatinae by Johannsen (1909), Enderlein (1911), Gagné (1980).

It was assigned to Mycetophilidae by Johannsen (1909), Enderlein (1911), Gagné (1980); to Mycetophiloidea by Matile (1981); to Diptera by Solórzano Kraemer and Evenhuis (2008); and to Sciaroidea by Yeates et al. (2007), Blagoderov et al. (2019), Blagoderov (2019).

Synonyms

Platyura ehrhardti was named by Loew (1850). It is an inclusion in amber. Its type locality is Baltic amber, Konigsberg collection, which is in a Priabonian terrestrial amber in the Russian Federation.

It was considered a nomen dubium by Evenhuis (2006).
Platyura kunowi was named by Meunier (1904). It is an inclusion in amber.

It was considered a nomen dubium by Evenhuis (2006).
Platyura verrali was named by Meunier (1904). It is an inclusion in amber. Its type locality is Baltic amber, Konigsberg collection, which is in a Priabonian terrestrial amber in the Russian Federation.

It was considered a nomen dubium by Evenhuis (2006).
Platyura graciosa was named by Meunier (1904). It is an inclusion in amber. Its type locality is Baltic amber, Konigsberg collection, which is in a Priabonian terrestrial amber in the Russian Federation.

It was considered a nomen dubium by Evenhuis (2006).
Platyura moniliformis was named by Meunier (1904). It is an inclusion in amber. Its type locality is Baltic amber, Konigsberg collection, which is in a Priabonian terrestrial amber in the Russian Federation.

It was considered a nomen dubium by Evenhuis (2006).
Platyura ectorsii was named by Meunier (1904). It is an inclusion in amber. Its type locality is Baltic amber, Konigsberg collection, which is in a Priabonian terrestrial amber in the Russian Federation.

It was considered a nomen dubium by Evenhuis (2006).
Platyura mikii was named by Meunier (1904). It is an inclusion in amber.

It was considered a nomen dubium by Evenhuis (2006).
Platyura distincta was named by Meunier (1904). It is an inclusion in amber. Its type locality is Baltic amber, Konigsberg collection, which is in a Priabonian terrestrial amber in the Russian Federation.

It was considered a nomen dubium by Evenhuis (2006).
Platyura ceroplatoides was named by Meunier (1904). It is an inclusion in amber. Its type locality is Baltic amber, Konigsberg collection, which is in a Priabonian terrestrial amber in the Russian Federation.

It was considered a nomen dubium by Evenhuis (2006).
Platyura ceroplatites was named by Meunier (1904). It is an inclusion in amber. Its type locality is Baltic amber, Konigsberg collection, which is in a Priabonian terrestrial amber in the Russian Federation.

It was considered a nomen dubium by Evenhuis (2006).
Platyura crassicornis was named by Meunier (1917). It is an inclusion in amber. Its type locality is Baltic amber, Konigsberg collection, which is in a Priabonian terrestrial amber in the Russian Federation.

It was considered a nomen dubium by Evenhuis (2006).
Platyura exigua was named by Meunier (1907). It is an inclusion in amber. Its type locality is Zanzibar Copal (Evers collection), which is in a Holocene terrestrial amber in Tanzania.

It was considered a nomen dubium by Evenhuis (2006).
Platyura armata was named by Meunier (1899). It is an inclusion in amber. Its type locality is Baltic amber, Konigsberg collection, which is in a Priabonian terrestrial amber in the Russian Federation.

It was considered a nomen dubium by Evenhuis (2006).
Platyura calcar was named by Meunier (1899). It is an inclusion in amber. Its type locality is Baltic amber, Konigsberg collection, which is in a Priabonian terrestrial amber in the Russian Federation.

It was considered a nomen dubium by Evenhuis (2006).
Platyura pusilla was named by Meunier (1899). It is an inclusion in amber. Its type locality is Baltic amber, Konigsberg collection, which is in a Priabonian terrestrial amber in the Russian Federation.

It was considered a nomen dubium by Evenhuis (2006).

Synonymy list

Year	Name and author
1850	Platyura ehrhardti Loew p. 35
1856	Keroplatidae Rondani
1899	Platyura armata Meunier p. 164
1899	Platyura calcar Meunier p. 164
1899	Platyura ehrhardti Meunier p. 165
1899	Platyura pusilla Meunier p. 165 figs. Pl 1, fig 10
1904	Platyura ehrhardti Meunier p. 106
1904	Platyura kunowi Meunier p. 107
1904	Platyura verrali Meunier p. 108
1904	Platyura graciosa Meunier p. 109
1904	Platyura moniliformis Meunier p. 110
1904	Platyura ectorsii Meunier p. 111
1904	Platyura distincta Meunier p. 112
1904	Platyura mikii Meunier p. 112
1904	Platyura ceroplatoides Meunier p. 113
1904	Platyura ceroplatites Meunier p. 114
1907	Platyura exigua Meunier p. 54 fig. 9
1909	Ceroplatinae Johannsen p. 13
1909	Platyura armata Johannsen p. 21
1909	Platyura calcar Johannsen p. 21
1909	Platyura ceroplatites Johannsen p. 21
1909	Platyura ceroplatoides Johannsen p. 21
1909	Platyura distincta Johannsen p. 22
1909	Platyura ectorsii Johannsen p. 22
1909	Platyura ehrhardti Johannsen p. 22
1909	Platyura exigua Johannsen p. 22
1909	Platyura graciosa Johannsen p. 23
1909	Platyura kunowi Johannsen p. 23
1909	Platyura mikii Johannsen p. 23
1909	Platyura moniliformis Johannsen p. 24
1909	Platyura pusilla Johannsen p. 24
1909	Platyura verrali Johannsen p. 25
1911	Ceroplatinae Enderlein p. 145
1917	Platyura crassicornis Meunier p. 91 figs. Pl 13, fig 52
1980	Ceroplatinae Gagné
1981	Keroplatidae Matile
2007	Keroplatidae Yeates et al.
2008	Keroplatidae Solórzano Kraemer and Evenhuis p. 57
2019	Keroplatidae Blagoderov p. 541
2019	Keroplatidae Blagoderov et al. p. 643

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Gravenhorst 1843
—	Deuteropoda
—	Mandibulata
—	Pancrustacea	Zrzavý and Štys 1997
—	Altocrustacea	Regier et al. 2010
superclass	Allotriocarida
—	Athalassocarida
—	Labiocarida	Schwentner et al. 2017
superclass	Hexapoda	Latreille 1825
—	Cercofilata

Rank	Name	Author
class	Insecta
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Holometabola
—	Aparaglossata
—	Panorpida
superorder	Antliophora
order	Diptera
—	Neodiptera	Michelsen 1994
infraorder	Bibionomorpha
superfamily	Sciaroidea
family	Keroplatidae	Rondani 1856
family	Keroplatidae	Rondani 1856

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Keroplatidae Rondani 1856 [predatory fungus gnat]

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Subfm. †Adamacrocerinae Ševčík et al. 2020

G. †Adamacrocera Ševčík et al. 2020

†Adamacrocera adami Ševčík et al. 2020

Subfm. Keroplatinae Rondani 1856

G. Antlemon Loew 1871

G. Asindulum Latreille 1805

†Asindulum curvipalpe Meunier 1904

†Asindulum elegantulum Meunier 1904

†Asindulum girschneri Meunier 1904

†Asindulum longipalpe Meunier 1904

†Asindulum pygmaeum Statz 1944

G. Keroplatus Bosc 1792

†Keroplatus major Meunier 1904

G. Orfelia Costa 1857

G. Proceroplatus Edwards 1925

†Proceroplatus hennigi Schmalfuss 1979

†Proceroplatus preziosii Evenhuis and Penney 2013

Invalid names: Keroplatini Rondani 1856 [invalid subgroup], Orfeliini Matile 1990 [invalid subgroup]

Subfm. Lygistorrhininae Edwards 1925 [long beaked fungus gnat]

G. †Archaeognoriste Blagoderov and Grimaldi 2004

†Archaeognoriste primitiva Blagoderov and Grimaldi 2004

G. †Indorrhina Stebner and Grimaldi 2017

†Indorrhina sahnii Stebner and Grimaldi 2017

G. †Lebanognoriste Blagoderov and Grimaldi 2004

†Lebanognoriste prima Blagoderov and Grimaldi 2004

G. †Leptognoriste Blagoderov and Grimaldi 2004

†Leptognoriste davisi Blagoderov and Grimaldi 2004

†Leptognoriste microstoma Blagoderov and Grimaldi 2004

G. Lygistorrhina Skuse 1890

†Lygistorrhina caribbiana Grund 2012

†Lygistorrhina indica Stebner and Grimaldi 2017

G. †Palaeognoriste Meunier 1904

†Palaeognoriste affine Meunier 1912

†Palaeognoriste affinis Meunier 1912

†Palaeognoriste orientale Stebner and Grimaldi 2017

†Palaeognoriste sciariforme Meunier 1904

G. †Parisognoriste Blagoderov et al. 2010

†Parisognoriste eocenica Blagoderov et al. 2010

G. †Plesiognoriste Blagoderov and Grimaldi 2004

†Plesiognoriste carpenteri Blagoderov and Grimaldi 2004

†Plesiognoriste zherikhini Blagoderov and Grimaldi 2004

G. †Protognoriste Blagoderov and Grimaldi 2004

†Protognoriste amplicauda Blagoderov and Grimaldi 2004

†Protognoriste goeleti Blagoderov and Grimaldi 2004

†Protognoriste nascifoa Blagoderov and Grimaldi 2004

Subfm. Macrocerinae Rondani 1856

G. Burmacrocera Cockerell 1917

†Burmacrocera petiolata Cockerell 1917

G. †Eomacroceritis Hong 2002

†Eomacroceritis melanopoda Hong 1974

Tr. Macrocerini Rondani 1856

G. †Hegalari Blagoderov and Arillo 2002

†Hegalari antzinako Blagoderov and Arillo 2002

†Hegalari minor Blagoderov and Arillo 2002

G. Hesperodes Coquillet 1900

†Hesperodes concinna Meunier 1917

Invalid names: Archaemacrocera Meunier 1917 [synonym]

G. Macrocera Meigen 1803

†Macrocera apithanos Kerr and Greenwalt 2022

†Macrocera archaica Armbruster 1938

†Macrocera electricornis Evenhuis 2006

Invalid names: Macrocera longicornis Meunier 1904 [replaced]

†Macrocera obliqua Cockerell 1921

†Macrocera umbonata Statz 1944

Invalid names: Macrocera grandis Meunier 1899 [nomen nudum], Macrocera minuta Meunier 1899 [nomen nudum], Promacrocera Armbruster 1938 [synonym]

G. †Schlueterimyia Matile 1981

†Schlueterimyia cenomanica Matile 1981

Tr. Robsonomyiini Matile 1990

G. †Kelneria Matile 1979

†Kelneria abundare Meunier 1904

†Kelneria ciliata Meunier 1904

†Kelneria filiformis Meunier 1904

†Kelneria setosa Matile 1979

G. Micrepimera Matile 1990

†Micrepimera elegantissima Meunier 1904

†Micrepimera neli Blagoderov and Skibińska 2019

G. Robsonomyia Matile and Vockeroth 1980

†Robsonomyia baltica Pełczyńska et al. 2024

†Robsonomyia henningseni Pełczyńska et al. 2024

Invalid names: Macrocera soccata Meunier 1899 [nomen dubium]

G. Paleoplatyura Meunier 1899

†Palaeoplatyura eocenica Cockerell 1921

†Palaeoplatyura loewi Meunier 1922

†Palaeoplatyura macrocera Meunier 1899

†Paleoplatyura agnieszkae Ševčík et al. 2021

†Paleoplatyura macrocera Meunier 1899

†Paleoplatyura magnifica Ševčík et al. 2021

†Paleoplatyura miae Ševčík et al. 2021

Subfm. Platyurinae Loew 1850

G. Platyura Meigen 1803

†Platyura conjuncta Loew 1850

Invalid names: Platyura difficilis Meunier 1899 [nomen nudum], Platyura filipes Meunier 1899 [nomen nudum], Platyura subaequalis Meunier 1899 [nomen nudum]

G. †Vastaplatyura Solórzano Kraemer and Evenhuis 2008

†Vastaplatyura electrica Solórzano Kraemer and Evenhuis 2008

G. †Vladelektra Evenhuis 2020

†Vladelektra blagoderovi Evenhuis 2020

Invalid names: Platyura armata Meunier 1899 [nomen dubium], Platyura calcar Meunier 1899 [nomen dubium], Platyura ceroplatites Meunier 1904 [nomen dubium], Platyura ceroplatoides Meunier 1904 [nomen dubium], Platyura crassicornis Meunier 1917 [nomen dubium], Platyura distincta Meunier 1904 [nomen dubium], Platyura ectorsii Meunier 1904 [nomen dubium], Platyura ehrhardti Loew 1850 [nomen dubium], Platyura exigua Meunier 1907 [nomen dubium], Platyura graciosa Meunier 1904 [nomen dubium], Platyura kunowi Meunier 1904 [nomen dubium], Platyura mikii Meunier 1904 [nomen dubium], Platyura moniliformis Meunier 1904 [nomen dubium], Platyura pusilla Meunier 1899 [nomen dubium], Platyura verrali Meunier 1904 [nomen dubium]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Late/Upper Berriasian to the top of the Holocene or 139.10000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 137.05 Ma

Collections (45 total)

Time interval	Ma	Country or state	Original ID and collection number
Late/Upper Berriasian	139.1 - 137.05	United Kingdom (England)	Keroplatidae indet. (104003)
Early/Lower Barremian	125.77 - 121.4	Lebanon	Lebanognoriste prima (122359)
Late/Upper Albian	106.3 - 100.5	Spain (Álava)	Hegalari antzinako, Hegalari minor (108769)
Early/Lower Cenomanian	100.5 - 93.9	France	Schlueterimyia cenomanica (123019)
Early/Lower Cenomanian	100.5 - 93.9	Myanmar	Adamacrocera adami (213072) Archaeognoriste primitiva, Protognoriste goeleti, Protognoriste nascifoa, Leptognoriste davisi, Leptognoriste microstoma (111688) Burmacrocera petiolata (123020) Vladelektra blagoderovi (237956)
Late/Upper Cenomanian	100.5 - 93.9	Russian Federation	Protognoriste amplicauda (35051)
Late/Upper Santonian	85.7 - 83.6	Russian Federation	Plesiognoriste zherikhini (35056)
Late/Upper Campanian	83.6 - 72.2	Canada (Manitoba)	Plesiognoriste carpenteri (122354)
Ypresian	56.0 - 48.07	USA (Colorado)	Palaeoplatyura eocenica (105005)
Ypresian	56.0 - 48.07	France (Oise)	Parisognoriste eocenica (123850)
Ypresian	56.0 - 48.07	China (Liaoning)	Macrocera melanopoda (125469) Platyura noda (124530)
Ypresian	56.0 - 48.07	India (Gujarat)	Palaeognoriste orientale, Lygistorrhina indica, Indorrhina sahnii (124463) Vastaplatyura electrica (124464)
Lutetian	48.07 - 41.03	USA (Montana)	Macrocera apithanos (135666 203699)
Priabonian	37.71 - 33.9	Russian Federation (Kaliningrad)	Archaemacrocera concinna (124189) Kelneria setosa (123936) Macrocera abundare (140082) Macrocera abundare, Platyura kunowi, Platyura mikii, Asindulum longipalpis, Ceroplatus major (124073) Palaeognoriste sciariforme, Kelneria sp., Macrocera abundare, Macrocera filiformis, Macrocera ciliata, Kelneria setosa, Macrocera longicornis, Macrocera elegantissima, Platyura ehrhardti, Platyura kunowi, Platyura verrali, Platyura graciosa, Platyura moniliformis, Platyura ectorsii, Platyura mikii, Platyura distincta, Platyura ceroplatoides, Platyura ceroplatites, Platyura conjuncta, Asindulum longipalpis, Asindulum girschneri, Asindulum curvipalpis, Asindulum elegantulus, Palaeognoriste affinis, Platyura crassicornis, Macrocera soccata, Paleoplatyura macrocera, Platyura armata, Platyura calcar, Platyura pusilla, Palaeoplatyura loewi (123927) Palaeoplatyura macrocera (152640) Robsonomyia baltica, Robsonomyia henningseni (234648)
Priabonian	37.71 - 33.9	Germany (Halle)	Antlemon sp. (124273)
Priabonian	37.71 - 33.9	Poland	Micrepimera neli, Paleoplatyura agnieszkae, Paleoplatyura miae, Paleoplatyura magnifica (123215)
Priabonian	37.71 - 33.9	United Kingdom (England)	Platyura obliqua (123959)
Chadronian	37.0 - 33.9	USA (Colorado)	Proapemon infernus (106381)
Chattian	27.3 - 23.04	Germany (North Rhine-Westphalia)	Macrocera umbonata, Asindulum pygmaea (122010)
Early/Lower Miocene	23.04 - 15.98	Mexico	Orfelia sp. 1, Orfelia sp. 2 (21977)
Early/Lower Miocene	23.04 - 15.98	Mexico (Chiapas)	Ceroplatinae indet. (124469 124474 138881 138882 140207 140215 140260 140368 140369)
Burdigalian - Langhian	20.45 - 13.82	Dominican Republic	Proceroplatus hennigi, Lygistorrhina caribbiana (122474) Proceroplatus preziosii (151922)
MN 5	15.98 - 13.82	Germany (Baden-Württemberg)	Promacrocera archaica (127613)
Holocene	0.0117 - 0.0	Tanzania	Platyura exigua (140788)