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Aradoidea

Insecta - Hemiptera

Synonymy list
YearName and author
1836Aradoidea Brullé p. 326
1986Aradoidea Popov p. 81
2006Aradoidea Wappler and Heiss p. 40
2010Aradoidea Cassis and Schuh
2018Aradoidea Johnson et al.
2018Aradoidea Szwedo p. 113

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
RankNameAuthor
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Clareocercaria
—Acercaria()
superorderPancondylognatha
—Condylognatha
orderHemiptera
suborderHeteroptera
infraorderPentatomomorphaLeston et al. 1954
superfamilyAradoidea
superfamilyAradoidea

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Superfm. Aradoidea Brullé 1836
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Fm. Aradidae Brullé 1836 [flat bug]
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Subfm. Aneurinae Douglas and Scott 1865
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G. Aneurus Curtis 1925
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Subg. Aneurus (Aneurodes) Heiss 1998
Subg. Aneurus (Aneurus) Curtis 1925
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Subg. †Aneurus (Neaneurosoma) Heiss 2001
+
Subg. †Aneurus (Paraneurosoma) Heiss 2012
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†Aneurus (Paraneurosoma) damzeni Heiss et al. 2021
†Aneurus incertus Heiss et al. 2015
†Aneurus opertus Zhang et al. 1994
Subfm. Aradinae Brullé 1836
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G. Aradus Fabricius 1803
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†Aradus andancensis Marchal et al. 2011
†Aradus antediluvianus Heer 1853
†Aradus assimilis Germar and Berendt 1856
†Aradus balticus Heiss 2002
†Aradus consimilis Germar and Berendt 1856
†Aradus creticus Kormilev and Popov 1986
†Aradus damzeni Heiss 2002
†Aradus frater Popov 1978
†Aradus frateroides Heiss 1998
†Aradus goellnerae Heiss 2002
†Aradus grabenhorsti Heiss 2013
†Aradus kotashevichi Heiss 2002
†Aradus lativentris Heiss 2002
†Aradus leptosomus Heiss 2014
†Aradus macrosomus Heiss 2014
†Aradus madagascariensis Bervoets 1909
†Aradus nicholasi Popov 1989
†Aradus penteneuros Heiss 2014
†Aradus popovi Heiss 1998
†Aradus rotundiventris Heiss 2014
†Aradus superstes Germar and Berendt 1856
†Aradus velteni Heiss 2002
†Aradus voigti Heiss 2002
†Aradus weitschati Heiss 2002
G. Quilnus Stål 1873
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†Quilnus rectinotus Heiss 2016
G. †Aradoleptus Heiss 2016
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†Aradoleptus birmanus Heiss 2016
G. †Archeaneurus Heiss 2019
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†Archeaneurus neli Heiss 2019
Subfm. †Archearadinae Heiss and Grimaldi 2002
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G. †Archearadus Heiss and Grimaldi 2001
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†Archearadus burmensis Heiss and Grimaldi 2001
†Archearadus elongatus Heiss 2016
G. †Archemezira Heiss and Chen 2023
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†Archemezira nuoxichenae Heiss and Chen 2023
G. †Cretopiesma Grimaldi and Engel 2008
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†Cretopiesma anticum Heiss and Poinar 2012
†Cretopiesma engelgrimaldii Azar et al. 2020
†Cretopiesma inexpectatum Azar et al. 2020
†Cretopiesma lini Azar et al. 2020
†Cretopiesma suukyiae Grimaldi and Engel 2008
Invalid names: Microaradus Heiss and Poinar 2012 [synonym]
G. †Cretozemira Heiss 2023
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†Cretozemira elongata Heiss 2023
†Cretozemira gregori Heiss 2025
G. †Sauronaradus Cumming and Mlynarek 2024
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†Sauronaradus meganae Cumming and Mlynarek 2024
Subfm. Calisiinae Stal 1873
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G. †Allocalisius Heiss 2002
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G. †Archecalisius Heiss 2019
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G. †Calisiomorpha Heiss 2016
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†Calisiomorpha herczeki Heiss 2023
G. Calisiopsis Champion 1898
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†Calisiopsis azteca Heiss 2016
†Calisiopsis brodzinskyorum Froeschner 1992
G. Calisius Stal 1860
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†Calisius balticus Usinger 1941
†Calisius hoffeinsorum Heiss 2002
†Calisius rietscheli Heiss 2000
†Calisius vonholti Heiss 2000
†Calisius weitschati Heiss 2000
Subfm. Carventinae Usinger and Matsuda 1959
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G. Acaricoris Harris and Drake 1948
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†Acaricoris robertae Heiss and Poinar 2012
G. Carventus Stal 1865
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†Carventus bechlyi Heiss 2012
G. Neoproxius Usinger and Matsuda 1959
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†Neoproxius latocanus Heiss and Poinar 2012
G. †Proneoproxius Heiss and Poinar 2012
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†Proneoproxius cornutus Heiss and Poinar 2012
G. †Ellenbergeria Heiss 2016
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G. †Kachinocoris Heiss 2012
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Subfm. Mezirinae Oshanin 1908
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G. †Brevisensoria Poinar 2011
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†Brevisensoria incrustata Poinar 2011
G. Mezira Amyot and Audinet-Serville 1843
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†Mezira crassifemur Wappler and Heiss 2006
†Mezira eckfeldensis Wappler and Heiss 2006
†Mezira eocenica Wappler and Heiss 2006
†Mezira parapetrificata Heiss et al. 2015
†Mezira petrificata Heiss et al. 2015
†Mezira scheveni Heiss 2000
†Mezira succinica Usinger 1941
G. †Myanmezira Heiss and Poinar 2012
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†Myanmezira longicornis Heiss and Poinar 2012
G. Neuroctenus Fieber 1860
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†Neuroctenus enspelensis Wappler and Heiss 2006
†Neuroctenus incompletus Wappler and Heiss 2006
†Neuroctenus kotejai Wappler and Heiss 2006
†Neuroctenus messelensis Wappler and Heiss 2006
G. †Pachytylaradus Heiss 2022
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Subfm. Prosympiestinae Usinger and Matsuda 1959
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G. †Shaykayatcoris Kóbor and Szabó 2025
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†Shaykayatcoris michalskii Kóbor and Szabó 2025
Fm. Termitaphididae Meyers 1924 [termite bug]
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G. Termitaradus Meyers 1924
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†Termitaradus avitinquilinus Grimaldi and Engel 2008
†Termitaradus dominicanus Poinar 2011
†Termitaradus mitnicki Engel 2009
†Termitaradus protera Poinar and Doyen 1992
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Aptian to the top of the Holocene or 121.40000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 113.2 Ma

Collections (51 total)


Time interval Ma Country or state Original ID and collection number
Aptian121.4 - 113.2Mongolia Aradus nicholasi (103657)
Early/Lower Cenomanian100.5 - 93.9Myanmar (Kachin) Myanmezira longicornis, Microaradus anticus (104053)
Early/Lower Cenomanian100.5 - 93.9Myanmar Archemezira nuoxichenae (229441) Cretopiesma anticum, Cretopiesma engelgrimaldii, Cretopiesma lini, Cretopiesma inexpectatum (161726) Cretopiesma suukyiae, Archearadus burmensis (111688) Cretopiesma suukyiae, Microaradus anticus, Calisiomorpha yuripopovi, Ellenbergeria oviventris, Aradoleptus birmanus, Archearadus elongatus, Kachinocoris brevipennis, Archecalisius longiventris, Archeaneurus neli, Cretopiesma anticum, Cretopiesma lini, Cretopiesma inexpectatum, Pachytylaradus cretaceous, Cretozemira elongata, Calisiomorpha herczeki, Cretozemira gregori (153736) Sauronaradus meganae (220289) Shaykayatcoris michalskii (217465)
Early/Lower Campanian83.6 - 72.2Russian Federation (Tenkinskii) Aradus creticus (105375)
Ypresian56.0 - 48.07France (Oise) Calisius balticus (123850)
MP 1148.07 - 41.03Germany (Hessen) Aneurus incertus (173355) Mezira eocenica (125466 144629) Mezira parapetrificata (68475) Mezira petrificata (166830 173356) Neuroctenus kotejai (144627) Neuroctenus messelensis (144628)
MP 1348.07 - 41.03Germany (Eifel) Aradidae indet. (135641) Mezira crassifemur, Aradidae indet. (138912) Neuroctenus incompletus, Aradidae indet. (135643)
Priabonian37.71 - 33.9Lithuania Aradus frateroides (123767)
Priabonian37.71 - 33.9Germany (Halle) Aneurus goitschenus (169965) Aradus grabenhorsti (147606) Calisius balticus (124273 149475 169966)
Priabonian37.71 - 33.9Russian Federation (Kaliningrad) Aradus damzeni, Calisius weitschati (123764) Aradus goellnerae, Aradus lativentris (123221) Aradus grabenhorsti (123770) Calisius balticus, Aneurus groehni (110735) Calisius balticus, Mezira succinica (124205) Calisius sp., Calisius balticus, Aneurus kotashevichi, Aneurus ancestralis, Allocalisius spiniventris, Calisius vonholti, Aradus weitschati, Aradus kotashevichi, Aradus velteni, Aradus voigti, Aradus balticus, Aradus frateroides, Calisius rietscheli, Aradus grabenhorsti, Aradus macrosomus, Aneurus ursulae, Quilnus rectinotus, Aradus leptosomus, Aradus rotundiventris, Aradus penteneuros, Aneurus damzeni (123862)
Priabonian37.71 - 33.9Russian Federation Allocalisius spiniventris, Calisius hoffeinsorum, Aradus kotashevichi (123419)
Priabonian37.71 - 33.9Poland Aradus frater, Aradus popovi (120813) Aradus superstes, Aradus assimilis, Aradus consimilis, Aradidae larva indet. (109625)
MP 2827.3 - 23.04Germany (Rheinland-Pfalz) Neuroctenus enspelensis (160097)
Aquitanian23.04 - 20.45New Zealand Aneurus sp. (50035)
Early/Lower Miocene23.04 - 15.98Mexico (Chiapas) Calisiopsis azteca (178678) Termitaradus protera (125757)
Early/Lower Miocene - Middle Miocene23.04 - 11.63Thailand Neuroctenus sp. (160098)
Burdigalian20.45 - 15.98China (Shandong) Aneurus opertus (138946)
Burdigalian - Langhian20.45 - 13.82Dominican Republic Calisiopsis brodzinskyorum (122473) Carventus bechlyi (122474) Mezira scheveni (124229) Termitaradus avitinquilinus (122982) Termitaradus dominicanus, Brevisensoria incrustata, Neoproxius latocanus, Acaricoris robertae, Proneoproxius cornutus (122472) Termitaradus mitnicki (123708)
Sarmatian12.8 - 11.63Croatia Aradus antediluvianus (117246)
Turolian8.725 - 5.09France (Ardèche) Aradus andancensis (113893)
Holocene0.0117 - 0.0Madagascar Aradus madagascariensis (151372)