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Spiloblattinidae

Insecta - Spiloblattinidae

Taxonomy

Spiloblattinidae was named by Handlirsch (1906).

It was reranked as the subfamily Spiloblattininae by Becker-Migdisova (1961).

It was assigned to Blattoidea by Handlirsch (1906); to Archimylacridae by Becker-Migdisova (1961); to Blattariae by Handlirsch (1922), Zimmerman (1962); to Blattodea by Schneider (1983); and to Eoblattodea by Nel et al. (2022), Garrouste et al. (2025).

Synonyms

Poroblattina longinqua was named by Scudder (1895). Its type specimen is Coll Huston 111, a tegmen (fragment), and it is an impression. Its type locality is Willis Creek near Steubenville, which is in a Stephanian terrestrial shale in the Conemaugh Formation of Ohio.

It was recombined as Ametroblatta longinqua by Handlirsch (1906), Handlirsch (1906), Handlirsch (1920), Handlirsch (1922); it was synonymized subjectively with Spiloblattina variegata by Schneider (1982); it was considered a nomen dubium by Belahmira et al. (2025).
Etoblattina aperta was named by Scudder (1895). Its type specimen is Coll Lacoe 2066 ab, a tegmen (fragment), and it is an impression. Its type locality is Cassville, which is in an Asselian terrestrial shale in the Cassville Shales Formation of West Virginia.

It was considered a nomen dubium by Handlirsch (1906), Handlirsch (1922).
Anthracoblattina wagneri was named by Kliver (1886). It is an impression. Its type locality is Löbejün, which is in a Stephanian C terrestrial shale in the Ottweiler Stufe Formation of Germany.

It was considered a nomen dubium by Handlirsch (1906), Handlirsch (1922).
Anthracoblattina triassica was named by Scudder (1890). It is an impression. Its type locality is Fairplay, which is in an Asselian terrestrial siliciclastic in the Maroon Formation of Colorado.

It was considered a nomen dubium by Handlirsch (1906), Handlirsch (1922).
Spiloblattina laxa was named by Sellards (1908). Its type specimen is KU 171, a tegmen, and it is an impression. Its type locality is Haverkampf farm , which is in a Kasimovian estuary/bay shale in the Lawrence Shale Formation of Kansas.

It was considered a nomen dubium by Handlirsch (1922).

Synonymy list

Year	Name and author
1886	Anthracoblattina wagneri Kliver p. 114 figs. Pl 14, fig 12
1890	Anthracoblattina triassica Scudder p. 464 figs. Pl 41, fig 9
1895	Etoblattina aperta Scudder pp. 80-81 figs. Pl V fig 9
1895	Poroblattina longinqua Scudder pp. 135-136 figs. Pl 11 fig 12
1906	Spiloblattinidae Handlirsch p. 240
1906	Ametroblatta longinqua Handlirsch p. 256 figs. Pl XXVII fig 2
1906	Spiloblattinidae Handlirsch p. 751
1906	Ametroblatta longinqua Handlirsch p. 764
1908	Spiloblattina laxa Sellards p. 523
1920	Ametroblatta longinqua Handlirsch p. 453
1922	Spiloblattinidae Handlirsch
1922	Ametroblatta longinqua Handlirsch p. 171
1961	Spiloblattininae Becker-Migdisova p. 147
1962	Spiloblattinidae Zimmerman p. 1328
1983	Spiloblattinidae Schneider p. 129
2022	Spiloblattinidae Nel et al. p. 3
2025	Spiloblattinidae Garrouste et al. p. 1640

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Gravenhorst 1843
—	Deuteropoda
—	Mandibulata
—	Pancrustacea	Zrzavý and Štys 1997
—	Altocrustacea	Regier et al. 2010
superclass	Allotriocarida
—	Athalassocarida
—	Labiocarida	Schwentner et al. 2017

Rank	Name	Author
superclass	Hexapoda	Latreille 1825
—	Cercofilata
class	Insecta
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Polyneoptera	Martynov 1938
—	Holopandictyoptera
—	Eoblattodea	()
family	Spiloblattinidae
family	Spiloblattinidae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Spiloblattinidae Handlirsch 1906

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G. †Ametroblatta Handlirsch 1906

†Ametroblatta strigosa Scudder 1890

G. †Arrhythmoblatta Handlirsch 1906

†Arrhythmoblatta detecta Scudder 1895

†Arrhythmoblatta scudderiana Handlirsch 1906

G. †Atactoblatta Handlirsch 1906

†Atactoblatta anomala Handlirsch 1906

G. †Dicladoblatta Handlirsch 1906

†Dicladoblatta defossa Scudder 1895

†Dicladoblatta marginata Scudder 1885

G. †Doryblatta Handlirsch 1906

†Doryblatta longipennis Handlirsch 1906

G. †Kinneyblatta Schneider et al. 2021

†Kinneyblatta huberi Schneider et al. 2021

G. †Olethroblatta Handlirsch 1906

†Olethroblatta americana Handlirsch 1906

†Olethroblatta lineolata Cockerell 1927

†Olethroblatta minuta Guthorl 1930

†Olethroblatta tomiensis Becker-Migdisova 1961

Subfm. †Permoblattininae Vishnyakova 1993

G. †Permoblattina Tillyard 1937

†Permoblattina curta Sellards 1908

Invalid names: Etoblattina pecta Sellards 1908 [synonym]

†Permoblattina permiana Sellards 1908

G. †Pycnoblattina Sellards 1908

†Pycnoblattina valeriea Garrouste et al. 2025

G. †Sysciolata Guthorl 1933

†Sysciolata vorkutensis Vishnyakova 1993

G. †Spiloblattina Scudder 1885

†Spiloblattina allegheniensis Carpenter 1934

†Spiloblattina homigtalensis Schneider and Werneburg 1993

†Spiloblattina lawrenceana Handlirsch 1906

Invalid names: Spiloblattina curvata Sellards 1908 [synonym], Spiloblattina lineata Sellards 1908 [synonym], Sysciophlebia sellardsi Handlirsch 1906 [synonym]

†Spiloblattina odernheimensis Schneider 1982

†Spiloblattina pygmaea Meunier 1904

Invalid names: Sysciophlebia lawrenceana Handlirsch 1906 [synonym]

†Spiloblattina sperbersbachensis Schneider and Werneburg 1993

†Spiloblattina steinbachi Guthörl 1934

†Spiloblattina variegata Scudder 1890

Invalid names: Etoblattina exsensa Scudder 1895 [synonym], Etoblattina fasciata Scudder 1890 [synonym], Etoblattina funesta Scudder 1895 [synonym], Etoblattina gracilenta Scudder 1895 [synonym], Etoblattina hastata Scudder 1895 [synonym], Etoblattina hustoni Scudder 1890 [synonym], Etoblattina maledicta Scudder 1895 [synonym], Etoblattina marginata Scudder 1890 [synonym], Etoblattina ramosa Scudder 1895 [synonym], Etoblattina stipata Scudder 1890 [synonym], Etoblattina tenuis Scudder 1890 [synonym], Etoblattina willsiana Scudder 1895 [synonym], Gerablattina apicalis Scudder 1895 [synonym]

†Spiloblattina weissigensis Geinitz 1873

Invalid names: Spiloblattina gardineri Scudder 1885 [synonym], Spiloblattina guttata Scudder 1885 [synonym], Spiloblattina perforata Handlirsch 1906 [synonym], Spiloblattina triassica Scudder 1885 [synonym], Sysciophlebia fenestrata Handlirsch 1906 [synonym]

†Spiloblattinidae gardinerana Handlirsch 1906

†Spiloblattinidae zinkeniana Schlechtendal 1906

G. †Syscioblatta Handlirsch 1906

†Syscioblatta allegheniensis Carpenter 1934

†Syscioblatta anomala Handlirsch 1906

†Syscioblatta dohrni Scudder 1879

Invalid names: Sysciophlebia steinbachi Guthörl 1934 [synonym]

†Syscioblatta euryptera Vishnyakova 1993

†Syscioblatta kaierkanensis Vishnyakova 1993

†Syscioblatta minor Handlirsch 1906

†Syscioblatta misera Handlirsch 1906

†Syscioblatta obscura Handlirsch 1906

†Syscioblatta pennsylvanica Carpenter 1934

†Syscioblatta slusiensis Schneider and Werneburg 1993

†Syscioblatta steubenvilleana Handlirsch 1906

†Syscioblatta teberdensis Vishnyakova 1993

G. †Sysciophlebia Handlirsch 1906

†Sysciophlebia acutipennis Handlirsch 1906

†Sysciophlebia adumbrata Handlirsch 1906

†Sysciophlebia affinis Handlirsch 1906

†Sysciophlebia alligans Schneider 1982

†Sysciophlebia angusta Langiaux and Parriat 1975

†Sysciophlebia arcuata Sellards 1904

†Sysciophlebia balteata Scudder 1879

Invalid names: Etoblattina occulta Scudder 1895 [synonym], Gerablattina cassvici Scudder 1895 [synonym], Gerablattina diversinervis Scudder 1895 [synonym], Sysciophlebia diversipennis Scudder 1895 [synonym], Sysciophlebia frankei Schlechtendal 1906 [synonym]

†Sysciophlebia benedicta Scudder 1895

†Sysciophlebia euglyptica Germar 1851

Invalid names: Dicladoblatta limbata Schlechtendal 1906 [synonym], Gerablattina producta Scudder 1879 [synonym], Gerablattina weissiana Goldenberg 1869 [synonym], Sysciophlebia agilis Schlechtendal 1906 [synonym], Sysciophlebia deperdita Schlechtendal 1906 [synonym], Sysciophlebia elegantissima Schlechtendal 1906 [synonym], Sysciophlebia elongata Schlechtendal 1906 [synonym], Sysciophlebia huysseni Schlechtendal 1906 [synonym], Sysciophlebia ignota Schlechtendal 1906 [synonym], Sysciophlebia laspeyresiana Schlechtendal 1906 [synonym], Sysciophlebia martiusana Schlechtendal 1906 [synonym], Sysciophlebia schlechtendali Handlirsch 1920 [synonym]

†Sysciophlebia grata Schneider 1982

†Sysciophlebia hercynica Scharf 1924

†Sysciophlebia hybrida Handlirsch 1906

†Sysciophlebia ilfeldensis Schlechtendal 1906

†Sysciophlebia intermedia Goldenberg 1877

†Sysciophlebia invisa Scudder 1895

†Sysciophlebia kovacsae Riek 1976

†Sysciophlebia nana Handlirsch 1906

†Sysciophlebia obtusa Handlirsch 1906

†Sysciophlebia oligoneura Handlirsch 1906

†Sysciophlebia patiens Scudder 1895

†Sysciophlebia picta Handlirsch 1906

†Sysciophlebia praepilata Meunier 1921

Invalid names: Sysciophlebia expansa Bolton 1925 [synonym]

†Sysciophlebia recidiva Scudder 1895

†Sysciophlebia rotundata Handlirsch 1906

†Sysciophlebia rubida Schneider 1982

†Sysciophlebia schucherti Handlirsch 1906

†Sysciophlebia scudderi Handlirsch 1906

†Sysciophlebia stulta Schlechtendal 1906

†Sysciophlebia subtilis Schlechtendal 1906

Invalid names: Berlichia wettinensis Schlechtendal 1906 [synonym], Sysciophlebia angustipennis Schlechtendal 1906 [synonym]

†Sysciophlebia variegata Scudder 1890

†Sysciophlebia whitei Handlirsch 1906

Subfm. †Vivablattinae Vršanský and Mostovsky 2000

G. †Vivablatta Vršanský and Mostovsky 2000

†Vivablatta orenburgensis Vishnyakova 1993

†Vivablatta pectinacea Vishnyakova 1993

Invalid names: Kargalia Vishnyakova 1993 [replaced]

Invalid names: Kargaliinae Vishnyakova 1993 [replaced]

G. †Zavjaloblatta Becker-Migdisova 1961

†Zavjaloblatta rotundata Becker-Migdisova 1961

Invalid names: Anthracoblattina triassica Scudder 1890 [nomen dubium], Anthracoblattina wagneri Kliver 1886 [nomen dubium], Etoblattina aperta Scudder 1895 [nomen dubium], Poroblattina longinqua Scudder 1895 [nomen dubium], Spiloblattina laxa Sellards 1908 [nomen dubium]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Bush and Bambach 2015, Kiessling 2004

Age range: base of the Westphalian C to the top of the Capitanian or 314.60000 to 259.51000 Ma

Collections (68 total)

Time interval	Ma	Country or state	Original ID and collection number
Moscovian	315.2 - 307.0	USA (Maryland)	Olethroblatta lineolata (124496)
Westphalian C - Westphalian D	314.6 - 307.4	USA (Pennsylvania)	Olethroblatta americana (114456)
Stephanian	307.4 - 300.3	Germany (Rheinland-Pfalz)	Blattina weissiana (122603)
Stephanian	307.4 - 300.3	Germany	Etoblattina pygmaea (125762) Sysciophlebia sp. (138004) Sysciophlebia steinbachi (155778)
Stephanian	307.4 - 300.3	USA (Ohio)	Etoblattina maledicta, Etoblattina benedicta, Sysciophlebia whitei, Sysciophlebia scudderi, Sysciophlebia hybrida, Sysciophlebia affinis, Etoblattina ramosa, Etoblattina variegata, Sysciophlebia schucherti, Sysciophlebia picta, Sysciophlebia adumbrata, Etoblattina funesta, Sysciophlebia rotundata, Sysciophlebia nana, Sysciophlebia obtusa, Sysciophlebia acutipennis, Etoblattina hastata, Etoblattina fasciata, Etoblattina marginata, Gerablattina apicalis, Etoblattina tenuis, Etoblattina willsiana, Etoblattina exsensa, Syscioblatta obscura, Syscioblatta anomala, Syscioblatta minor, Etoblattina hustoni, Etoblattina gracilenta, Syscioblatta steubenvilleana, Syscioblatta misera, Etoblattina strigosa, Poroblattina longinqua, Atactoblatta anomala, Doryblatta longipennis, Etoblattina stipata (123812) Gerablattina balteata (122605)
Stephanian	307.4 - 300.3	Germany (Saarland)	Blattina intermedia (122600)
Kasimovian	307.0 - 303.7	USA (Kansas)	Gerablattina arcuata, Spiloblattina lineata, Spiloblattina curvata, Spiloblattina laxa, Sysciophlebia lawrenceana, Sysciophlebia sellardsii, Spiloblattina maledicta (113003) Spiloblattina maledicta (147045)
Kasimovian - Gzhelian	307.0 - 298.9	Russian Federation	Syscioblatta teberdensis (126062)
Gzhelian	303.7 - 298.9	Russian Federation	Zavjaloblatta rotundata (124583)
Gzhelian	303.7 - 298.9	Spain	Sysciophlebia sp. (227128)
Gzhelian	303.7 - 298.9	France	Etoblattina praepilata (121979) Sysciophlebia expansa (124916)
Stephanian B	301.4 - 300.9	Czechia	Sysciophlebia grata (156567)
Stephanian B	301.4 - 300.9	France	Sysciophlebia angusta (172760)
Stephanian B	301.4 - 300.9	Morocco	Spiloblattina pygmaea (155288 155290 155291) Sysciophlebia grata (155293)
Stephanian C	300.9 - 300.3	Germany	Berlichia wettinensis (172340)
Stephanian C	300.9 - 300.3	USA (Pennsylvania)	Syscioblatta allegheniensis (144528) Syscioblatta pennsylvanica (155383)
Stephanian C	300.9 - 300.3	Germany (Saxony-Anhalt)	Blattina euglyptica (153702) Spiloblattinidae indet., Blattina euglyptica, Etoblattina dohrnii, Gerablattina producta, Berlichia wettinensis, Sysciophlebia elongata, Sysciophlebia angustipennis, Sysciophlebia deperdita, Sysciophlebia agilis, Sysciophlebia elegantissima, Sysciophlebia stulta, Sysciophlebia ignota, Dicladoblatta subtilis, Dicladoblatta limbata, S. zinkeniana (122578) Syscioblatta dohrni, Sysciophlebia subtilis (125736) Sysciophlebia huysseni, Sysciophlebia agilis, Sysciophlebia oligoneura, Sysciophlebia laspeyresiana, Syscioblatta dohrni, Anthracoblattina wagneri, Sysciophlebia sp. 3, Sysciophlebia sp. 2 (122579) Sysciophlebia sp., Sysciophlebia martiusana (125761)
Stephanian C	300.9 - 300.3	Czechia (Hradec Králové)	Spiloblattina lawrenceana, Sysciophlebia rubida (156568)
Asselian	298.9 - 293.52	USA (Colorado)	Spiloblattina gardineri, Spiloblattina triassica, Spiloblattina guttata, Spiloblattina marginata, Sysciophlebia fenestrata, Anthracoblattina triassica, S. gardinerana, Spiloblattina perforata (106345)
Asselian	298.9 - 293.52	Germany	Spiloblattina odernheimensis (156564 156565) Sysciophlebia balteata (192084)
Asselian	298.9 - 293.52	Germany (Thuringia)	Sysciophlebia hercynica (155777) Sysciophlebia ilfeldensis (128321)
Asselian	298.9 - 293.52	USA (West Virginia)	Gerablattina balteata, Etoblattina patiens, Etoblattina detecta, Etoblattina invisa, Etoblattina occulta, Etoblattina defossa, Etoblattina recidiva, Gerablattina diversinervis, Gerablattina cassvici, Sysciophlebia balteata, Arrhythmoblatta scudderiana, Etoblattina fasciata (122604)
Asselian	298.9 - 293.52	Austria	Sysciophlebia sp 1, Sysciophlebia sp 2, Sysciophlebia sp 3 (146278)
Asselian	298.9 - 293.52	Germany (Rhineland-Palatinate)	Sysciophlebia sp. B (156570 156571)
Asselian	298.9 - 293.52	Germany (Thüringer Wald)	Spiloblattina weissigensis (128478) Sysciophlebia balteata (167765 167768)
Asselian	298.9 - 293.52	Czechia	Spiloblattina weissigensis (114141) Sysciophlebia alligans (133680 133681 156569)
Asselian - Sakmarian	298.9 - 290.1	Russian Federation (Krasnoyarsk)	Syscioblatta euryptera (126061) Syscioblatta kaierkanensis (126063)
Asselian - Sakmarian	298.9 - 290.1	Germany (Rhineland-Palatinate)	Olethroblatta minuta (122606)
Asselian - Sakmarian	298.9 - 290.1	Germany	Sysciophlebia frankei, Sysciophlebia balteata (127934)
Sakmarian	293.52 - 290.1	Germany	Blattina weissigensis, Blattina mahri (122580) Spiloblattina odernheimensis (156566)
Sakmarian	293.52 - 290.1	Germany (Thüringer Wald)	Sysciophlebia balteata (167766) Sysciophlebia balteata, Spiloblattina weissigensis (160734) Sysciophlebia balteata, Syscioblatta slusiensis (167767)
Artinskian	290.1 - 283.3	USA (Kansas)	Etoblattina curta, Etoblattina pecta, Etoblattina permiana (21979) Spiloblattinidae indet., Permoblattina curta, Pycnoblattina sp., Permoblattina permiana (111219)
Artinskian	290.1 - 283.3	Russian Federation	Olethroblatta tomiensis (124594)
Artinskian	290.1 - 283.3	USA (Oklahoma)	Permoblattina curta (104826)
Artinskian - Wordian	290.1 - 264.28	France (Languedoc)	Pycnoblattina valeriea (192423)
Kungurian	283.3 - 274.4	Russian Federation	Spiloblattinidae indet. (102504)
Kungurian	283.3 - 274.4	South Africa (Gauteng)	Sysciophlebia kovacsae (122863)
Kungurian	283.3 - 274.4	Russian Federation (Komi Republic)	Sysciolata vorkutensis (126064)
Roadian	274.4 - 266.9	Russian Federation (Kirov)	Kitjakia longifurcata (105058)
Guadalupian	274.4 - 259.51	Spain	Spiloblattinidae indet. (241076)
Capitanian	264.28 - 259.51	Russian Federation (Orenburg)	Kargalia orenburgensis, Kargalia pectinacea (104057)