PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Mirinae

Insecta - Hemiptera - Miridae

Taxonomy

Mirinae was named by Hahn (1833). It is extant.

It was assigned to Miridae by Statz and Wagner (1950), Popov (1968), Popov and Herczek (1998), Herczek and Popov (2009), Kim and Jung (2019), Cazorla (2021), Kim et al. (2021).

Subtaxa

Armanomirini, Cretamirini, Herdoniini, Mecistoscelini, Mirini (syn. Capsini), Phytocoris consobrinus, Phytocoris involutus, Scutelliferini

Synonyms

Phytocoris consobrinus was named by Germar and Berendt (1856). Its type specimen is MfN Inv. 7297, an exoskeleton, and it is an inclusion in amber. Its type locality is Baltic Amber, Berendt collection, which is in a Priabonian terrestrial amber in Poland.

It was considered a nomen dubium by Herczek and Popov (1997).
Phytocoris involutus was named by Germar and Berendt (1856). Its type specimen is MfN Inv. 7304, an exoskeleton, and it is an inclusion in amber. Its type locality is Baltic Amber, Berendt collection, which is in a Priabonian terrestrial amber in Poland.

It was considered a nomen dubium by Herczek and Popov (1997).

Synonymy list

Year	Name and author
1833	Mirinae Hahn
1856	Phytocoris consobrinus Germar and Berendt p. 25 figs. Pl 3, fig 2
1856	Phytocoris involutus Germar and Berendt p. 28 figs. Pl 3, fig 9
1950	Mirinae Statz and Wagner p. 104
1968	Mirinae Popov p. 107
1998	Mirinae Popov and Herczek p. 41
2009	Mirinae Herczek and Popov p. 94
2019	Mirinae Kim and Jung
2021	Mirinae Cazorla p. 33
2021	Mirinae Kim et al. p. 572

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Gravenhorst 1843
—	Deuteropoda
—	Mandibulata
—	Pancrustacea	Zrzavý and Štys 1997
—	Altocrustacea	Regier et al. 2010
superclass	Allotriocarida
—	Athalassocarida
—	Labiocarida	Schwentner et al. 2017
superclass	Hexapoda	Latreille 1825
—	Cercofilata
class	Insecta

Rank	Name	Author
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Clareocercaria
—	Acercaria	()
superorder	Pancondylognatha
—	Condylognatha
order	Hemiptera
suborder	Heteroptera
infraorder	Cimicomorpha	Leston et al. 1954
superfamily	Miroidea
family	Miridae	()
subfamily	Mirinae
subfamily	Mirinae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. Mirinae Hahn 1833

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Tr. †Armanomirini Popov and Herczek 1998

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G. †Armanomiris Popov and Herczek 1998

†Armanomiris cretaceous Popov and Herczek 1998

†Armanomiris klimaszewskii Popov and Herczek 1998

Tr. †Cretamirini Popov and Herczek 1998

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G. †Cretamiris Popov and Herczek 1998

†Cretamiris zherikhini Popov and Herczek 1998

Tr. Herdoniini Distant 1904

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G. Closterocoris Uhler 1890

†Closterocoris elegans Scudder 1890

Tr. Mecistoscelini Reuter 1910

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G. †Cretamystilus Kim and Jung 2021

†Cretamystilus herczeki Kim and Jung 2021

Tr. Mirini Hahn 1833

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G. †Baltiomiris Kim et al. 2021

†Baltiomiris herczeki Kim et al. 2021

G. Calocoris Fieber 1858

G. Capsus Fabricius 1803

†Capsus lacus Scudder 1890

†Capsus obsolefactus Scudder 1890

G. Lygus Hahn 1831

†Lygus oligocaenica Statz and Wagner 1950

G. Mermitelocerus Reuter 1908

†Mermitelocerus antennatus Statz and Wagner 1950

G. †Mixocapsus Herczek 1991

†Mixocapsus eocenicus Herczek 1991

G. Phytocoris Fallen 1814

†Phytocoris balticus Germar and Berendt 1856

†Phytocoris euglotta Germar and Berendt 1856

†Phytocoris merus Germar and Berendt 1856

†Phytocoris poissoni Theobald 1937

†Phytocoris sendelii Germar and Berendt 1856

G. Poecilocapsus Reuter 1876

†Poecilocapsus fremontii Scudder 1890

†Poecilocapsus ostentus Scudder 1890

†Poecilocapsus tabidus Scudder 1890

†Poecilocapsus veterandus Scudder 1890

†Poecilocapsus veternosus Scudder 1890

G. †Stomatomiris Maldonado Capriles and Poinar 1995

†Stomatomiris funebris Maldonado Capriles and Poinar 1995

G. †Tschirnhausia Herczek and Popov 2011

†Tschirnhausia sambiensis Herczek and Popov 2009

Invalid names: Stenopterna Herczek and Popov 2009 [replaced]

Invalid names: Capsini Burmeister 1835 [synonym]

Tr. †Scutelliferini Popov 1968

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G. †Scutellifer Popov 1968

†Scutellifer karatavicus Popov 1968

Invalid names: Phytocoris consobrinus Germar and Berendt 1856 [nomen dubium], Phytocoris involutus Germar and Berendt 1856 [nomen dubium]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Callovian to the top of the Langhian or 165.30000 to 13.82000 Ma
Minimum age of oldest fossil (stem group age): 154.8 Ma

Collections (13 total)

Time interval	Ma	Country or state	Original ID and collection number
Callovian - Oxfordian	165.3 - 154.8	Kazakhstan (Chimkent)	Scutellifer karatavicus (105971)
Early/Lower Cenomanian	100.5 - 93.9	Myanmar	Cretamystilus herczeki (219180)
Early/Lower Campanian	83.6 - 72.2	Russian Federation (Tenkinskii)	Armanomiris klimaszewskii, Armanomiris cretaceous, Cretamiris zherikhini (105375)
Late/Upper Eocene	37.71 - 33.9	USA (Colorado)	Poecilocapsus fremontii (122093)
Priabonian	37.71 - 33.9	Poland	Mirinae indet., Lygus sp. (120813) Phytocoris larva sp., Phytocoris sendelii, Phytocoris euglotta, Phytocoris consobrinus, Phytocoris balticus, Phytocoris involutus (109625)
Priabonian	37.71 - 33.9	Russian Federation (Kaliningrad)	Baltiomiris herczeki (224001) Mixocapsus eocenicus (123458) Stenopterna sambiensis (110735)
Chadronian	37.0 - 33.9	USA (Colorado)	Closterocoris elegans, Poecilocapsus fremontii, Poecilocapsus veterandus, Poecilocapsus veternosus, Poecilocapsus tabidus, Poecilocapsus ostentus, Capsus obsolefactus, Capsus lacus (106384)
MP 23	33.9 - 27.3	France	Phytocoris poissoni (122535)
Chattian	27.3 - 23.04	Germany (North Rhine-Westphalia)	Calocoris antennatus, Lygus oligocaenica (122010)
Burdigalian - Langhian	20.45 - 13.82	Dominican Republic	Stomatomiris funebris (122472)