PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Hydrophilini

Insecta - Coleoptera - Hydrophilidae

Taxonomy

Hydrophilini was named by Latreille (1802). It is extant. Its type is Hydrophilus.

It was assigned to Hydrophilidae by Fikáček et al. (2010); and to Hydrophilinae by Bouchard et al. (2011), Short and Fikáček (2013).

Subtaxa

Hydrophilina, Hydrophilopsis, Hydrophilopsis elongata, Hydrous ovalis

Synonyms

Hydrophilopsis was named by Heer (1862).

It was considered a nomen dubium by Fikáček and Schmied (2013).

It was assigned to Hydrophilidae by Carpenter (1992).
Coccinella protogaeae was named by Germar (1837). Its type specimen is GPIBO A-598, an exoskeleton, and it is an impression. Its type locality is Orsberg (Goldfuss Bonn collection), which is in a MP 30 lacustrine - large shale in the Rott Formation of Germany.

It was recombined as Escheria protogaeae by Heer (1847); it was misspelled as Escheria protogeae by Handlirsch (1908) and Statz (1939); it was considered a nomen dubium by Fikáček et al. (2010).
Hydrophilina was named by Latreille (1802). It is extant. Its type is Hydrophilus.

It was considered an invalid subgroup of Hydrophilini by Short and Fikáček (2013).

It was assigned to Hydrophilini by Fikáček et al. (2010), Bouchard et al. (2011).
Hydrophilopsis elongata was named by Heer (1862). Its type specimen is ETHZ NO-Z51.3-39-4:4-Pl.I144, an exoskeleton, and it is a compression fossil. Its type locality is Oeningen (Zurich collection), which is in a Sarmatian crater lake limestone in the Upper Freshwater-Molasse Formation of Germany.

It was considered a nomen dubium by Fikáček and Schmied (2013).
Hydrous ovalis was named by Heer (1862). Its type specimen is ETHZ NO-Z51.3-39-4:4-Pl.I149b, an exoskeleton, and it is a compression fossil. Its type locality is Oeningen (Zurich collection), which is in a Sarmatian crater lake limestone in the Upper Freshwater-Molasse Formation of Germany.

It was considered a nomen dubium by Fikáček and Schmied (2013).

Synonymy list

Year	Name and author
1802	Hydrophilina Latreille p. 136
1802	Hydrophilini Latreille p. 136
1837	Coccinella protogaeae Germar figs. Pl. 15
1847	Escheria protogaeae Heer p. 60
1862	Hydrophilopsis Heer p. 68
1862	Hydrous ovalis Heer p. 68 figs. Pl 6, fig 1
1862	Hydrophilopsis elongata Heer p. 69 figs. Pl 5, figs 18,18b
1864	Hydrophilopsis elongata Heer p. 381 fig. 274
1908	Escheria protogeae Handlirsch p. 767
1939	Escheria protogaea Statz
1992	Hydrophilopsis Carpenter
2010	Hydrophilina Fikáček et al. p. 740
2010	Hydrophilini Fikáček et al. p. 740
2011	Hydrophilini Bouchard et al. p. 155
2011	Hydrophilina Bouchard et al. p. 156
2013	Hydrophilini Short and Fikáček p. 738

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Gravenhorst 1843
—	Deuteropoda
—	Mandibulata
—	Pancrustacea	Zrzavý and Štys 1997
—	Altocrustacea	Regier et al. 2010
superclass	Allotriocarida
—	Athalassocarida
—	Labiocarida	Schwentner et al. 2017
superclass	Hexapoda	Latreille 1825
—	Cercofilata
class	Insecta
—	Dicondylia

Rank	Name	Author
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Holometabola
—	Aparaglossata
—	Neuropteriformia
—	Coleopterida
order	Coleoptera
—	Mesocoleoptera
—	Metacoleoptera
suborder	Polyphaga	Emery 1886
infraorder	Staphyliniformia
superfamily	Hydrophiloidea
family	Hydrophilidae
subfamily	Hydrophilinae	()
tribe	Hydrophilini
tribe	Hydrophilini

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Tr. Hydrophilini Latreille 1802

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Invalid names: Hydrophilina Latreille 1802 [invalid subgroup], Hydrophilopsis Heer 1862 [nomen dubium], Hydrophilopsis elongata Heer 1862 [nomen dubium], Hydrous ovalis Heer 1862 [nomen dubium]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the MP 11 to the top of the Late/Upper Pleistocene or 48.07000 to 0.01170 Ma
Minimum age of oldest fossil (stem group age): 46.2 Ma

Collections (38 total)

Time interval	Ma	Country or state	Original ID and collection number
Bridgerian	50.5 - 46.2	USA (Wyoming)	Tropisternus saxialis, Tropisternus sculptilis (121322)
MP 11	48.07 - 41.03	Germany (Hessen)	Hydrobiomorpha eopalpalis (165522) Hydrophilus sp., Hydrobiomorpha eopalpalis (120036) Hydrophilus sp., Hydrobiomorpha sp. 1, Hydrochara sp., Hydrophilina indet., Hydrobiomorpha eopalpalis (120798)
Priabonian	37.71 - 33.9	United Kingdom (England)	Hydrochara woodwardi (21976 123959 123960)
Chadronian	37.0 - 33.9	USA (Colorado)	Tropisternus limitatus, Tropisternus vanus, Hydrocharis extricatus (106384)
Chattian	27.3 - 23.04	Germany (North Rhine-Westphalia)	Hydrophilus fraternus, Hydrophilus neptunus, Hydrous rottensis (122010)
Chattian	27.3 - 23.04	France (Provence-Alpes-Côte d'Azur)	Hydrophilopsis incerta (122487)
Chattian	27.3 - 23.04	France (Bouches-Du-Rhone)	Hydrophilus antiquus (123944)
Late/Upper Oligocene	27.3 - 23.04	France (Puy-de-Dome)	Hydrophilus magnificus (155204)
MP 30	27.3 - 23.04	Germany	Coccinella protogaeae (115802) Hydrous neptunus (123945)
MP 30	27.3 - 23.04	Germany (Nordrhein-Westfalen)	Hydrophilus fraternus, Hydrous miserandus (127628)
Chattian - Aquitanian	27.3 - 20.45	Germany (Sachsen)	Hydrobiomorpha enspelense (26765)
Aquitanian	23.04 - 20.45	Germany	Hydrous sp., Hydrophilus sp. (123872)
Aquitanian	23.04 - 20.45	Switzerland	Hydrophilus gaudini, Hydrophilus ruminianus (154053)
Burdigalian	20.45 - 15.98	Greece	Hydrophilus vexatorius (144526)
Burdigalian	20.45 - 15.98	China (Shandong)	Hydrous pergrandis (128552)
MN 3	20.0 - 16.9	Czechia	Hydrophilus pistaceus (105508)
Badenian	13.82 - 12.8	Austria (Steiermark)	Hydrophilus carbonarius (115306)
Sarmatian	12.8 - 11.63	Germany (Baden-Württemberg)	Hydrobiomorpha heeri (144245) Hydrophilus rehmanni (117338) Hydrophilus spectabilis (172253) Hydrophilus spectabilis, Hydrous escheri, Hydrobiomorpha heeri, Hydrophilus giganteus, Hydrophilopsis elongata, Hydrous ovalis, Hydrous rehmanni, Hydrous braunii (117252) Hydrophilus vexatorius, Hydrophilus knorrii, Hydrophilus braunii (115305) Hydrophilus vexatorius, Hydrophilus spectabilis, Hydrophilus noachicus (115304)
Sarmatian	12.8 - 11.63	Germany (Wurttemberg)	Hydrophilus kuntzeni, Hydrophilus caraboides (113752)
Pliocene	5.333 - 2.58	Japan	Hydrochara libera (165744)
Piacenzian	3.6 - 2.58	Japan	Hydrophilus sp., Hydrochara sp. (165738)
Calabrian	1.8 - 0.774	Japan	Hydrochara sp. (160539)
Chibanian	0.774 - 0.129	Japan	Hydrophilus sp., Hydrochara sp., Hydrochara libera (165700)
Middle Pleistocene	0.774 - 0.129	United Kingdom	Hydrophilus caraboides (168267)
Late/Upper Pleistocene	0.129 - 0.0117	Russian Federation (Astrakhan)	Hydrochara sp., Hydrochara caraboides, Hydrochara flavipes (125794)
Late/Upper Pleistocene	0.129 - 0.0117	Peru	Hydrophilus sp., Gyrinidae indet. (71346)
Late/Upper Pleistocene	0.129 - 0.0117	USA (California)	Hydrophilus sp. (44954)
Late/Upper Pleistocene	0.129 - 0.0117	Japan	Hydrochara libera (166082)