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Panaeshnida

Insecta - Odonata

Taxonomy
Panaeshnida was named by Bechly et al. (2001). It is extant.

It was assigned to Aeshnomorpha by Bechly et al. (2001), Bechly (2007).

Synonymy list
YearName and author
2001Panaeshnida Bechly et al. p. 66
2007Panaeshnida Bechly

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
infraclassHydropalaeopteraRohdendorf 1969
—Euhydropalaeoptera
—OdonatopteraMartynov 1932
—Palaeodonatoptera
RankNameAuthor
—Plesiodonatoptera
—Apodonatoptera
—Paneodonatoptera
—Neodonatoptera
—Euodonatoptera
—OdonatocladaBechly 1999
—Nodialata
—Discoidalia
—Stigmoptera
—Lobodonata
—Panodonata
orderOdonata
—Epiproctophora
—EuepiproctophoraBechly 1998
—Anisopteromorpha
—Trigonoptera
—Pananisoptera
—NeoanisopteraBechly 1998
suborderAnisopteraSelys-Longchamps and Hagen 1854
—Euanisoptera
—Aeshnoptera
—Aeshnomorpha
—Panaeshnida
—Panaeshnida

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Panaeshnida Bechly et al. 2001
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Unr. Aeshnida Bechly 1996
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Superfm. †Cymatophlebioidea Handlirsch 1906
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Fm. †Cymatophlebiidae Handlirsch 1906
+
Subfm. †Cymatophlebiinae Handlirsch 1906
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G. †Cymatophlebia Deichmüller 1886
Subfm. †Valdaeshninae Bechly et al. 2001
+
G. †Hoyaeshna Nel and Martínez-Delclòs 1993
G. †Prohoyaeshna Bechly et al. 2001
G. †Valdaeshna Jarzembowski 1988
Invalid names: Libellulium Westwood 1854 [nomen dubium], Libellulium agrias Westwood 1854 [nomen dubium]
Fm. †Rudiaeschnidae Bechly et al. 2001
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G. †Fuxiaeschna Lin et al. 2004
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†Fuxiaeschna hsiufunia Lin et al. 2004
G. †Rudiaeschna Ren and Guo 1996
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†Rudiaeschna jarzembowskii Huang et al. 2022
†Rudiaeschna limnobia Ren and Guo 1996
Unr. Paneuaeshnida Bechly et al. 2001
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Unr. Euaeshnida Bechly 1996
+
Fm. †Eumorbaeschnidae Bechly et al. 2001
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G. †Eumorbaeschna Bechly et al. 2001
Unr. Neoaeshnida Bechly 1996
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Unr. Aeshnodea Bechly 1996
Fm. Gomphaeschnidae Tillyard and Fraser 1940
Fm. †Paracymatophlebiidae Bechly et al. 2001
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G. †Linqibinia Pinkert et al. 2017
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†Linqibinia panae Pinkert et al. 2017
G. †Paracymatophlebia Bechly et al. 2001
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†Paracymatophlebia splendida Bechly et al. 2001
G. †Sinacymatophlebia Nel and Huang 2009
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†Sinacymatophlebia mongolica Nel and Huang 2009
Fm. †Progobiaeshnidae Bechly et al. 2001
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G. †Decoraeshna Li et al. 2012
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†Decoraeshna preciosa Li et al. 2012
G. †Gobiaeshna Pritykina 1977
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†Gobiaeshna occulta Pritykina 1977
G. †Mongoliaeshna Nel and Huang 2010
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†Mongoliaeshna exiguusens Li et al. 2012
†Mongoliaeshna hadrens Li et al. 2012
†Mongoliaeshna sinica Nel and Huang 2010
G. †Paradecoraeshna Zheng et al. 2018
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†Paradecoraeshna liaoningensis Zheng et al. 2018
G. †Progobiaeshna Bechly et al. 2001
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†Progobiaeshna liaoningensis Bechly et al. 2001
G. †Telmaeshna Zhang et al. 2008
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†Telmaeshna paradoxica Zhang et al. 2008
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitino
Form: roller-shapedo
Environment: freshwater, terrestrialo
Locomotion: fast-movingo
Life habit: volanto
Diet: carnivoreo
Diet 2: insectivoreo
Vision: well-developedo
Dispersal: watero
Created: 2011-03-07 03:41:10
Modified: 2011-03-06 10:41:10
Source: o = order
Reference: Ren and Guo 1996

Age range

Maximum range based only on fossils: base of the Callovian to the top of the Holocene or 165.30000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 154.8 Ma

Collections (163 total)


Time interval Ma Country or state Original ID and collection number
Callovian - Oxfordian165.3 - 154.8China (Nei Mongol) Sinacymatophlebia mongolica, Linqibinia panae, Cymatophlebia daohugouensis (113371)
Early/Lower Kimmeridgian154.8 - 152.21Germany (Baden-Wurttemberg) Cymatophlebia herrienae (105965) Cymatophlebia suevica (105964)
Kimmeridgian154.8 - 149.2Kazakhstan Paracymatophlebia splendida (105426)
Late/Upper Kimmeridgian152.21 - 149.2Germany (Swabian Alb) Libellulium longialatum (106402)
Early/Lower Tithonian149.2 - 145.06Germany (Bavaria) Aeschna multicellulosa (152880) Cymatophlebia longialata (152603 152884) Cymatophlebia longialata, Cymatophlebia jurassica (139876) Cymatophlebia longialata, Cymatophlebia pumilio, Cymatophlebia kuempeli, Eumorbaeschna jurassica, Morbaeschna muensteri (122586) Cymatophlebia longialata, Eumorbaeschna jurassica (123994)
Early/Lower Tithonian149.2 - 145.06Germany Cymatophlebia longialata (122880) Cymatophlebia longialata, Eumorbaeschna jurassica (105799)
Early/Lower Tithonian149.2 - 145.06Germany (Bayern) Cymatophlebia densa (152881) Cymatophlebia longialata (105797 105798 123144 123906 131959 133578 139548 139550 152882 152901) Cymatophlebia longialata, Cymatophlebia pumilio, Cymatophlebia kuempeli, Eumorbaeschna jurassica (139551) Cymatophlebia longialata, Eumorbaeschna jurassica (122730 152883) Cymatophlebia pumilio (105541 117534) Eumorbaeschna jurassica (139547) Libellula longialata, Eumorbaeschna jurassica (123985)
Early/Lower Tithonian149.2 - 145.06Poland Eumorbaeschna adriankini (140049)
Late/Upper Tithonian - Early/Lower Berriasian145.06 - 141.0Spain Cymatophlebia longialata (224315)
Berriasian143.1 - 137.05United Kingdom (England) Libellulium agrias (127481)
Middle Berriasian141.0 - 139.1United Kingdom (England) Cymatophlebia purbeckensis (153648)
Late/Upper Berriasian139.1 - 137.05United Kingdom (England) Cretalloaeschna cliffordae (103833) Prohoyaeshna milleri (104003)
Valanginian - Early/Lower Hauterivian137.05 - 125.77China (Gansu) Fuxiaeschna hsiufunia (156355)
Hauterivian132.6 - 125.77Algeria Gomphaeschnidae indet. (123148)
Late/Upper Hauterivian132.6 - 125.77United Kingdom (Surrey) Valdaeshna surreyensis (160653)
Late/Upper Hauterivian132.6 - 125.77United Kingdom (England) Valdaeshna andressi, Plesigomphaeschnaoides pindelskii (106281)
Barremian125.77 - 121.4Russian Federation (Chita) Gomphaeschna sibirica (105968)
Early/Lower Barremian125.77 - 121.4United Kingdom (England) Libellulium standingae, Libellulium zdrzaleki (104005) Valdaeshna mikei (45193)
Early/Lower Barremian125.77 - 121.4Spain Palaeaeschna vidali (104052 118362 123437)
Late/Upper Barremian125.77 - 121.4China (Liaoning) Aeschna acrodonta (148368) Paradecoraeshna liaoningensis (111725) Progobiaeshna liaoningensis (105795) Rudiaeschna limnobia (111716 138436) Rudiaeschna limnobia, Sinojagoria imperfecta (105809) Sinojagoria cancellosa, Sinojagoria magna, Decoraeshna preciosa (124759) Sinojagoria magna, Cymatophlebia yixianensis (176229) Telmaeshna paradoxica (111722)
Late/Upper Barremian125.77 - 121.4China (Nei Mongol) Mongoliaeshna sinica, Rudiaeschna jarzembowskii (125655) Sophoaeschna frigida, Falsisophoaeschna generalis, Mongoliaeshna hadrens, Mongoliaeshna exiguusens (125656)
Late/Upper Barremian125.77 - 121.4Spain (Cuenca) Hoyaeshna cretacica (152992)
Aptian121.4 - 113.2Mongolia Gobiaeshna occulta (105796) Gomphaeschna sibirica (103655) Plesigomphaeschnaoides mongolensis (103657)
Aptian121.4 - 113.2Russian Federation (Buryatia) Baissaeshna zherikhini (105969) Gomphaeschna inferna, Baissaeshna prisca (105536)
Late/Upper Aptian119.57 - 113.2Brazil (Ceará) Anomalaeschna berndschusteri, Paramorbaeschna araripensis (122924) Gomphaeschna obliqua (122352) Gomphaeschna obliqua, Gomphaeschnaoides betoreti (123011) Gomphaeschna obliqua, Gomphaeschnaoides magnus, Gomphaeschnaoides petersi (122919) Gomphaeschnaoides magnus (123012) Paramorbaeschna araripensis (122553) Paramorbaeschna araripensis, Gomphaeschna obliqua (122925) Paramorbaeschna araripensis, Gomphaeschna obliqua, Gomphaeschnaoides obliquus, Gomphaeschnaoides magnus (122557) Primumaeshna britta (215116) Progomphaeschnaoides staniczeki, Gomphaeschnaoides magnus, Gomphaeschnaoides petersi (123010) Progomphaeschnaoides ursulae (122556)
Early/Lower Cenomanian100.5 - 93.9Myanmar Cretagomphaeschnaoides jarzembowskae, Kachinaeshna zhuoi (161726)
Selandian61.66 - 59.24France (Auvergne) Aeschnidiidae indet. (113685)
Tiffanian60.9 - 57.5USA (North Dakota) Gomphaeschna schrankii (15000)
Tiffanian60.9 - 57.5Canada (Alberta) Aeshnidae indet., Gomphaeschninae indet., Alloaeschna paskapooensis, Alloaeschna quadrata (119839) Alloaeschna paskapooensis (119828 119832) Alloaeschna paskapooensis, Gomphidae indet. (119816) Alloaeschna sp. (232706)
Thanetian - Ypresian59.24 - 48.07USA (Alaska) Basiaeschna alaskaensis (204785)
Ypresian56.0 - 48.07Denmark Gomphaeschna danica (120023) Gomphaeschna paleocenica (113029) Parabaissaeshna ejerslevense (199521)
Ypresian56.0 - 48.07Canada (British Columbia) Aeshnidae indet. (226231) Ypshna latipennata (147816) Ypshna latipennata, Eoshna thompsonensis (103205)
Ypresian56.0 - 48.07Argentina (Chubut) Huncoaeshna corrugata (75178)
Ypresian56.0 - 48.07USA (Washington) Aeshnidae indet. A, Antiquiala snyderae, Idemlinea versatilis, Ypshna brownleei (103206)
Lutetian48.07 - 41.03USA (Montana) Kishenehna prima (203699)
Lutetian48.07 - 41.03United Kingdom (England) Triaeschna gossi (113860)
MP 1148.07 - 41.03Germany (Hessen) Aeschnidae indet. (120798)
MP 1348.07 - 41.03Germany (Eifel) Gomphaeschnidae indet. (135641)
Priabonian37.71 - 33.9Poland Aeshnidae indet. specimen 1 (123215)
Priabonian37.71 - 33.9China (Xizang) Oligaeschna sinica (224485)
Priabonian37.71 - 33.9United Kingdom (England) Aeschnophlebia andreasi, Anglogomphaeschna eocenica (207258) Anglogomphaeschna eocenica, Gynacanthinae indet. A (123960) Oligoaeschna anglica, Aeschnophlebia andreasi, Anglogomphaeschna eocenica (21976) Oligoaeschna anglica, Aeschnophlebia andreasi, Oligaeschna wedmanni (127743) Oligoaeschna anglica, Oplonaeschna vectensis, Anglogomphaeschna eocenica, Oligaeschna wedmanni (123959)
Priabonian37.71 - 33.9Russian Federation (Kaliningrad) Elektrogomphaeschna annekeae (123936) Elektrogomphaeschna peterthieli (120814)
Chadronian37.0 - 33.9USA (Colorado) Aeschna solida, Aeschna larvata, Aeschna separata (106384) Aeshna larvata (123119) Lithaeschna needhami, Oplonaeschna separata, Aeschna solida (106381) Oplonaeschna lapidaria (106380)
Early/Lower Oligocene33.9 - 27.3Spain Aeshnidae indet., Aeshna sp. (113990)
MP 2133.9 - 27.3France Aeshna caseneuvensis (233239)
MP 2333.9 - 27.3France Aeschna ollivieri (122984)
Rupelian33.9 - 27.3Germany Aeshnidae indet., Aeschna heterofasciata (122407)
Rupelian33.9 - 27.3France (Alpes-de-Haute) Aeschna paleocyanea (114016)
Rupelian33.9 - 27.3Czechia (Moravia) Oligaeschna kvaceki (104941)
Rupelian33.9 - 27.3Czechia Aeshna sp. (181764) Kvacekia infuscata (31643)
Oligocene33.9 - 23.04France Oligaeschna jungi (127946)
Chattian27.3 - 23.04France (Bouches-du-Rhone) Aeschna palaeocoerulea (155869)
Chattian27.3 - 23.04Kazakhstan (Saisan) Epacantha magnifica, Basiaeschna ashutasica, Projagoria conjuncta, Necracantha sp., Necracantha composita (114018) Necracantha proxima (120121)
Chattian27.3 - 23.04France (Bouches-Du-Rhone) Aeshna oligocenica (127415)
Late/Upper Oligocene27.3 - 23.04France (Puy-de-Dome) Oligaeschna jungi (155204)
Late/Upper Oligocene27.3 - 23.04France (Aix-en-Provence) Epiaeschna pseudoheros, Jagoria oligocenica (114117)
MP 2827.3 - 23.04Germany (Rheinland-Pfalz) Oligaeschna jungi, Epiaeschna wisseri (71942)
MP 3027.3 - 23.04Germany Aeschna dido (123945)
Aquitanian23.04 - 20.45Germany Projagoria conjuncta, Aeschnidiidae indet. (123872)
Aquitanian23.04 - 20.45Denmark Oplonaeschna staurophlebioides (164572)
Early/Lower Miocene23.04 - 15.98Spain (Castellon) Aeshnidae indet. (140798) Oligaeschna saurai (122583)
Early/Lower Miocene23.04 - 15.98Azerbaijan Heliaeschna parallela (127434)
Burdigalian20.45 - 15.98Czechia Aeshna zlatkokvaceki (181763)
Burdigalian20.45 - 15.98China (Shandong) Epiaeschna matutina, Aeshna shanwangensis, Aeshna forficatum (160472) Jagoria sinica (128550) Mediaeschna lucida (128552 138944) Mediaeschna lucida, Aeschna ignivora (128551) Mediaeschna matutina (128549)
MN 320.0 - 16.9Czechia Aeshnidae indet. (150934) Merlax bohemicus, Gomphaeschna miocenica (105508)
Langhian15.98 - 13.82USA (Idaho) Gomphaeschninae indet. (130391)
Middle Miocene15.98 - 11.63Russian Federation (Stavropol) Epiaeschna stauropolitana (114118)
Middle Miocene15.98 - 11.63Russian Federation Aeshna stavropolensis (105571)
Middle Miocene15.98 - 11.63Bulgaria Oligaeschna bulgariensis (181730)
Middle Miocene - Late/Upper Miocene15.98 - 5.333China (Yunnan) Jingguaeshna taoae (229451)
Badenian13.82 - 12.8Austria Gomphaeschna carinthiae (184615)
Sarmatian12.8 - 11.63Germany (Baden-Württemberg) Aeschna polydore (115305) Aeschna tyche (115304)
Sarmatian12.8 - 11.63Croatia Aeschna metis (117250)
Sarmatian12.8 - 11.63Germany (Wurttemberg) Aeschnidae indet. (113752)
Vallesian - Turolian11.63 - 5.09Spain Aeshna cerdanica (113995)
Turolian8.725 - 5.09France (Ardèche) Aeshna andancensi, Aeschnophlebia miocenica (113893) Aeshna sp., Aeshna turoliana (156164) Nasiaeschna miocenica (227089)
Messinian7.246 - 5.333Italy Anax sp., Aeshna sp., Anax cryptus, Anax imperator, Anax ephippiger, Anax parthenope, Aeshna messiniana, Aeshna ghiandonii, Aeshna multicellulata (113886)
Messinian7.246 - 5.333Austria Aeschna voesendorfensis (127417)
Messinian7.246 - 5.333France (Cantal) Allopetalia europaea (122195)
Blancan4.7 - 1.4USA (Texas) Aeshnidae indet. (154909)
Late/Upper Pliocene3.6 - 2.58Japan Oligoaeschna pryeri (127452)
Piacenzian3.6 - 2.58Germany (Niedersachsen) Aeschnidae indet. (109196)
Gelasian2.58 - 1.8France Gomphaeschnini indet., Aeschna theobaldi (113996)
Pleistocene2.58 - 0.0117Slovakia Aeshna grandis (127418)
Holocene0.0117 - 0.0France Aeshna isosceles (127419)