PBDB Taxon

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Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Caloneuridae

Insecta - Caloneurodea - Caloneuridae

Taxonomy

Caloneuridae was named by Handlirsch (1906). It is not extant.

It was assigned to Protorthoptera by Handlirsch (1906), Pruvost (1919), Handlirsch (1919), Handlirsch (1922), Bolton (1925); and to Caloneurodea by Martynov (1938), Beckemeyer (2009).

Subtaxa

Amboneura, Anomalogramma, Caloneura (syn. Confusio, Boltonaloneura), Caloneurella, Eohymen, Euthygramma (syn. Paleuthygrammella, Pseudogramma), Isadistica, Ligogramma, Pleisiogramma, Vilvia, Vilviopsis

Synonyms

Euthygrammidae was named by Martynov (1928). It is not extant.

It was synonymized subjectively with Caloneuridae by Rasnitsyn (2004).

It was assigned to Protorthoptera by Martynov (1928); and to Caloneurodea by Martynov (1938), Zimmerman (1962).
Paleuthygrammidae was named by Carpenter (1943).

It was synonymized subjectively with Caloneuridae by Rasnitsyn (2004).

It was assigned to Caloneurodea by Carpenter (1943).
Pleisiogrammidae was named by Carpenter (1943).

It was corrected as Pleisiogrammatidae by Martynova (1961); it was synonymized subjectively with Caloneuridae by Rasnitsyn (2004).

It was assigned to Caloneurodea by Carpenter (1943), Martynova (1961).
Anomalogrammidae was named by Carpenter (1943).

It was synonymized subjectively with Caloneuridae by Rasnitsyn (2004).

It was assigned to Caloneurodea by Carpenter (1943).
Eohymenidae was named by Martynov (1937). It is not extant.

It was considered an invalid subgroup of Palaeoptera by Carpenter (1992); it was synonymized subjectively with Caloneuridae by Rasnitsyn (2004), Rasnitsyn and Aristov (2013).

It was assigned to Protohymenoptera by Martynov (1937).
Amboneuridae was named by Carpenter (1980).

It was synonymized subjectively with Caloneuridae by Rasnitsyn (2004).

It was assigned to Caloneurodea by Carpenter (1980).
Sthenaroceridae was named by Handlirsch (1906). Its type is Sthenarocera.

It was synonymized subjectively with Protokollariidae by Carpenter (1992); it was synonymized subjectively with Caloneuridae by Rasnitsyn (2004).

It was assigned to Protorthoptera by Handlirsch (1906), Handlirsch (1919), Handlirsch (1922).

Synonymy list

Year	Name and author
1906	Caloneuridae Handlirsch p. 140
1906	Sthenaroceridae Handlirsch p. 149
1919	Caloneuridae Handlirsch p. 544
1919	Sthenaroceridae Handlirsch p. 551
1919	Caloneuridae Pruvost p. 115
1922	Caloneuridae Handlirsch p. 81
1922	Sthenaroceridae Handlirsch p. 86
1925	Caloneuridae Bolton p. 14
1928	Euthygrammidae Martynov p. 49
1937	Eohymenidae Martynov p. 8
1938	Caloneuridae Martynov p. 78
1938	Euthygrammidae Martynov p. 78
1943	Paleuthygrammidae Carpenter p. 70
1943	Pleisiogrammidae Carpenter p. 73
1943	Anomalogrammidae Carpenter p. 74
1961	Pleisiogrammatidae Martynova p. 269
1962	Euthygrammidae Zimmerman p. 1327
1980	Amboneuridae Carpenter p. 111
2009	Caloneuridae Beckemeyer p. 301

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Gravenhorst 1843
—	Deuteropoda
—	Mandibulata
—	Pancrustacea	Zrzavý and Štys 1997
—	Altocrustacea	Regier et al. 2010
superclass	Allotriocarida
—	Athalassocarida
—	Labiocarida	Schwentner et al. 2017

Rank	Name	Author
superclass	Hexapoda	Latreille 1825
—	Cercofilata
class	Insecta
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Polyneoptera	Martynov 1938
superorder	Archaeorthoptera
—	Panorthoptera	Crampton 1928
order	Caloneurodea
family	Caloneuridae
family	Caloneuridae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Caloneuridae Handlirsch 1906

show all | hide all

G. †Caloneura Brongniart 1885

†Caloneura dawsoni Brongniart 1885

Invalid names: Caloneura longicornis Handlirsch 1919 [synonym], Caloneura major Handlirsch 1919 [synonym], Caloneura picta Handlirsch 1919 [synonym], Caloneura robusta Handlirsch 1919 [synonym], Caloneura subtilis Bolton 1925 [synonym], Homaloneura royeri Meunier 1911 [synonym]

†Caloneura dawsonii Brongniart 1885

Invalid names: Boltonaloneura Rasnitsyn 2002 [synonym], Confusio Handlirsch 1919 [synonym]

G. †Caloneurella Carpenter 1934

†Caloneurella carbonaria Carpenter 1934

G. †Eohymen Martynov 1937

†Eohymen maculipennis Martynov 1937

G. †Euthygramma Martynov 1928

†Euthygramma parallellum Martynov 1928

Invalid names: Euthygramma aberrans Martynov 1938 [synonym], Euthygramma curvatum Martynov 1931 [synonym]

†Euthygramma storozhenkoi Rasnitsyn 2025

Invalid names: Paleuthygrammella Engel 1995 [synonym], Pseudogramma Carpenter 1943 [synonym]

G. †Isadistica Rasnitsyn and Aristov 2013

†Isadistica issada Rasnitsyn and Aristov 2013

†Isadistica longa Rasnitsyn and Aristov 2013

G. †Ligogramma Béthoux et al. 2004

†Ligogramma sinuosa Béthoux et al. 2004

†Ligogramma wichita Beckemeyer 2009

G. †Pleisiogramma Carpenter 1943

†Pleisiogramma maueri Novokshonov 1998

†Pleisiogramma medialis Carpenter 1943

Invalid names: Pleisiogramma reducta Carpenter 1943 [synonym]

G. †Vilviopsis Martynov 1938

†Vilviopsis extensa Martynov 1938

†Vilviopsis picta Rasnitsyn 2004

Invalid names: Amboneuridae Carpenter 1980 [synonym], Anomalogrammatidae Carpenter 1943 [synonym], Eohymenidae Martynov 1937 [synonym], Euthygrammatidae Martynov 1928 [synonym], Paleuthygrammatidae Carpenter 1943 [synonym], Plesiogrammatidae Carpenter 1943 [synonym], Sthenaroceridae Handlirsch 1906 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Bush and Bambach 2015, Kiessling 2004

Age range: base of the Westphalian D to the top of the Changhsingian or 309.80000 to 251.90200 Ma

Collections (25 total)

Time interval	Ma	Country or state	Original ID and collection number
Westphalian D	309.8 - 307.4	USA (Pennsylvania)	Amboneura klosei (122502) Caloneurella carbonaria (124504)
Gzhelian	303.7 - 298.9	France	Caloneura dawsoni, Caloneura subtilis (124916) Caloneura picta, Caloneura major, Caloneura longicornis, Caloneura robusta, Caloneura dawsonii (103635) Homaloneura royeri (121979)
Asselian	298.9 - 293.52	USA (New Mexico)	Vilviopsis picta (124385 124387)
Artinskian	290.1 - 283.3	Russian Federation (Perm)	Vilvia densinervosa (153669)
Artinskian	290.1 - 283.3	USA (Oklahoma)	Anomalogramma parva, Ligogramma sinuosa (107405) Ligogramma wichita (104821)
Artinskian	290.1 - 283.3	USA (Kansas)	Anomalogramma parva (114151) Plesiogramma medialis (111319) Plesiogramma reducta (111217)
Kungurian	283.3 - 274.4	Russian Federation	Pleisiogramma maueri (107787)
Roadian	274.4 - 266.9	Russian Federation	Euthygramma parallellum (116935)
Roadian	274.4 - 266.9	Russian Federation (Kazan)	Euthygramma curvatum (116803)
Roadian	274.4 - 266.9	Russian Federation (Arkangelsk)	Euthygramma aberrans (104993) Euthygramma parallellum (104716 116535 116933) Euthygramma parallellum, Vilviopsis extensa (116934) Euthygramma parallelum (104992)
Capitanian	264.28 - 259.51	Russian Federation (Orenburg)	Eohymen maculipennis (116607)
Wuchiapingian	259.51 - 254.14	Russian Federation (Vologda)	Isadistica issada, Isadistica longa (104754)
Changhsingian	254.14 - 251.902	Russian Federation (Orenburg)	Euthygramma storozhenkoi (233223)