PBDB Taxon

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Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Empis

Insecta - Diptera - Empididae

Taxonomy

Empis was named by Linnaeus (1758). It is extant. It is the type genus of Empididae, Empidinae, Empidini, Empidoidea.

It was assigned to Halterata by Scopoli (1763); to Diptera by Linnaeus (1758), Linnaeus (1761), Müller (1764), Schaeffer (1766), Linnaeus (1767), Müller (1776), Harris (1780), Latreille (1796); to Antliata by Fabricius (1775), Fabricius (1798); to Asilici by Latreille (1802); to Empidinae by Statz (1940); to Empididae by Aldrich (1905), James (1937), Carpenter (1992); and to Empidini by Waters (1989), Sinclair and Cumming (2006), Wahlberg and Johanson (2018).

Synonyms

Empis bulbirostris was named by Meunier (1899).

It was considered a nomen nudum by Meunier (1899).
Empis stilicornis was named by Meunier (1899).

It was considered a nomen nudum by Meunier (1899).
Empis tibialis was named by Meunier (1899).

It was considered a nomen nudum by Meunier (1899).

Synonymy list

Year	Name and author
1758	Empis Linnaeus p. 603
1761	Empis Linnaeus p. 466
1763	Empis Scopoli p. 368
1764	Empis Müller p. 87
1766	Empis Schaeffer
1767	Empis Linnaeus p. 1003
1775	Empis Fabricius p. 801
1776	Empis Müller p. 182
1780	Empis Harris p. 149
1796	Empis Latreille p. 155
1798	Empis Fabricius p. 568
1802	Empis Latreille p. 435
1899	Empis bulbirostris Meunier p. 178
1899	Empis stilicornis Meunier p. 178
1899	Empis tibialis Meunier p. 178
1905	Empis Aldrich p. 321
1937	Empis James p. 247
1940	Empis Statz p. 132
1989	Empis Waters
1992	Empis Carpenter
2006	Empis Sinclair and Cumming p. 74
2018	Empis Wahlberg and Johanson

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997
—	Panarthropoda
phylum	Arthropoda	Gravenhorst 1843
—	Deuteropoda
—	Mandibulata
—	Pancrustacea	Zrzavý and Štys 1997
—	Altocrustacea	Regier et al. 2010
superclass	Allotriocarida
—	Athalassocarida
—	Labiocarida	Schwentner et al. 2017
superclass	Hexapoda	Latreille 1825
—	Cercofilata
class	Insecta
—	Dicondylia

Rank	Name	Author
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Holometabola
—	Aparaglossata
—	Panorpida
superorder	Antliophora
order	Diptera
—	Neodiptera	Michelsen 1994
suborder	Brachycera	Zetterstedt 1842
—	Heterodactyla
—	Eremoneura
superfamily	Empidoidea	Latreille 1804
family	Empididae	Latreille 1804
subfamily	Empidinae	Latreille 1804
tribe	Empidini	Latreille 1804
genus	Empis

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. Empis Linnaeus 1758

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Subg. Empis (Acallomyia) Melander 1928

†Empis carbonum Germar 1837

†Empis exilis Meunier 1908

†Empis florissantana Cockerell 1915

†Empis infossa Melander 1949

†Empis inscita Meunier 1908

†Empis macilenta Meunier 1908

†Empis macrophthalma Förster 1891

†Empis mala Meunier 1908

†Empis malefica Meunier 1908

†Empis melia von Heyden 1870

†Empis miocenica Cockerell 1915

†Empis mordax Meunier 1908

†Empis morosella Melander 1928

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Invalid names: Empis morosa Meunier 1908 [replaced]

†Empis orapaensis Waters 1989

†Empis perdita Cockerell 1916

†Empis personata Meunier 1908

†Empis poeppigi Giebel 1856

†Empis pulvillata Loew 1850

†Empis spinifera Statz 1940

†Empis tritava Melander 1928

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Invalid names: Empis tristis Meunier 1908 [replaced]

†Empis umbonata Statz 1940

Invalid names: Empis bulbirostris Meunier 1899 [nomen nudum], Empis stilicornis Meunier 1899 [nomen nudum], Empis tibialis Meunier 1899 [nomen nudum]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Turonian to the top of the Late/Upper Miocene or 93.90000 to 5.33300 Ma
Minimum age of oldest fossil (stem group age): 89.8 Ma

Collections (13 total)

Time interval	Ma	Country or state	Original ID and collection number
Turonian	93.9 - 89.8	Botswana	E. orapaensis (123430)
Priabonian	37.71 - 33.9	Poland	E. pulvillata (109625)
Priabonian	37.71 - 33.9	Russian Federation (Kaliningrad)	E. malefica, E. mala, E. morosa (124073) E. poeppigi (147344) Hilara macilenta, E. exilis, E. inscita, E. personata, E. mordax, E. tristis, E. pulvillata, E. morosa (123927)
Chadronian	37.0 - 33.9	USA (Colorado)	E. infossa (124624) E. miocenica, E. florissantana (113749) E. perdita (124692)
Early/Lower Oligocene	33.9 - 27.3	France	E. macrophthalma (121473)
Chattian	27.3 - 23.04	Germany (North Rhine-Westphalia)	E. spinifera, E. umbonata (122010)
MP 30	27.3 - 23.04	Germany (Nordrhein-Westfalen)	E. melia (138696)
MP 30	27.3 - 23.04	Germany	E. melia (123945)
Late/Upper Miocene	11.63 - 5.333	Germany	E. carbonum (117341)