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Berothidae (beaded lacewing)

Insecta - Neuroptera - Berothidae

Synonyms
Synonymy list
YearName and author
1906Berothidae Handlirsch p. 42
1955Proberothidae Riek p. 674
1991Mesoberothidae Carpenter p. 87
1992Berothidae Carpenter
1992Mesoberothidae Carpenter
2002Berothidae Martins-Neto p. 37
2008Berothidae Engel and Grimaldi
2008Mesoberothidae Engel and Grimaldi
2018Berothidae Engel et al.
2018Mesoberothidae Engel et al.

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
—Deuteropoda
—Mandibulata
—PancrustaceaZrzavý and Štys 1997
—AltocrustaceaRegier et al. 2010
superclassAllotriocarida
—Athalassocarida
—LabiocaridaSchwentner et al. 2017
superclassHexapodaLatreille 1825
—Cercofilata
classInsecta
RankNameAuthor
—Dicondylia
—Paranotalia
subclassPterygota()
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
superorderNeuropterida
—Metaxyneuroptera
—Eidoneuroptera
orderNeuroptera
—Euneuroptera
superfamilyMantispoidea(Leach 1815)
familyBerothidae
familyBerothidae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Berothidae Handlirsch 1906 [beaded lacewing]
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G. †Aggregataberotha Wang et al. 2022
+
†Aggregataberotha paucipunctata Chen et al. 2024
†Aggregataberotha punctata Wang et al. 2022
G. †Ansoberotha Yang et al. 2019
+
†Ansoberotha jiewenae Yang et al. 2019
G. †Araripeberotha Martins-Neto and Vulcano 1990
+
†Araripeberotha fairchildi Martins-Neto and Vulcano 1990
G. †Banoberotha Whalley 1980
+
†Banoberotha enigmatica Whalley 1980
G. Berotha Walker 1860
Subfm. Berothinae Handlirsch 1906
+
G. †Elektroberotha Makarkin and Ohl 2015
+
†Elektroberotha groehni Makarkin and Ohl 2015
G. †Xenoberotha Makarkin 2017
+
†Xenoberotha angustialata Makarkin 2017
G. †Cantabroberotha Pérez-de la Fuente et al. 2020
+
†Cantabroberotha soplaensis Pérez-de la Fuente et al. 2020
G. †Caririberotha Martins-Neto and Vulcano 1990
+
†Caririberotha martinsi Martins-Neto and Vulcano 1990
G. †Cornoberotha Yang et al. 2019
+
†Cornoberotha anomala Yang et al. 2019
†Cornoberotha aspoeckae Yang et al. 2019
†Cornoberotha monogona Yang et al. 2019
G. †Danoberotha Makarkin et al. 2024
+
†Danoberotha verkleijorum Makarkin et al. 2024
G. †Dasyberotha Engel and Grimaldi 2008
+
†Dasyberotha eucharis Engel and Grimaldi 2008
G. †Dolichoberotha Yang et al. 2019
+
†Dolichoberotha bifurcata Yang et al. 2019
†Dolichoberotha burmana Yang et al. 2019
G. †Epimesoberotha Jepson et al. 2012
+
†Epimesoberotha parva Jepson et al. 2012
G. †Ethiroberotha Engel and Grimaldi 2008
+
†Ethiroberotha elongata Engel and Grimaldi 2008
G. †Ferganoberotha Khramov 2023
+
G. †Haploberotha Engel and Grimaldi 2008
+
†Haploberotha carsteni Makarkin 2018
†Haploberotha persephone Engel and Grimaldi 2008
G. †Iceloberotha Engel and Grimaldi 2008
+
†Iceloberotha kachinensis Engel and Grimaldi 2008
†Iceloberotha simulatrix Engel and Grimaldi 2008
G. Isoscelipteron Costa 1863
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†Isoscelipteron bertrandi Boderau et al. 2024
G. †Jersiberotha Grimaldi 2000
+
†Jersiberotha luzzii Grimaldi 2000
†Jersiberotha musivum Zhang et al. 2022
†Jersiberotha myanmarensis Engel and Grimaldi 2008
†Jersiberotha similis Grimaldi 2000
†Jersiberotha tauberorum Engel and Grimaldi 2008
G. †Khasurtoberotha Khramov 2020
+
G. †Krokhathone Khramov 2015
+
†Krokhathone parva Khramov 2015
†Krokhathone tristis Khramov 2015
G. †Maculaberotha Yuan et al. 2016
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†Maculaberotha nervosa Yuan et al. 2016
G. †Magniberotha Yuan et al. 2016
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†Magniberotha recurrens Yuan et al. 2016
Subfm. †Mesithoninae Panfilov 1980
+
G. †Berothone Khramov 2015
+
†Berothone gracilis Panfilov 1980
†Berothone protea Panfilov 1980
G. †Mesithone Panfilov 1980
+
†Mesithone angusta Makarkin 1999
†Mesithone grandis Panfilov 1980
†Mesithone maculata Panfilov 1980
†Mesithone maculosa Makarkin 1999
†Mesithone magna Panfilov 1980
†Mesithone proberotha Makarkin 1999
G. †Pseudosisyra Makarkin 1999
+
†Pseudosisyra minima Makarkin 1999
G. †Mesoberotha Carpenter 1991
+
†Mesoberotha superba Riek 1955
Invalid names: Proberotha Riek 1955 [replaced]
G. †Microberotha Archibald and Makarkin 2004
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†Microberotha macculloughi Archibald and Makarkin 2004
G. †Nascimberotha Grimaldi 2000
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†Nascimberotha picta Grimaldi 2000
G. †Oloberotha Ren and Guo 1996
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†Oloberotha sinica Ren and Guo 1996
G. †Osmyloberotha Khramov 2021
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†Osmyloberotha simpla Khramov 2021
G. †Protoberotha Huang et al. 2019
+
†Protoberotha minuta Huang et al. 2019
G. †Sejunctaberotha Chen et al. 2024
+
†Sejunctaberotha sphaerica Chen et al. 2024
†Sejunctaberotha tenuis Chen et al. 2024
†Sejunctaberotha transversa Chen et al. 2024
G. †Sibelliberotha Azar and Nel 2013
+
†Sibelliberotha rihanensis Azar and Nel 2013
G. †Sinosmylites Hong 1983
+
†Sinosmylites auliensis Khramov 2015
†Sinosmylites fumosus Makarkin et al. 2011
†Sinosmylites hotgoricus Khramov 2015
†Sinosmylites karatavicus Khramov 2015
†Sinosmylites rasnitsyni Makarkin et al. 2011
G. †Systenoberotha Engel and Grimaldi 2008
+
†Systenoberotha magillae Engel and Grimaldi 2008
G. †Telistoberotha Engel and Grimaldi 2008
+
†Telistoberotha libitina Engel and Grimaldi 2008
G. †Triassoberotha Khramov et al. 2023
+
†Triassoberotha japonica Khramov et al. 2023
G. †Xiaoberotha Shi et al. 2019
+
†Xiaoberotha bipunctata Shi et al. 2019
†Xiaoberotha dilatapoda Wang et al. 2022
Invalid names: Cyrenoberothinae Macleod and Adams 1967 [empty], Mesoberothidae Carpenter 1991 [synonym], Rhachiberothinae Tjeder 1959 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Ladinian to the top of the Messinian or 241.46400 to 5.33300 Ma
Minimum age of oldest fossil (stem group age): 237.0 Ma

Collections (52 total)


Time interval Ma Country or state Original ID and collection number
Ladinian241.464 - 237.0Kyrgyzstan (Osh) Ferganoberotha minutissima (105007)
Carnian237.0 - 227.3Japan (Yamaguchi) Berothidae indet., Triassoberotha japonica (106482)
Norian227.3 - 205.7Australia (Queensland) Proberotha superba, Proberothella elongata (106484)
Callovian - Oxfordian165.3 - 154.8China (Nei Mongol) Sinosmylites sp., Sinosmylites rasnitsyni, Sinosmylites fumosus (117610)
Callovian - Oxfordian165.3 - 154.8China (Liaoning) Sinosmylites pectinatus (154730)
Callovian - Oxfordian165.3 - 154.8Kazakhstan (Chimkent) Berothone protea (106279) Krokhathone tristis (106101) Mesithone gracilis (105970) Mesithone maculata, Mesithone magna, Berothone protea (113439) Mesithone protea, Mesithone grandis, Krokhathone parva, Sinosmylites auliensis (105971) Sinosmylites karatavicus (105426)
Tithonian149.2 - 143.1Mongolia (Dund-Gov’ Aymag) Sinosmylites hotgoricus (122321)
Early/Lower Berriasian143.1 - 141.0United Kingdom (England) Epimesoberotha parva (122911 122913)
Early/Lower Barremian125.77 - 121.4Lebanon Berothidae indet., Banoberotha enigmatica (123001) Sibelliberotha rihanensis (153333)
Late/Upper Barremian125.77 - 121.4China (Liaoning) Oloberotha sinica (138436)
Early/Lower Aptian121.4 - 119.57Russian Federation (Chita) Mesithone sp. (119499)
Aptian121.4 - 113.2Russian Federation (Buryatia) Khasurtoberotha bellissima (113350) Mesithone angusta (154573) Mesithone maculosa (122548) Mesithone proberotha (119498 123465) Pseudosisyra minima (122549)
Late/Upper Aptian119.57 - 113.2Brazil (Ceará) Araripeberotha fairchildi (123013) Caririberotha martinsi (125511)
Early/Lower Albian113.2 - 110.1Spain Cantabroberotha soplaensis (109438)
Late/Upper Albian106.3 - 100.5Spain (Álava) Berothidae indet., Berothidae indet. 1, Berothidae indet. 2, Berothidae indet. 3, Berothidae indet. 4 (108769)
Late/Upper Albian106.3 - 100.5Spain (Aragón) Berothidae indet. (91626)
Early/Lower Cenomanian100.5 - 93.9Myanmar Ansoberotha jiewenae, Cornoberotha aspoeckae, Cornoberotha monogona, Dolichoberotha bifurcata, Dolichoberotha burmana, Xiaoberotha sp. (161726) Berothidae indet., Berothidae larva indet., Jersiberotha myanmarensis, Jersiberotha tauberorum, Iceloberotha kachinensis, Iceloberotha simulatrix, Haploberotha persephone, Telistoberotha libitina, Dasyberotha eucharis, Systenoberotha magillae, Ethiroberotha elongata (111688) Cornoberotha anomala, Cornoberotha aspoeckae, Dolichoberotha bifurcata, Xiaoberotha bipunctata (178121) Haploberotha carsteni (172900) Haploberotha persephone (123116) Jersiberotha musivum (216019) Maculaberotha nervosa, Magniberotha recurrens, Protoberotha minuta, Ansoberotha jiewenae, Aggregataberotha punctata, Xiaoberotha dilatapoda, Aggregataberotha paucipunctata, Sejunctaberotha sphaerica, Sejunctaberotha tenuis, Sejunctaberotha transversa (165561) Osmyloberotha simpla (190961) Xiaoberotha bipunctata (178159)
Turonian93.9 - 89.8USA (New Jersey) Jersiberotha luzzii, Jersiberotha similis, Nascimberotha picta (110895)
Late/Upper Campanian83.6 - 72.2Myanmar Berothidae indet. (195455)
Middle Campanian83.6 - 72.2Canada (Alberta) Berothidae indet. (104932) Chrysopidae indet. (122384)
Ypresian56.0 - 48.07Denmark Danoberotha verkleijorum (236577)
Ypresian56.0 - 48.07Canada (British Columbia) Microberotha macculloughi (123795)
Uintan46.2 - 39.7USA (Colorado) Xenoberotha angustialata (184083)
Priabonian37.71 - 33.9Russian Federation (Kaliningrad) Berothidae indet. larva A, Berothidae indet. larva B (140480) Berothidae indet. larva D (152019) Elektroberotha groehni (110735 147612)
Priabonian37.71 - 33.9Russian Federation Berothidae indet. larva C (123419)
Priabonian37.71 - 33.9Ukraine Berothidae indet. (122997)
Messinian7.246 - 5.333France (Cantal) Isoscelipteron bertrandi (122195)