PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Tylostoma

Gastropoda - Tylostomatidae

Taxonomy

Tylostoma was named by Sharpe (1849) [Sepkoski's age data: J Kimm T Dani Sepkoski's reference number: 914]. Its type is Tylostoma torrubiae.

It was assigned to Gastropoda by Acquaotta (1929); to Euspirinae by Allison (1955); to Naticidae by Kase (1984); to Neotaenioglossa by Sepkoski (2002); and to Tylostomatidae by Squires and Saul (2004), Kollmann (2009).

Species

T. (Tylostoma), T. aequatoriale, T. allisoni, T. cossoni, T. garzana, T. globosum, T. laharpi, T. materinum, T. minimum, T. ovatum, T. pallaryi, T. pechueli, T. sergipensis, T. torrubiae (type species), T. whitei, Varicigera choffati

Synonyms

Varigera was named by d'Orbigny (1850).

It was synonymized subjectively with Tylostoma by Squires and Saul (2004).
Varicigera was named by Douvillé (1916) [Synonym of Tylostoma according to Squires and Saul (2004)].

It was synonymized subjectively with Tylostoma by Squires and Saul (2004).

Synonymy list

Year	Name and author
1849	Tylostoma Sharpe p. 24
1850	Varigera d'Orbigny
1916	Varicigera Douvillé
1929	Thylostoma Acquaotta
1955	Tylostoma Allison p. 428
1984	Tylostoma Kase p. 157
2002	Tylostoma Sepkoski
2004	Tylostoma Squires and Saul p. 24
2009	Tylostoma Kollmann p. 58

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Spiralia
superphylum	Lophotrochozoa
phylum	Mollusca

Rank	Name	Author
class	Gastropoda
subclass	Caenogastropoda	(Cox 1959)
—	Sorbeoconcha	(Ponder and Lindberg 1997)
—	Campanilimorpha	(Haszprunar 1988)
superfamily	Campaniloidea	Douvillé 1904
family	Tylostomatidae	Stoliczka 1868
genus	Tylostoma	Sharpe 1849

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Tylostoma Sharpe 1849

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Subg. †Tylostoma (Tylostoma) Sharp 1849

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†Tylostoma (Tylostoma) mutabilis Gabb 1869

†Tylostoma aequatoriale Kossmat 1893

†Tylostoma allisoni Squires and Saul 2004

†Tylostoma cossoni Thomas and Peron 1889

†Tylostoma garzana Anderson 1958

†Tylostoma globosum Sharpe 1849

†Tylostoma laharpi Pictet and Campiche 1862

†Tylostoma materinum White 1887

†Tylostoma minimum White 1887

†Tylostoma ovatum Sharpe 1849

†Tylostoma pallaryi Peron and Fortau 1904

†Tylostoma pechueli Choffat 1888

†Tylostoma sergipensis Maury 1936

†Tylostoma torrubiae Sharpe 1849

†Tylostoma whitei Ihering 1907

Invalid names: Varicigera Douvillé 1916 [synonym], Varigera d'Orbigny 1850 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

aragonitec

Locomotion:

actively mobilesuperf

Life habit:

epifaunalsuperf

Diet:

grazersuperf

Vision:

limitedc

Created:

2009-09-19 08:44:14

Modified:

2009-09-19 10:44:14

Source: superf = superfamily, c = class

References: Kiessling 2004, Hendy et al. 2009

Age range: base of the Late/Upper Aalenian to the top of the Selandian or 174.70000 to 59.24000 Ma

Collections (171 total)

Time interval	Ma	Country or state	Original ID and collection number
Late/Upper Aalenian	174.7 - 170.9	Morocco	T. globosum (141219)
Bajocian - Bathonian	170.9 - 165.3	USA (Utah)	T. sp. (171916 171928 171941 171947 171948 172024 172027 172038 172041 172042 172045)
Early/Lower Bajocian - Late/Upper Oxfordian	170.9 - 154.8	Romania (Brasov)	T. sp. (81442)
Middle Oxfordian	161.5 - 154.8	France	T. ponderosum (11085)
Oxfordian - Kimmeridgian	161.5 - 149.2	Yemen	T. sp. (94504)
Berriasian - Valanginian	143.1 - 132.6	Chile (Coquimbo)	T. sp. (221666)
Early/Lower Valanginian	137.05 - 132.6	Switzerland	T. fallax, T. naticoide, T. laharpi (2225)
Late/Upper Valanginian	137.05 - 132.6	Hungary	T. bulimoides (25152)
Valanginian	137.05 - 132.6	Switzerland	T. fallax, T. laharpi (2217 2224) T. fallax, T. naticoide, T. laharpi (2247) T. fallax., T. laharpi (2240) T. naticoide (2215) T. sp. (2243)
Valanginian	137.05 - 132.6	France	T. villersense (2222)
Early/Lower Hauterivian	132.6 - 125.77	Argentina (Neuquen)	T. jaworskii (47202)
Hauterivian	132.6 - 125.77	Mexico (Coahuila)	T. laharpi (2145)
Barremian	125.77 - 121.4	Venezuela	T. sp. (90531)
Early/Lower Barremian	125.77 - 121.4	Chile (Copiapó)	T. jaworskii (223061)
Late/Upper Barremian - Aptian	125.77 - 113.2	Japan (Honshu)	T. sanchuense (50307)
Early/Lower Aptian	121.4 - 119.57	Switzerland	T. rochatiana (2175 2232)
Early/Lower Aptian	121.4 - 119.57	Ethiopia	T. rochatianum (46586)
Early/Lower Aptian	121.4 - 119.57	Spain (Valenciana)	T. sp. (45154)
Aptian	121.4 - 113.2	Portugal	T. sp. (73507)
Aptian	121.4 - 113.2	Mexico	T. aguilerai (69900)
Aptian	121.4 - 113.2	Switzerland	T. depressum (2229) T. ellipticum, T. depressum (2188) T. rochatiana (2173 2231)
Aptian - Albian	121.4 - 100.5	China (Xizang)	T. sp. (61399)
Late/Upper Aptian	119.57 - 113.2	Mexico (Baja California)	T. allisoni (1890) T. materinum (1881) T. mutabilis (1878)
Late/Upper Aptian	119.57 - 113.2	Japan	T. hanaii (91261) T. miyakoense (91258 91283 91287 91288 91291 91292 91293 91308) T. miyakoense, T. hanai (91286) T. miyakoensis (91260 91262)
Late/Upper Aptian	119.57 - 113.2	Angola	T. pechueli (210857)
Late/Upper Aptian	119.57 - 113.2	Egypt	T. magharensis (151611 151613)
Early/Lower Albian	113.2 - 110.1	Mexico	T. sp. (161077)
Early/Lower Albian	113.2 - 110.1	Egypt	T. magharensis, T. canaliculata (98325)
Early/Lower Albian	113.2 - 110.1	Gabon	T. pollaryi (209483)
Albian	113.2 - 100.5	Mexico (Coahuila de Zaragoza)	T. kentense (88630 88631)
Albian	113.2 - 100.5	Switzerland	T. gaultinum (2166 2167)
Middle Albian	110.1 - 106.3	Egypt	T. magharensis (98326)
Middle Albian	110.1 - 106.3	Brazil (Sergipe)	T. materinum (87860) T. whitei, T. minimum (87861 87866)
Late/Upper Albian	106.3 - 100.5	Angola	T. globosum (210728 210738)
Late/Upper Albian	106.3 - 100.5	Egypt	T. cossoni, T. pallaryi (140606) T. pallaryi (140611) T. sp. 1, T. sp. 2, T. globosum, T. choffati, T. magharensis (98327) Varicigera choffati (171310)
Late/Upper Albian	106.3 - 100.5	Spain	T. escragnollensis (66129)
Late/Upper Albian	106.3 - 100.5	Brazil (Sergipe)	T. sergipensis (87857)
Late/Upper Albian	106.3 - 100.5	Iran	T. sp. (185775 185784 185794)
Late/Upper Albian - Cenomanian	106.3 - 93.9	Egypt	T. cossoni, T. pallaryi (150677 150737) T. pallaryi (150874)
Cenomanian	100.5 - 93.9	Spain	T. ovatum (175376) T. torrubiae, T. ovatum (175374)
Cenomanian	100.5 - 93.9	Serbia	T. sp., T. minimum (239139)
Cenomanian	100.5 - 93.9	Germany (Nordrhein-Westfalen)	T. sp. (94265)
Cenomanian	100.5 - 93.9	Italy (Campania)	T. schiosensis (84801)
Early/Lower Cenomanian	100.5 - 93.9	Algeria (Batna)	T. cossoni (234831) T. elevatum, T. cossoni, T. globosum (234835) T. globosum (234827) T. torrubiae, T. cossoni, T. globosum (234823)
Early/Lower Cenomanian	100.5 - 93.9	USA (New Mexico)	T. harrisi (179775)
Early/Lower Cenomanian	100.5 - 93.9	Egypt	T. cossoni (140114) T. cossoni, T. pallaryi (140613) T. pallaryi (114872)
Early/Lower Cenomanian - Middle Cenomanian	100.5 - 93.9	Egypt	T. cossoni (150614) T. cossoni, T. globosum (150675)
Late/Upper Cenomanian	100.5 - 93.9	Egypt	T. cossoni (114797 114798 114810 140579) T. pallaryi (114794 140578)
Late/Upper Cenomanian	100.5 - 93.9	Portugal (Região do Centro)	T. ovatum (224527)
Late/Upper Cenomanian	100.5 - 93.9	Algeria (Batna)	T. globosum (234838) T. pallaryi (234825 234826 234829)
Late/Upper Cenomanian	100.5 - 93.9	Libya	T. cossoni (208627)
Middle Cenomanian	100.5 - 93.9	Algeria (Batna)	T. cossoni (234824)
Middle Cenomanian	100.5 - 93.9	Egypt	T. cossoni (140233)
Late/Upper Cenomanian - Early/Lower Turonian	100.5 - 89.8	Egypt	T. globosum (150676)
Early/Lower Turonian	93.9 - 89.8	Tunisia	T. cossoni (88843 88846 88853 88854)
Early/Lower Turonian	93.9 - 89.8	Brazil	T. sp. (98673)
Early/Lower Turonian	93.9 - 89.8	Portugal (Região do Centro)	T. ovatum (224528)
Early/Lower Turonian	93.9 - 89.8	Egypt	T. cossoni, T. pallaryi (114799) T. globosum (114815 114897 140245 140264 140317 140587 140622)
Early/Lower Turonian	93.9 - 89.8	Mexico	T. ovatum (73576)
Early/Lower Turonian	93.9 - 89.8	Spain	T. torrubiae, T. ovatum (175375)
Early/Lower Turonian - Middle Turonian	93.9 - 89.8	Egypt	T. globosum (150612)
Late/Upper Turonian	93.9 - 89.8	Brazil	T. sp. (140890)
Late/Upper Turonian	93.9 - 89.8	Egypt	T. globosum (114823 114824 140592 140594 140595 150674)
Late/Upper Turonian	93.9 - 89.8	France	T. cossoni (59230)
Middle Turonian	93.9 - 89.8	Kyrgyzstan (Batken)	T. ferganense, T. subglobosum, T. globosum (3808)
Middle Turonian	93.9 - 89.8	Egypt	T. cossoni (114800 140319 140320 140623) T. globosum (114818 114820 114901)
Middle Turonian - Late/Upper Turonian	93.9 - 89.8	Egypt	T. cossoni (114801)
Turonian	93.9 - 89.8	Spain	T. torrubiae, T. ovatum (175378)
Turonian	93.9 - 89.8	Gabon	T. aequatoriale (209641) T. cossoni (209529 209532 209535) T. globosum (209534) T. sp. A (209533) T. sp. B, T. cossoni (209530)
Turonian	93.9 - 89.8	Egypt	T. cossoni (150613)
Early/Lower Coniacian	89.8 - 85.7	Brazil	T. spp. (140901)
Late/Upper Coniacian	89.8 - 85.7	Egypt	T. athleticum (114804)
Santonian	85.7 - 83.6	France	T. globosa, T. sp. (3731)
Late/Upper Campanian	83.6 - 72.2	Saudi Arabia	T. sp. (184844)
Late/Upper Campanian - Maastrichtian	83.6 - 66.0	United Arab Emirates	T. sp. (67456)
Early/Lower Maastrichtian - Late/Upper Maastrichtian	72.2 - 66.0	USA (California)	Phasianella garzana (215406)
Maastrichtian	72.2 - 66.0	Brazil (Paraíba)	T. materinum (88070)
Maastrichtian	72.2 - 66.0	USA (California)	Phasianella garzana (83790 83791 185450 185452)
Danian	66.0 - 61.66	France	T. ampullariaeforme (20816)
Danian	66.0 - 61.66	Greenland	T. ampullariaeformis (22943)
Selandian	61.66 - 59.24	Greenland	T. ampullariaeformis (22944)