Subfamily Scarabaeinae Latreille 1802 (scarab beetle)

Insecta - Coleoptera - Scarabaeidae

Full reference: P. A. Latreille. 1802. Histoire Naturelle, Générale et Particulière des Crustacés et des Insectes, Tome Troisième 1-467

Parent taxon: Scarabaeidae according to F. Tello et al. 2023

See also Bouchard et al. 2011, Cai et al. 2022, Krell 2000, Lawrence and Newton 1995, Nikolajev 2007, Tarasov and Dimitrov 2016, Tarasov et al. 2016 and Wappler 2003

Sister taxa: none

Subtaxa: Anachalcos Ateuchini Canthochilum Coprini Deltochilini Gymnopleurini Heliocopris Metacatharsius Oniticellini Onitini Onthophagini Onthophagus spitsbergeniensis Phanaeini Scarabaeini

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Type: Scarabaeus

Ecology:

Distribution:

• Quaternary of Chile (2 collections), Ecuador (1), Japan (7), Peru (1), the United Kingdom (9), United States (6: California, Pennsylvania, Virginia)

• Pliocene of Georgia (1), Germany (1), Japan (3)

• Miocene of China (1), the Dominican Republic (1), Germany (2), Japan (1), Kenya (2)

• Oligocene of France (1), Germany (1)

• Eocene of France (1), Germany (2), the United Kingdom (1)

• Paleocene of Svalbard and Jan Mayen (1)

Total: 45 collections including 70 occurrences

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