Subfamily Ursinae Swainson 1835 (bear)

Mammalia - Carnivora - Ursidae

Parent taxon: Ursidae according to R. M. Hunt 1998

See also Bjork 1970, Jin et al. 2007 and Kurten and Anderson 1980

Sister taxa: Agriotheriinae, Ailuropodinae, Amphicynodontinae, Harpaleocyon, Hemicyon, Hemicyoninae, Metarctos, Myadina, Phoberocyoninae, Proarctotherium, Subparictidae, Taxina, Thaumastocyon, Tremarctinae, Ursulus, Zaragocyon

Subtaxa: Melursus Tremarctini Ursavini Ursini

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Ecology: ground dwelling herbivore-carnivore

Distribution:

• Quaternary of Austria (1 collection), Belgium (2), Canada (15: British Columbia, Northwest Territories, Nunavut, Ontario, Quebec, Yukon), China (11), Croatia (2), the Czech Republic (3), Denmark (2), France (24), Georgia (2), Germany (20), Greece (9), Greenland (10), Hungary (6), Indonesia (1), Israel (3), Italy (36), Japan (4), Kazakhstan (1), Libya (1), Mexico (2), Monaco (3), Mongolia (1), Morocco (4), the Netherlands (4), Norway (3), Romania (5), the Russian Federation (10), Spain (19), Svalbard and Jan Mayen (1), Sweden (3), Taiwan (1), Turkey (3), the United Kingdom (14), United States (96: Alabama, Alaska, Arkansas, California, Colorado, Florida, Georgia, Idaho, Illinois, Indiana, Iowa, Kentucky, Louisiana, Maine, Maryland, Massachusetts, Michigan, Mississippi, Montana, Nevada, New Mexico, Ohio, Pennsylvania, Tennessee, Texas, Virginia, West Virginia, Wyoming), Vietnam (1)

• Blancan of United States (5: California, Idaho, Nevada, Texas)

• Pliocene to Pleistocene of France (2), Greece (3), Hungary (1), Italy (2), Turkey (1)

• Pliocene of Bulgaria (2), Canada (1: Northwest Territories), China (1), Ethiopia (1), France (3), Georgia (2), Greece (6), Hungary (6), Italy (2), Libya (1), Mexico (1), Moldova (3), Morocco (1), Poland (1), Romania (2), the Russian Federation (3), South Africa (3), Spain (2), Tajikistan (4), Ukraine (2), the United Kingdom (2)

• Hemphillian of Mexico (6), United States (25: Arizona, California, Florida, Kansas, Nebraska, Nevada, Oklahoma, Oregon, Texas)

• Miocene to Pliocene of Ethiopia (1), South Africa (1)

• Neogene of Italy (1)

• Miocene of Algeria (1), Bosnia and Herzegovina (1), Canada (1: Saskatchewan), China (8), France (8), Germany (1), Greece (1), Hungary (1), Italy (1), Kazakhstan (1), Libya (2), Poland (1), Slovakia (2), Spain (1), Switzerland (1), Turkey (5), United States (13: Arizona, Colorado, Florida, Nebraska, Nevada, Oregon, South Dakota)

• Arikareean of United States (1: Oregon)

Total: 470 collections including 517 occurrences

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