Subfamily Testudininae Gray 1825 (turtle)

Reptilia - Testudines - Pantestudinidae

Full reference: J. E. Gray. 1825. A synopsis of the genera of Reptilia and Amphibia. Annals of Philosophy, n. s., London 10:193-217

Parent taxon: Testudinidae according to A. Perez-Garcia et al. 2020

See also de Broin and de la Fuente 1993, Estes and Tihen 1964, Hervet and de Broin 2000, Holman 1985 and Kuhn 1966

Sister taxa: Achilemys, Agrionemys, Aldabrachelys, Anhuichelys, Astrochelys, Cachuga, Caryoderma, Centrochelys, Cheirogaster, Chelonoidis, Chelys patagonica, Chersina, Chersine, Cylindraspis, Damonia, Emys sombrerensis, Ergilemys, Floridemys, Floridemys hurdi, Fontainechelon, Geochelone, Geochelone (Chelonoidis), Geochelone (Hesperotestudo) equicomes, Gigantochersina, Gopherus, Gopherus praecedens, Hesperotestudo, Homopus, Impregnochelys, Indotestudo, Juvemys, Kansuchelys, Kinixys, Malacochersus, Manouria, Megalochelys, Mesochersus, Miotestudo, Namibchersus, Nicoria, Palaeoemys, Paleotestudo, Pelorochelon, Platysterninae, Protestudo, Psammobates, Pyxis, Sinohadrianus, Stigmochelys, Stylemys, Stylemys calaverensis, Taraschelon, Testudininei, Testudinites sellovii, Testudo, Testudo annae, Testudo atascosae, Testudo castrensis, Testudo cultrata, Testudo exornata, Testudo formosa, Testudo francisi, Testudo hayi, Testudo klettiana, Testudo laticaudata, Testudo louisekressmani, Testudo luciae, Testudo ocalana, Testudo paranensis, Testudo peragrans, Testudo quadratus, Testudo undata, Titanochelon

Subtaxa: Geochelona Geoemydinei

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Ecology: ground dwelling herbivore-herbivore

Distribution:

• Pliocene of Spain (1 collection)

• Miocene of United States (1: California)

• Deseadan of Argentina (1)

• Oligocene of Oman (1)

Total: 4 collections each including a single occurrence

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