Full reference: F. Welker, M. J. Collins, J. A. Thomas, M. Wadsley, S. Brace, E. Cappellini, S. T. Turvey, M. Reguero, J. N. Gelfo, A. Kramarz, J. Burger, J. Thomas-Oates, D. A. Ashford, P. D. Ashton, K. Rowsell, D. M. Porter, B. Kessler, R. Fischer, C. Baessmann, S. Kaspar, J. V. Olsen and P. Kiley. 2015. Ancient proteins resolve the evolutionary history of Darwin’s South American ungulates. Nature 522:81-84
Parent taxon: Euungulata according to F. Welker et al. 2015
Sister taxa: Artiodactylamorpha, Haloboidea, Amblypoda, Diplarthra, Cetungulata, Barypoda, Kharmerungulatum, Protungulata, Sivatheridae, Tipotheria, Polydactyli, Trichenae, Barytheria, Monochenae, Capridae, Palaeocladosictis, Ruminantes, Procreodi, Altungulata, Ganungulatum, Mesodactyli, Hydrophorae, Hyraces, Tesserachenae, Antilopidae
Subtaxa: Panameriungulata Perissodactylamorpha Phenacodonta
Ecology: scansorial insectivore