Reptilia - Testudines - Testudinidae

This is a clade name that includes the so-called "Geochelone complex"

Parent taxon: Testudininae according to A. Pérez-García et al. 2020

Sister taxa: Agrionemys, Centrochelys, Cheirogaster, Ergilemys, Geochelone, Geoemydinei, Gigantochersina, Impregnochelys, Indotestudo, Kinixys, Mesochersus, Namibchersus, Paleotestudo, Stigmochelys, Stylemys, Testudinini

Subtaxa: Alatochelon

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Ecology: ground dwelling herbivore-herbivore

Distribution: found only at Puerto de la Cadena (Pliocene of Spain)

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