Mammalia - Artiodactyla - Bovidae
Parent taxon: Caprinae according to A. W. Gentry 2010
See also Grubb 2001
Sister taxa: Aragoral, Benicerus, Capricornis, Damalavus, Hemitragus, Myotragus, Naemorhedini, Ovibos, Pseudois, Rupicapra, Rupicaprini
Subtaxa: Makapania Nitidarcus
Ecology: ground dwelling grazer-browser
Distribution:
• Quaternary of Canada (2: Yukon collections), Ethiopia (3), France (1), Italy (1), the Russian Federation (1), South Africa (2), Spain (1), United States (3: California, Tennessee, Virginia)
• Pliocene to Pleistocene of China (1), South Africa (1)
• Pliocene of Greece (1), Kenya (1), South Africa (3), Spain (1)
• Miocene to Pleistocene of Spain (1)
• Miocene to Pliocene of South Africa (1)
• Miocene of Iraq (1)
Total: 25 collections including 27 occurrences