Full reference: R. J. Tillyard. 1924. Upper Permian Coleoptera and a new order from the Belmont Beds, New South Wales. Proceedings of the Linnean Society of New South Wales 49:429-435
Parent taxon: Coleoptera according to P. Bouchard et al. 2011
Sister taxa: Acmaeodera perreaui, Adelidium, Adikia, Adynasia, Allognosis, Allopliosilpha, Alphacoleoptera, Anancaeon, Anapiptus, Ancylocheira liasina, Anepismus, Angelinella, Anhuistoma, Anhydrophilus, Anomerus, Anypostatus, Apheloodes, Apicasia, Apioderes, Apiopyrenides, Apistotes, Apsychus, Aptilotitus, Archosyne, Auchenophorites, Babuskaya, Bareus, Bathygerus, Bellingera, Bellingeropsis, Biadelater, Blaptoides, Bothroptera, Brachylaimon, Brachytrachelites, Brevispinnatus, Brochocoleus indibili, Brodiola, Bucklandula, Buprestium, Campopsis, Carabicina, Carabiques, Carabocera, Cardioides, Cardiosyne, Cebrionates, Cerylonopsis, Chimaerocoleites, Chlaeniopsis, Chrysomelidium, Chrysomelinae, Chrysomelophana, Cicindeletae, Cistela, Cistelites, Clathropenna, Clerii, Clinomerus, Coccinellophana, Coleopterites, Coleopteron, Coleopteron depressus, Coleopteron interpunctatus, Coleopteron laevigatus, Coleopteron scabriusculus, Colymbetopsis, Coprophagi, Cretaceites, Cretofalselaterium, Cretogermen, Cretohelophorus, Criocerina pristina, Critoderma, Cryptauchenia, Curculidium, Curculionites liasinus, Curculiopsidae, Curculium, Cydnopsis nigromembranacea, Cyphospheron, Darbiopelta, Diachoristes, Diaperialae, Diaperidium, Diatarastus, Diatrypamene, Dicyphelus, Diphymation, Diplocelides, Diplothece, Doggeria, Dysarestus, Ecthlimma, Elateridopsidae, Elateridopsis, Elaterina, Elytrulum, Enamma, Entomocantharus, Entomophagi, Eocoleopteron, Eodromeus viriosus, Episcepes, Epomenus, Erotilenae, Erotylites, Etheridgea, Euchroma arvernica, Eucoleoptera, Eurynotellus, Eurysphinctus, Eusarcantarus, Explansimarginis, Fengningia, Flichea, Fuscicupes, Gastrodelus, Gastroratus, Gemelina, Glaphoptera, Glaphyropterites, Glaphyropterula, Grahamelytron, Grasselites, Gurvanocoleus, Gyrinites, Gyrinulopsis, Halticophana, Harpalidium, Heeriaopsis, Helophoropsis, Helophorus exilis, Helophorus magnus, Helopidium, Helopii, Helopium, Helops atticus, Hubeicoleus, Hydrobius godeti, Hydrobius obsoletus, Hydrocanthares, Hydroicetes, Hydrophilites, Hydrophilus stenopterus, Hydroporopsis, Hydrous ebeninus, Hylesinus lineatus, Hyperomima, Ipidae, Isomira aurora, Kamaroma, Katapiptus, Keleusticus, Kibdelia, Kryptophagus, Laimocenos, Lamiites, Lamiophanes, Laodiscis, Latridiites, Leioodes, Leptomites, Leptopodocoleidae, Leptosolenophorus, Longimaxilla, Loxocamarotus, Macrogastri, Macrotrachelites, Malacodermi, Malacodermidae, Masselytron, Megacentrus, Megachorites, Megopis lineolatus, Melanophilites, Melanophilopsis, Menephiloides, Mesocoleoptera, Mesoncus, Mesorhynchophora, Mesotricupes, Mesotylites, Metanastes, Metaxycoleoptera, Micrelaterium, Microcoleopteron, Microentomus, Mimelater, Nannoodes, Nebrioides, Necrophagi, Necticus minutus, Nitidulariae, Nubili, Omogongylus, Onkedodimus, Onthophagus luteus, Ooidellus, Ooperiglyptus, Ooperioristus, Oropseini, Oxycephalites, Oxytoroptera, Pachycoleon, Pachypleurites, Paklapwania, Palaeotrachys, Palaeotylinae, Pallax, Palpatores, Pantodapus, Parabuprestium, Paracurculium, Paragrilium, Paragyrinus, Parasilphites, Parnidium, Parnosoma, Pedinosoma, Peltocoleops, Periboloptera, Peridosoma, Petalocupes, Petrorophus, Phaulogyrinus, Pholipheron, Phytoplesion, Pimeliodes, Plastobuprestites, Plastonebria, Platyprosopi, Pleuralocista, Polypamon, Praedodromeus, Prionophana, Prochrysomela, Proheuristes, Prophasis, Prosthecogmus, Prosthenostictus, Protocuneus, Prototoma, Pseudobuprestites, Pseudocarabites, Pseudocurculionites, Pseudocymindis, Pseudocyphon, Pseudoelateropsis, Pseudohydrophilites, Pseudoprionites, Pseudotelephorus, Pseudotenebrio, Pseudus, Rhomaleus, Rhynchophori, Saperda valdensis, Saprophagi, Scalopoides, Semiglobus, Sideriosemion, Silphidium, Sitonites, Smodicoptera, Sojanocoleidae, Sphaericites, Staphyliniae, Stenelytron, Sternoxi, Stictulus, Strongylites, Strullendorfia, Telephorium, Tenebrionites, Tetillopsis, Tetragonides, Thurmannia, Toksunius, Tolype, Triadocoleopteron, Trialarva, Trichelepturgetes, Tridigitati, Tripsalis, Trixagites, Trochiscites, Trochmalus, Trogossitarii, Trypocoleus, Tshekardocoleidae, Tychon, Ulomites, Velenovskya, Versicolores, Wollastonia, Xenogyrinus, Xinbinia, Xylophagi, Xylotupia, Yanqingia, Zacoleoptera, Zetemenos, Tachypus
Subtaxa: none
Ecology:
Distribution: there are no occurrences of Protocoleoptera in the database