Insecta - Coleoptera - Carabidae
Full reference: F. A. Bonelli. 1810. Observations Entomologiques. Première Partie (Cicindélètes et Portion des Carabiques) 1-58
Parent taxon: Harpalinae according to R. P. Webster 2016
See also Bouchard et al. 2011, LeConte 1861 and Tamutis et al. 2011
Sister taxa: Abacetini, Aethocarabus, Chlaeniini, Dryptini, Galeritini, Helluonini, Lebiini, Licinini, Odacanthini, Panagaeini, Pseudomorphini, Pterostichini, Sinis, Sphodrini, Zabrini, Zuphiini, Oodini
Subtaxa: Anisodactylina Harpalina Stenolophina
Type: Harpalus
Ecology:
Distribution:
• Quaternary of Canada (1: Ontario collection), the Russian Federation (26), Ukraine (1), the United Kingdom (4), United States (2: Colorado, Pennsylvania)
• Blancan of United States (1: Texas)
• Miocene of China (1), Croatia (1), Germany (3), the Russian Federation (1)
• Oligocene of France (7), Germany (2), United States (1: Montana)
• Eocene of France (1), United States (5: Colorado)
• Paleocene of France (1), Svalbard and Jan Mayen (1)
Total: 59 collections including 92 occurrences