Full reference: D. M. Unwin. 2003. On the phylogeny and evolutionary history of pterosaurs. Evolution and Palaeobiology of Pterosaurs, Geological Society of London, Special Publications 217:139-190
Parent taxon: Novialoidea according to B. Andres et al. 2010
See also Codorniú et al. 2016, Lü et al. 2010, O'Sullivan and Martill 2018, Unwin 2003 and Wei et al. 2021
Sister taxon: Lonchognatha
Subtaxa: Allkaruen Rhamphorhynchidae
Ecology: volant piscivore
Distribution:
• Cretaceous of China (1 collection)
• Jurassic to Cretaceous of Morocco (2)
• Jurassic of Argentina (1), China (6), Cuba (2), France (6), Germany (52), Kazakhstan (1), Kyrgyzstan (1), Morocco (1), Portugal (5), the Russian Federation (1), Switzerland (1), Tanzania (1), the United Kingdom (16)
Total: 97 collections including 113 occurrences