Parent taxon: Ungulata according to H. Gadow 1898
See also Ameghino 1894, Bond et al. 2006, Carroll 1988, Gregory 1910, Kramarz et al. 2010, McKenna 1975 and Shockey et al. 2009
Sister taxa: Altungulata, Amblypoda, Artiodactyla, Barypoda, Barytheria, Bulbulodentata, Cetungulata, Condylarthra, Diplarthra, Ganungulatum, Haloboidea, Hydrophorae, Hyraces, Kharmerungulatum, Meridiungulata, Monochenae, Palaeocladosictis, Procreodi, Prolatidens, Protungulata, Ruminantes, Tesserachenae, Tipotheria, Trichenae
Subtaxa: Adianthidae Eolicaphrium Eolitopterna Lambdaconus elegans Lopholipterna Macraucheniidae Mesorhinidae Notonychopidae Paulogervaisia celata Proacrodon Proacrodon transformatus Proadiantus gibbus Protolipternidae Sparnotheriodontidae Thoatheriopsis
Ecology: scansorial herbivore